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<mods:title id="6988D347F3244C77016D8FB7AC1C9058">New insights on the systematics and reproductive behaviour in tree frogs of the genus Feihyla, with description of a new related genus from Asia (Anura, Rhacophoridae)</mods:title>
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<mods:affiliation id="DBCDC3E3C4D96D6A86B363B42689AD4A">Systematics Lab, Department of Environmental Studies, University of Delhi, Delhi, India &amp; gopikacjnv 33 @ gmail. com; https: // orcid. org / 0000 - 0002 - 0047 - 0438</mods:affiliation>
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<treatment id="03F8BC2EFFB3FFFACBA7FBDEB77F9CBD" ID-DOI="http://doi.org/10.5281/zenodo.4425413" ID-GBIF-Taxon="180323983" ID-Zenodo-Dep="4425413" LSID="urn:lsid:plazi:treatment:03F8BC2EFFB3FFFACBA7FBDEB77F9CBD" httpUri="http://treatment.plazi.org/id/03F8BC2EFFB3FFFACBA7FBDEB77F9CBD" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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<taxonomicName id="4C5176BBFFB3FFFBCBA7FBDEB25A983B" authority="(Wilkinson, Win, Thin, Lwin, Shein, and Tun, 2003)" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" baseAuthorityName="Wilkinson, Win, Thin, Lwin, Shein, and Tun" baseAuthorityYear="2003" box="[151,1086,1080,1107]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="punctatus" status="comb. nov.">
<emphasis id="B925D12AFFB3FFFBCBA7FBDEB7D4983B" bold="true" box="[151,432,1080,1107]" italics="true" pageId="42" pageNumber="43">Rohanixalus punctatus</emphasis>
(
<bibRefCitation id="EFC070C9FFB3FFFBCAF0FBDFB251983B" author="Wilkinson, J. A. &amp; Win, H. &amp; Thin, T. &amp; Lwin, K. S. &amp; Shein, A. K. &amp; Tun, H." box="[448,1077,1081,1107]" pageId="42" pageNumber="43" pagination="17 - 26" refId="ref41264" refString="Wilkinson, J. A., Win, H., Thin, T., Lwin, K. S., Shein, A. K. &amp; Tun, H. (2003) A new species of Chirixalus (Anura: Rhacophoridae) from western Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (2), 17 - 26." type="journal article" year="2003">Wilkinson, Win, Thin, Lwin, Shein, and Tun, 2003</bibRefCitation>
)
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<paragraph id="8BEE0D38FFB3FFFBCBA7FBBBB7D39810" blockId="42.[151,1227,1080,1180]" box="[151,439,1117,1144]" pageId="42" pageNumber="43">Spotted Bubble-nest Frog</paragraph>
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<paragraph id="8BEE0D38FFB3FFFBCBA7FB67B7B698F3" blockId="42.[151,1227,1080,1180]" box="[151,466,1153,1180]" pageId="42" pageNumber="43">
(
<figureCitation id="136A11BDFFB3FFFBCBAFFB67B76F98F3" box="[159,267,1153,1179]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="18.[151,250,1817,1841]" captionStartId-1="19.[151,250,1411,1435]" captionTargetBox-0="[182,1408,187,1783]" captionTargetBox-1="[156,1427,182,1368]" captionTargetId-0="figure-29@18.[174,1412,181,1793]" captionTargetId-1="figure-17@19.[151,1436,181,1372]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURE 1. Maximum Likelihood tree showing phylogenetic relationships among 124 representative taxa from all recognised genera of the subfamily Rhacophorinae. Relationships are inferred based on 1,937 bp of mitochondrial (12SrRNA, tRNAVAL, 16SrRNA) and nuclear (RHO and RAG1) genes. Numbers above and below the branches indicate Bayesian Posterior Probabilities and RAxML bootstrap support values, respectively." captionText-1="FIGURE 2. AB. DNA Barcoding based on mitochondrial 16S rRNA sequences: A. genus Feihyla; B. genus Rohanixalus gen. nov.; C. distribution of the new genus Rohanixalus. Images represent the corresponding species and their broad geographical ranges." figureDoi-0="http://doi.org/10.5281/zenodo.4424574" figureDoi-1="http://doi.org/10.5281/zenodo.4424576" httpUri-0="https://zenodo.org/record/4424574/files/figure.png" httpUri-1="https://zenodo.org/record/4424576/files/figure.png" pageId="42" pageNumber="43">Figs. 12</figureCitation>
,
<figureCitation id="136A11BDFFB3FFFBCA27FB67B74398F3" box="[279,295,1153,1179]" captionStart="FIGURE 4" captionStartId="21.[151,250,1852,1876]" captionTargetBox="[160,1427,669,1828]" captionTargetId="figure-228@21.[159,1428,669,1828]" captionTargetPageId="21" captionText="FIGURE 4. Distribution of genetically confirmed records of all known Feihyla and Rohanixalus members (indicated by circles) and their respective type localities (indicated by triangles). Numbers corresponding to each species are indicated alongside the distribution spots." figureDoi="http://doi.org/10.5281/zenodo.4424582" httpUri="https://zenodo.org/record/4424582/files/figure.png" pageId="42" pageNumber="43">4</figureCitation>
,
<figureCitation id="136A11BDFFB3FFFBCA03FB64B72798F4" box="[307,323,1154,1180]" captionStart="FIGURE 7" captionStartId="27.[151,250,1588,1612]" captionTargetBox="[151,1436,777,1565]" captionTargetId="figure-298@27.[151,1436,777,1565]" captionTargetPageId="27" captionText="FIGURE 7. Members of the genus Rohanixalus. A. Dorsolateral view of R. hansenae from the type locality, Nong Khor, Thailand; B. dorsolateral view of R. marginis (holotype) from Perlis State Park chalets, Perlis, Peninsular Malaysia; C. dorsal view of R. cf. nauli, from Batang Gadis National Park, North Sumatra, Indonesia with a jelly-nest; D. dorsolateral view of R. cf. nauli, from Batang Gadis National Park, North Sumatra, Indonesia; E. dorsolateral view of R. cf. nauli, from Bukit Barisan Forest Park, North Sumatra, Indonesia; F. dorsolateral view of R. punctatus (Paratype, JBS 9274) from Gwa, Rakhine State, Myanmar (CC Public domain 3.0; AmphibiaWeb); G. dorsolateral view of R. vittatus from Rangat, Middle Andaman Islands, India; H. dorsolateral view of R. senapatiensis from Manipur, India; I. dorsolateral view of R. shyamrupus from Namdapha, Arunachal Pradesh." figureDoi="http://doi.org/10.5281/zenodo.4424590" httpUri="https://zenodo.org/record/4424590/files/figure.png" pageId="42" pageNumber="43">7</figureCitation>
;
<tableCitation id="C6D33883FFB3FFFBCA7FFB67B7C998F3" box="[335,429,1153,1179]" captionStart="TABLE 1" captionStartId="4.[151,239,462,488]" captionTargetPageId="4" captionText="TABLE 1. List of taxa and DNA sequences included in the study. (1) Indicates taxa used for the multi-gene phylogenetic analysis of family Rhacophoridae; (2) indicates taxa used for the 16S barcoding analyses of Feihyla and Rohanixalus gen. nov. Institutional abbreviations: SCUM (Zoological Museum of the School of Life Science, Sichuan University, China); NMNS (National Museum of Natural Science, Taiwan); KIZ (Kunming Institute of Zoology, China); KUHE (Kyoto University, Graduate School of Human and Environmental Studies, Japan); ZFMK (Zoologisches Forschungsinstitut und Museum Alexander Koenig, Germany); ROM (Royal Ontario Museum, Department of Herpetology, Toronto, Canada); THNHM (Thailand Natural History Museum, Thailand); HNNU (Department of Biology, Hunan Normal University, Changsha, China); WHT (Wildlife Heritage Trust, Colombo, Sri Lanka); MVZ (Museum of Vertebrate Zoology, University of California Berkeley, USA); ZMH (Zoologisches Museum Hamburg, Universität Hamburg, Germany); NMBE (Naturhistorisches Museum Bern, Switzerland); ZMKU (Zoological Museum, Kasetsart University, Bangkok, Thailand); MNHN (Muséum National dhistoire Naturelle, Paris, France); SN (Shunqing field number); ACD (Arvin C. Deismos field number); SDBDU (Systematics Lab, University of Delhi, India); CUMZ (Chulalongkorn University Museum of Natural History, Thailand); MZB (Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Sciences, Indonesia)." httpUri="http://table.plazi.org/id/DF2E5DB0FF9DFFD5CBA7FE28B1779F60" pageId="42" pageNumber="43" tableUuid="DF2E5DB0FF9DFFD5CBA7FE28B1779F60">Tables 1</tableCitation>
,
<tableCitation id="C6D33883FFB3FFFBCA8BFB67B7AD98F3" box="[443,457,1153,1179]" captionStart="TABLE 3" captionStartId="41.[151,239,660,686]" captionTargetPageId="41" captionText="TABLE 3. Inter and intraspecific uncorrected p-distances (in percent) in the genus Rohanixalus gen. nov. for the mitochondrial 16S rRNA gene sequences. Values indicate average distances followed by range in parentheses." httpUri="http://table.plazi.org/id/DF2E5DB0FFB0FFF8CBA7FD72B40C9EBA" pageId="42" pageNumber="43" tableUuid="DF2E5DB0FFB0FFF8CBA7FD72B40C9EBA">3</tableCitation>
)
</paragraph>
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<emphasis id="B925D12AFFB3FFFBCBA7FB2FB469988B" bold="true" box="[151,525,1225,1251]" pageId="42" pageNumber="43">Original name and description.</emphasis>
<taxonomicName id="4C5176BBFFB3FFFBC924FB2FB359988B" authority="Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003" authorityName="Wilkinson, Win, Thin, Lwin, Shein, and Tun" authorityYear="2003" box="[532,1341,1225,1252]" class="Amphibia" family="Rhacophoridae" genus="Chirixalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B925D12AFFB3FFFBC924FB2FB49B988B" box="[532,767,1225,1251]" italics="true" pageId="42" pageNumber="43">Chirixalus punctatus</emphasis>
<bibRefCitation id="EFC070C9FFB3FFFBC836FB2FB359988B" author="Wilkinson, J. A. &amp; Win, H. &amp; Thin, T. &amp; Lwin, K. S. &amp; Shein, A. K. &amp; Tun, H." box="[774,1341,1225,1252]" pageId="42" pageNumber="43" pagination="17 - 26" refId="ref41264" refString="Wilkinson, J. A., Win, H., Thin, T., Lwin, K. S., Shein, A. K. &amp; Tun, H. (2003) A new species of Chirixalus (Anura: Rhacophoridae) from western Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (2), 17 - 26." type="journal article" year="2003">Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003</bibRefCitation>
</taxonomicName>
. Wilkinson J.A., Win, H., Thin T., Lwin K.S., Shein A.K. and Tun H. 2003. A new species of
<taxonomicName id="4C5176BBFFB3FFFBCF5EFB0BB280996F" baseAuthorityName="Frost" baseAuthorityYear="2020" box="[1134,1252,1261,1287]" class="Amphibia" family="Rhacophoridae" genus="Chirixalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="B925D12AFFB3FFFBCF5EFB0BB280996F" box="[1134,1252,1261,1287]" italics="true" pageId="42" pageNumber="43">Chirixalus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C5176BBFFB3FFFBCFC4FB0BB324996F" authorityName="Fischer von Waldheim" authorityYear="1813" box="[1268,1344,1261,1287]" class="Magnoliopsida" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="order">Anura</taxonomicName>
:
<taxonomicName id="4C5176BBFFB3FFFBCE7BFB0BB69F9944" authorityName="Hoffman" authorityYear="1932" class="Amphibia" family="Rhacophoridae" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="family">Rhacophoridae</taxonomicName>
) from western
<collectingCountry id="F3464DA8FFB3FFFBCA97FAF4B4709944" box="[423,532,1298,1324]" name="Myanmar" pageId="42" pageNumber="43">Myanmar</collectingCountry>
(
<collectingCountry id="F3464DA8FFB3FFFBC913FAF7B4129943" box="[547,630,1297,1323]" name="Myanmar" pageId="42" pageNumber="43">Burma</collectingCountry>
). Proceedings of the California Academy of Sciences, 54: 1726.
<emphasis id="B925D12AFFB3FFFBCE6BFAF7B3F89943" bold="true" box="[1371,1436,1297,1323]" pageId="42" pageNumber="43">Type.</emphasis>
<typeStatus id="54EAB39AFFB3FFFBCBA7FAD3B7669927" box="[151,258,1333,1359]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
, CAS 221555, by original designation.
<emphasis id="B925D12AFFB3FFFBC98AFAD3B53C9927" bold="true" box="[698,856,1333,1359]" pageId="42" pageNumber="43">Type locality.</emphasis>
<collectingRegion id="4995C3DAFFB3FFFBC857FAD3B5A39927" box="[871,967,1333,1359]" country="Myanmar" name="Rakhine" pageId="42" pageNumber="43">Rakhine</collectingRegion>
Yoma Elephant Wildlife Sanctuary Headquarters (
<geoCoordinate id="EE656BFFFFB3FFFBCA30FABFB7CB991C" box="[256,431,1369,1396]" degrees="17" direction="north" minutes="36" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="48.8" value="17.613556">17° 36 48.8” N</geoCoordinate>
<geoCoordinate id="EE656BFFFFB3FFFBCA85FABFB407991C" box="[437,611,1369,1396]" degrees="94" direction="east" minutes="36" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="50.2" value="94.613945">94° 36 50.2” E</geoCoordinate>
), Gwa, Gwa Township,
<collectingRegion id="4995C3DAFFB3FFFBC85EFABFB26A991B" box="[878,1038,1369,1395]" country="Myanmar" name="Rakhine" pageId="42" pageNumber="43">Rakhine State</collectingRegion>
”,
<collectingCountry id="F3464DA8FFB3FFFBCF13FABCB2F6991C" box="[1059,1170,1370,1396]" name="Myanmar" pageId="42" pageNumber="43">Myanmar</collectingCountry>
.
<emphasis id="B925D12AFFB3FFFBCFABFABFB77C99FF" bold="true" pageId="42" pageNumber="43">Current status of specific name.</emphasis>
Valid name as
<taxonomicName id="4C5176BBFFB3FFFBCAF3FA9BB36A99FF" authority="(Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003)" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" baseAuthorityName="Wilkinson, Win, Thin, Lwin, Shein, and Tun" baseAuthorityYear="2003" box="[451,1294,1405,1432]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="punctatus" status="comb. nov.">
<emphasis id="B925D12AFFB3FFFBCAF3FA9BB4A799FF" box="[451,707,1405,1431]" italics="true" pageId="42" pageNumber="43">Rohanixalus punctatus</emphasis>
(
<bibRefCitation id="EFC070C9FFB3FFFBC9E1FA9BB36299FF" author="Wilkinson, J. A. &amp; Win, H. &amp; Thin, T. &amp; Lwin, K. S. &amp; Shein, A. K. &amp; Tun, H." box="[721,1286,1405,1432]" pageId="42" pageNumber="43" pagination="17 - 26" refId="ref41264" refString="Wilkinson, J. A., Win, H., Thin, T., Lwin, K. S., Shein, A. K. &amp; Tun, H. (2003) A new species of Chirixalus (Anura: Rhacophoridae) from western Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (2), 17 - 26." type="journal article" year="2003">Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003</bibRefCitation>
)
</taxonomicName>
<emphasis id="B925D12AFFB3FFFBCE25FA9BB3F199FF" bold="true" box="[1301,1429,1405,1431]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="A2166C51FFB3FFFBCE25FA9BB3F199FF" box="[1301,1429,1405,1431]" pageId="42" pageNumber="43" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BEE0D38FFB3FFFBCBF7FA47B5ED9B4B" blockId="42.[151,1437,1225,2044]" pageId="42" pageNumber="43">
<emphasis id="B925D12AFFB3FFFBCBF7FA47B7D999D3" bold="true" box="[199,445,1441,1467]" pageId="42" pageNumber="43">Taxonomic remarks.</emphasis>
Based on morphological affinity, this species is closely related to members of the genus
<taxonomicName id="4C5176BBFFB3FFFBCBA7FA23B74799B7" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" box="[151,291,1477,1503]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="B925D12AFFB3FFFBCBA7FA23B74799B7" box="[151,291,1477,1503]" italics="true" pageId="42" pageNumber="43">Rohanixalus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC070C9FFB3FFFBCA02FA23B45D99B7" author="Wilkinson, J. A. &amp; Win, H. &amp; Thin, T. &amp; Lwin, K. S. &amp; Shein, A. K. &amp; Tun, H." box="[306,569,1477,1504]" pageId="42" pageNumber="43" pagination="17 - 26" refId="ref41264" refString="Wilkinson, J. A., Win, H., Thin, T., Lwin, K. S., Shein, A. K. &amp; Tun, H. (2003) A new species of Chirixalus (Anura: Rhacophoridae) from western Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (2), 17 - 26." type="journal article" year="2003">
Wilkinson
<emphasis id="B925D12AFFB3FFFBCA9DFA20B7BB99B7" box="[429,479,1477,1503]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
. (2003)
</bibRefCitation>
diagnosed the species on the basis of differences in dorsal colour and markings with the closely related
<taxonomicName id="4C5176BBFFB3FFFBCAEBFA0CB4289A6B" box="[475,588,1514,1539]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="B925D12AFFB3FFFBCAEBFA0CB4289A6B" box="[475,588,1514,1539]" italics="true" pageId="42" pageNumber="43">R. vittatus</emphasis>
</taxonomicName>
, but subsequently stated in the variations section that the extent of spotting varies, being “sparser in some individuals, restricted primarily to the dorsal aspect of the head whereas in others to the entire dorsum. The accompanying
<typeStatus id="54EAB39AFFB3FFFBC976F9D4B4C29A24" box="[582,678,1586,1612]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
photograph (Joseph B. Slowinski field voucher number JBS 9274, deposited in CAS;
<bibRefCitation id="EFC070C9FFB3FFFBCA45F9B3B40B9A07" author="Wilkinson, J. A. &amp; Win, H. &amp; Thin, T. &amp; Lwin, K. S. &amp; Shein, A. K. &amp; Tun, H." box="[373,623,1621,1648]" pageId="42" pageNumber="43" pagination="17 - 26" refId="ref41264" refString="Wilkinson, J. A., Win, H., Thin, T., Lwin, K. S., Shein, A. K. &amp; Tun, H. (2003) A new species of Chirixalus (Anura: Rhacophoridae) from western Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (2), 17 - 26." type="journal article" year="2003">
Wilkinson
<emphasis id="B925D12AFFB3FFFBCAC2F9B0B4419A07" box="[498,549,1621,1647]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
. 2003
</bibRefCitation>
) also shows close similarities to another species,
<taxonomicName id="4C5176BBFFB3FFFBCF9AF9B0B34A9A07" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" baseAuthorityName="Chan, Grismer, Anuar, Quah, Grismer, Wood, Muin, and Ahmad" baseAuthorityYear="2011" box="[1194,1326,1622,1647]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="marginis">
<emphasis id="B925D12AFFB3FFFBCF9AF9B0B34A9A07" box="[1194,1326,1622,1647]" italics="true" pageId="42" pageNumber="43">R. marginis</emphasis>
</taxonomicName>
, that was described more recently from Peninsular
<collectingCountry id="F3464DA8FFB3FFFBC955F99FB4AB9AFB" box="[613,719,1657,1683]" name="Malaysia" pageId="42" pageNumber="43">Malaysia</collectingCountry>
. Based on available evidence, we formally reallocate
<taxonomicName id="4C5176BBFFB3FFFBCE16F99FB5229ADF" authority="Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003" authorityName="Wilkinson, Win, Thin, Lwin, Shein, and Tun" authorityYear="2003" class="Amphibia" family="Rhacophoridae" genus="Chirixalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B925D12AFFB3FFFBCE16F99FB7609ADF" italics="true" pageId="42" pageNumber="43">Chirixalus punctatus</emphasis>
<bibRefCitation id="EFC070C9FFB3FFFBCA3CF97BB5229ADF" author="Wilkinson, J. A. &amp; Win, H. &amp; Thin, T. &amp; Lwin, K. S. &amp; Shein, A. K. &amp; Tun, H." box="[268,838,1693,1720]" pageId="42" pageNumber="43" pagination="17 - 26" refId="ref41264" refString="Wilkinson, J. A., Win, H., Thin, T., Lwin, K. S., Shein, A. K. &amp; Tun, H. (2003) A new species of Chirixalus (Anura: Rhacophoridae) from western Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (2), 17 - 26." type="journal article" year="2003">Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003</bibRefCitation>
</taxonomicName>
to the genus
<taxonomicName id="4C5176BBFFB3FFFBC8D0F97BB2089ADF" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" box="[992,1132,1693,1719]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="B925D12AFFB3FFFBC8D0F97BB2089ADF" box="[992,1132,1693,1719]" italics="true" pageId="42" pageNumber="43">Rohanixalus</emphasis>
</taxonomicName>
, as
<taxonomicName id="4C5176BBFFB3FFFBCFABF97BB4BC9AB3" authority="(Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003)" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" baseAuthorityName="Wilkinson, Win, Thin, Lwin, Shein, and Tun" baseAuthorityYear="2003" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="punctatus" status="comb. nov.">
<emphasis id="B925D12AFFB3FFFBCFABF97BB3F89ADF" box="[1179,1436,1693,1719]" italics="true" pageId="42" pageNumber="43">Rohanixalus punctatus</emphasis>
(
<bibRefCitation id="EFC070C9FFB3FFFBCBAFF927B4B49AB3" author="Wilkinson, J. A. &amp; Win, H. &amp; Thin, T. &amp; Lwin, K. S. &amp; Shein, A. K. &amp; Tun, H." box="[159,720,1729,1756]" pageId="42" pageNumber="43" pagination="17 - 26" refId="ref41264" refString="Wilkinson, J. A., Win, H., Thin, T., Lwin, K. S., Shein, A. K. &amp; Tun, H. (2003) A new species of Chirixalus (Anura: Rhacophoridae) from western Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (2), 17 - 26." type="journal article" year="2003">Wilkinson, Win, Thin, Lwin, Shein, and Tun 2003</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2166C51FFB3FFFBC9EEF927B5049AB4" box="[734,864,1729,1756]" pageId="42" pageNumber="43" rank="species">
<emphasis id="B925D12AFFB3FFFBC9EEF927B53D9AB3" bold="true" box="[734,857,1729,1755]" pageId="42" pageNumber="43">comb. nov</emphasis>
.
</taxonomicNameLabel>
Further molecular evidence can clarify the systematic relationship of
<taxonomicName id="4C5176BBFFB3FFFBCA44F900B4659A97" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" baseAuthorityName="Wilkinson, Win, Thin, Lwin, Shein, and Tun" baseAuthorityYear="2003" box="[372,513,1766,1791]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B925D12AFFB3FFFBCA44F900B4659A97" box="[372,513,1766,1791]" italics="true" pageId="42" pageNumber="43">R. punctatus</emphasis>
</taxonomicName>
with other
<taxonomicName id="4C5176BBFFB3FFFBC9B5F903B5759A97" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" box="[645,785,1765,1791]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="B925D12AFFB3FFFBC9B5F903B5759A97" box="[645,785,1765,1791]" italics="true" pageId="42" pageNumber="43">Rohanixalus</emphasis>
</taxonomicName>
members, especially
<taxonomicName id="4C5176BBFFB3FFFBCF37F900B2EE9A97" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" baseAuthorityName="Chan, Grismer, Anuar, Quah, Grismer, Wood, Muin, and Ahmad" baseAuthorityYear="2011" box="[1031,1162,1766,1791]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="marginis">
<emphasis id="B925D12AFFB3FFFBCF37F900B2EE9A97" box="[1031,1162,1766,1791]" italics="true" pageId="42" pageNumber="43">R. marginis</emphasis>
</taxonomicName>
, which could be a close congener based on morphological and geographical considerations.
</paragraph>
</subSubSection>
<subSubSection id="C34B5EB3FFB3FFFBCBF7F8CBB5E99B94" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BEE0D38FFB3FFFBCBF7F8CBB5E99B94" blockId="42.[151,1437,1225,2044]" pageId="42" pageNumber="43">
<emphasis id="B925D12AFFB3FFFBCBF7F8CBB7269B2F" bold="true" box="[199,322,1837,1863]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
Small-sized adults (male, SVL
<quantity id="4CA9A0DDFFB3FFFBC981F8CBB5629B20" box="[689,774,1837,1864]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="42" pageNumber="43" unit="mm" value="23.0">23 mm</quantity>
), with slender body; snout sub-elliptical to nearly pointed in dorsal view; tympanum externally obscure; dorsal skin smooth; the entire dorsum, lateral surfaces, and dorsal surface of limbs covered with fine dark brown speckles; dorsum and limbs with or without prominent spots; a pair of continuous dorsolateral stripes starting from the snout tip, extending over the eyes, and ending close to the vent; webbing absent between fingers I, II, and III, basal webbing present between fingers III and IV; foot webbing moderate, up to the second subarticular tubercle on either side of toe IV.
</paragraph>
</subSubSection>
<subSubSection id="C34B5EB3FFB2FFFACBF7FF70B77F9CBD" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BEE0D38FFB2FFFACBF7FF70B77F9CBD" blockId="43.[151,1436,150,213]" pageId="43" pageNumber="44">
<emphasis id="B925D12AFFB2FFFACBF7FF70B7059CD8" bold="true" box="[199,353,150,176]" pageId="43" pageNumber="44">Distribution.</emphasis>
<taxonomicName id="4C5176BBFFB2FFFACA58FF71B40C9CD8" authorityName="Biju &amp; Garg &amp; Gokulakrishnan &amp; Chandrakasan &amp; Thammachoti &amp; Ren &amp; Gopika &amp; Bisht &amp; Hamidy &amp; Shouche" authorityYear="2020" baseAuthorityName="Wilkinson, Win, Thin, Lwin, Shein, and Tun" baseAuthorityYear="2003" box="[360,616,151,177]" class="Amphibia" family="Rhacophoridae" genus="Rohanixalus" kingdom="Animalia" order="Anura" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B925D12AFFB2FFFACA58FF71B40C9CD8" box="[360,616,151,177]" italics="true" pageId="43" pageNumber="44">Rohanixalus punctatus</emphasis>
</taxonomicName>
is currently known only from western
<collectingCountry id="F3464DA8FFB2FFFACF2FFF71B2EA9CD9" box="[1055,1166,151,177]" name="Myanmar" pageId="43" pageNumber="44">Myanmar</collectingCountry>
, the place of its original description.
</paragraph>
</subSubSection>
</treatment>
</document>