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<document id="8729CD29259E2559F3B04E4106FBCC42" ID-DOI="10.11646/zootaxa.5506.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747442" ID-ZooBank="44E64616-6FA5-44E7-820D-59EF94E7D6F3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726055075908" checkinUser="plazi" docAuthor="Ermilov, Sergey G." docDate="2024" docId="03E88946B82AC622FF41C4A808F46D3A" docLanguage="en" docName="zootaxa.5506.4.5.pdf" docOrigin="Zootaxa 5506 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratorchestes (Ceratorchestes) duocornutus Ermilov, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="561" masterDocId="FFD1F13EB82EC624FFD6C701091E6F07" masterDocTitle="New Ceratorchestes (Acari, Oribatida, Peloppiidae) from Venezuela" masterLastPageNumber="566" masterPageNumber="555" pageNumber="559" updateTime="1726055901464" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="86E5548FB087FCB6AAC698BD5D6A3208">New Ceratorchestes (Acari, Oribatida, Peloppiidae) from Venezuela</mods:title>
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<mods:namePart id="616A752F9DCCC80C82449E04A455BD49">Ermilov, Sergey G.</mods:namePart>
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<treatment id="03E88946B82AC622FF41C4A808F46D3A" ID-DOI="http://doi.org/10.5281/zenodo.13747359" ID-Zenodo-Dep="13747359" LSID="urn:lsid:plazi:treatment:03E88946B82AC622FF41C4A808F46D3A" httpUri="http://treatment.plazi.org/id/03E88946B82AC622FF41C4A808F46D3A" lastPageId="6" lastPageNumber="561" pageId="4" pageNumber="559">
<subSubSection id="C35B6BDBB82AC620FF41C4A80A006CC3" box="[151,798,937,964]" pageId="4" pageNumber="559" type="nomenclature">
<paragraph id="8BFE3850B82AC620FF41C4A80A006CC3" blockId="4.[151,798,937,1000]" box="[151,798,937,964]" pageId="4" pageNumber="559">
<heading id="D0B68F3CB82AC620FF41C4A80A006CC3" bold="true" box="[151,798,937,964]" fontSize="11" level="1" pageId="4" pageNumber="559" reason="1">
<emphasis id="B935E442B82AC620FF41C4A80A006CC3" bold="true" box="[151,798,937,964]" pageId="4" pageNumber="559">
<taxonomicName id="4C4143D3B82AC620FF41C4A80BA76CC3" authorityName="Ermilov" authorityYear="2024" box="[151,697,937,964]" class="Arachnida" family="Ceratoppiidae" genus="Ceratorchestes" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="duocornutus" status="sp. nov." subGenus="Ceratorchestes">
<emphasis id="B935E442B82AC620FF41C4A808546CC3" bold="true" box="[151,330,937,964]" italics="true" pageId="4" pageNumber="559">Ceratorchestes</emphasis>
(
<emphasis id="B935E442B82AC620FE8AC4A80B116CC3" bold="true" box="[348,527,937,964]" italics="true" pageId="4" pageNumber="559">Ceratorchestes</emphasis>
)
<emphasis id="B935E442B82AC620FDF6C4A80BA76CC3" bold="true" box="[544,697,937,964]" italics="true" pageId="4" pageNumber="559">duocornutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2065939B82AC620FD16C4AB0A006CC3" box="[704,798,938,964]" pageId="4" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B6BDBB82AC620FF41C4CF09F56CEF" box="[151,235,974,1000]" pageId="4" pageNumber="559" type="description">
<paragraph id="8BFE3850B82AC620FF41C4CF09F56CEF" blockId="4.[151,798,937,1000]" box="[151,235,974,1000]" pageId="4" pageNumber="559">
(
<figureCitation id="137A24D5B82AC620FF49C4CF09FC6CEF" box="[159,226,974,1000]" captionStart="FIGURE 3" captionStartId="5.[151,250,1652,1676]" captionTargetBox="[165,1422,194,1621]" captionTargetId="figure-18@5.[151,1436,181,1629]" captionTargetPageId="5" captionText="FIGURE 3. Ceratorchestes (Ceratorchestes) duocornutus sp. nov., adult, male (A, B) and female (C, D): A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted); D—anogenital region. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13747448" httpUri="https://zenodo.org/record/13747448/files/figure.png" pageId="4" pageNumber="559">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35B6BDBB82AC620FF41C3140A736BC7" pageId="4" pageNumber="559" type="materials_examined">
<paragraph id="8BFE3850B82AC620FF41C3140BD06B53" blockId="4.[151,1437,1045,2008]" pageId="4" pageNumber="559">
<emphasis id="B935E442B82AC620FF41C31408186B28" bold="true" box="[151,262,1045,1071]" pageId="4" pageNumber="559">Material.</emphasis>
<materialsCitation id="3B29320DB82AC620FED9C3170BD06B53" collectingDate="1995-02-26" collectionCode="R" collectorName="W. Brooks" country="Venezuela" elevation="1450" location="Rancho Grande" pageId="4" pageNumber="559" specimenCount="4" specimenCount-male="2" stateProvince="Aragua" typeStatus="holotype">
<typeStatus id="54FA86F2B82AC620FED9C31708696B37" box="[271,375,1046,1072]" pageId="4" pageNumber="559" type="holotype">Holotype</typeStatus>
(female) and
<specimenCount id="9D47F3D9B82AC620FDCCC3170BA96B37" box="[538,695,1046,1072]" count="2" pageId="4" pageNumber="559" type="generic" typeStatus="paratype">two paratypes</specimenCount>
(
<specimenCount id="9D47F3D9B82AC620FD1FC3170A5A6B37" box="[713,836,1046,1072]" count="2" pageId="4" pageNumber="559" type="male">two males</specimenCount>
): northern
<collectingCountry id="F35678C0B82AC620FC16C3170D286B37" box="[960,1078,1046,1072]" name="Venezuela" pageId="4" pageNumber="559">Venezuela</collectingCountry>
,
<collectingRegion id="4985F6B2B82AC620FB97C3170DC96B37" box="[1089,1239,1046,1072]" country="Venezuela" name="Aragua" pageId="4" pageNumber="559">Aragua State</collectingRegion>
,
<location id="8E9E6E8BB82AC620FB32C3170C866B37" LSID="urn:lsid:plazi:treatment:03E88946B82AC622FF41C4A808F46D3A:8E9E6E8BB82AC620FB32C3170C866B37" box="[1252,1432,1046,1072]" country="Venezuela" name="Rancho Grande" pageId="4" pageNumber="559" stateProvince="Aragua">Rancho Grande</location>
,
<elevation id="006CDF63B82AC620FF41C33808376B53" box="[151,297,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="559" unit="m" value="1450.0">
<quantity id="4CB995B5B82AC620FF41C33809F26B53" box="[151,236,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="559" unit="m" value="1450.0">1450 m</quantity>
a.s.l.
</elevation>
, leaf litter,
<date id="FFFF1E90B82AC620FE7EC3380B026B53" box="[424,540,1081,1108]" pageId="4" pageNumber="559" value="1995-02-26">
<collectingDate id="EFBBE778B82AC620FE7EC3380B026B53" box="[424,540,1081,1108]" pageId="4" pageNumber="559" value="1995-02-26">26.II.1995</collectingDate>
</date>
(
<collectionCode id="ED50A095B82AC620FDFBC33B0B5E6B53" box="[557,576,1082,1108]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="559">R</collectionCode>
.
<collectorName id="26B45D86B82AC620FD95C33B0BDC6B53" box="[579,706,1082,1108]" pageId="4" pageNumber="559">W. Brooks</collectorName>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFE3850B82AC620FF11C35F0A736BC7" blockId="4.[151,1437,1045,2008]" pageId="4" pageNumber="559">
The
<typeStatus id="54FA86F2B82AC620FF2FC35F08456B7F" box="[249,347,1118,1144]" pageId="4" pageNumber="559" type="holotype">holotype</typeStatus>
is deposited in the collection of the Canadian National Collection, Ottawa,
<collectingCountry id="F35678C0B82AC620FB49C35F0DE96B7F" box="[1183,1271,1118,1144]" name="Canada" pageId="4" pageNumber="559">Canada</collectingCountry>
;
<materialsCitation id="3B29320DB82AC620FAD7C35F0DEB6B9B" country="Russia" location="University Museum of Zoology" pageId="4" pageNumber="559" specimenCount="2" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount id="9D47F3D9B82AC620FAD7C35F0C826B7F" box="[1281,1436,1118,1144]" count="2" pageId="4" pageNumber="559" type="generic" typeStatus="paratype">two paratypes</specimenCount>
are deposited in the collection of the
<collectingRegion id="4985F6B2B82AC620FDE7C3830BD56B9B" box="[561,715,1154,1180]" country="Russia" name="Tyumen'" pageId="4" pageNumber="559">Tyumen State</collectingRegion>
<location id="8E9E6E8BB82AC620FD07C3830D2A6B9B" LSID="urn:lsid:plazi:treatment:03E88946B82AC622FF41C4A808F46D3A:8E9E6E8BB82AC620FD07C3830D2A6B9B" box="[721,1076,1154,1180]" country="Russia" name="University Museum of Zoology" pageId="4" pageNumber="559" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="4985F6B2B82AC620FBE8C3830D826B9B" box="[1086,1180,1154,1180]" country="Russia" name="Tyumen'" pageId="4" pageNumber="559">Tyumen</collectingRegion>
,
<collectingCountry id="F35678C0B82AC620FB71C3830DEB6B9B" box="[1191,1269,1154,1180]" name="Russia" pageId="4" pageNumber="559">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
<subSubSection id="C35B6BDBB82AC620FF11C3C808566AE7" pageId="4" pageNumber="559" type="diagnosis">
<paragraph id="8BFE3850B82AC620FF11C3C808566AE7" blockId="4.[151,1437,1045,2008]" pageId="4" pageNumber="559">
<emphasis id="B935E442B82AC620FF11C3C8085C6BE4" bold="true" box="[199,322,1225,1251]" pageId="4" pageNumber="559">Diagnosis.</emphasis>
Body length: 375390. Rostrum tripartite; rostral protrusions triangular, simple, similar in size. Lateral tooth of prodorsum strong. Lamellar cusp with well-developed lateral tooth and oblique medial part; translamella absent. Rostral and lamellar setae medium-sized, setiform, barbed; interlamellar seta long, slightly thickened, stiff, acuminate, barbed; bothridial seta long, setiform, shortly ciliated. All notogastral setae (except minute
<emphasis id="B935E442B82AC620FF3BC25B09E76A74" box="[237,249,1370,1395]" italics="true" pageId="4" pageNumber="559">c</emphasis>
) slightly thickened, stiff, acuminate, barbed;
<emphasis id="B935E442B82AC620FD2DC25B0A176A74" box="[763,777,1370,1395]" italics="true" pageId="4" pageNumber="559">p</emphasis>
<subScript id="17C53A15B82AC620FCDFC26D0A0F6A7D" attach="left" box="[777,785,1388,1402]" fontSize="6" pageId="4" pageNumber="559">1</subScript>
<emphasis id="B935E442B82AC620FCC9C25B0A336A74" box="[799,813,1370,1395]" italics="true" pageId="4" pageNumber="559">p</emphasis>
<subScript id="17C53A15B82AC620FCFBC26D0A2B6A7D" attach="left" box="[813,821,1388,1402]" fontSize="6" pageId="4" pageNumber="559">3</subScript>
shorter than others. Cheliceral seta
<emphasis id="B935E442B82AC620FB1AC25B0DEA6A73" box="[1228,1268,1370,1396]" italics="true" pageId="4" pageNumber="559">cha</emphasis>
comparatively short. Epimeral setae
<emphasis id="B935E442B82AC620FE50C27F08BD6A9F" box="[390,419,1406,1432]" italics="true" pageId="4" pageNumber="559">1b</emphasis>
,
<emphasis id="B935E442B82AC620FE79C27F08D46A9F" box="[431,458,1406,1432]" italics="true" pageId="4" pageNumber="559">1c</emphasis>
,
<emphasis id="B935E442B82AC620FE00C27F08EC6A9F" box="[470,498,1406,1432]" italics="true" pageId="4" pageNumber="559">3b</emphasis>
medium-sized, longer than others, slightly thickened, stiff, acuminate;
<emphasis id="B935E442B82AC620FAD0C27F0C3C6A9F" box="[1286,1314,1406,1432]" italics="true" pageId="4" pageNumber="559">3a</emphasis>
inserted in the same oblique row with setae of epimere IV. Adanal lyrifissure oblique to anal plate. Leg trochanter IV rounded dorsoanteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C35B6BDBB82AC622FF11C2E80BB86E8E" lastPageId="6" lastPageNumber="561" pageId="4" pageNumber="559" type="description">
<paragraph id="8BFE3850B82AC620FF11C2E80B10692F" blockId="4.[151,1437,1045,2008]" pageId="4" pageNumber="559">
<emphasis id="B935E442B82AC620FF11C2E808A56903" bold="true" box="[199,443,1513,1540]" pageId="4" pageNumber="559">Description of adult</emphasis>
.
<emphasis id="B935E442B82AC620FE1BC2EB0B716904" box="[461,623,1514,1539]" italics="true" pageId="4" pageNumber="559">Measurements</emphasis>
. Body length: 390 (
<typeStatus id="54FA86F2B82AC620FC8DC2EB0ADD6903" box="[859,963,1514,1540]" pageId="4" pageNumber="559" type="holotype">holotype</typeStatus>
), 375 (male
<typeStatus id="54FA86F2B82AC620FB8EC2EB0DD76903" box="[1112,1225,1514,1540]" pageId="4" pageNumber="559" type="paratype">paratypes</typeStatus>
); body width: 285 (
<typeStatus id="54FA86F2B82AC620FF49C10F0819692F" box="[159,263,1550,1576]" pageId="4" pageNumber="559" type="holotype">holotype</typeStatus>
), 270 (male
<typeStatus id="54FA86F2B82AC620FE44C10F0B1D692F" box="[402,515,1550,1576]" pageId="4" pageNumber="559" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="8BFE3850B82AC620FF11C1330B506993" blockId="4.[151,1437,1045,2008]" pageId="4" pageNumber="559">
<emphasis id="B935E442B82AC620FF11C133085B694C" box="[199,325,1586,1611]" italics="true" pageId="4" pageNumber="559">Integument</emphasis>
. Body color light brown. Surface densely microgranulate sculpturing (visible only under high magnification in dissected specimens; × 1000); lateral part of prodorsum between bothridium and acetabula IIII with larger granules (diameter up to 1).
</paragraph>
<paragraph id="8BFE3850B82AC620FF11C19F0A54686B" blockId="4.[151,1437,1045,2008]" pageId="4" pageNumber="559">
<emphasis id="B935E442B82AC620FF11C19F085A69BF" box="[199,324,1694,1720]" italics="true" pageId="4" pageNumber="559">Prodorsum</emphasis>
(
<figureCitation id="137A24D5B82AC620FE85C19F08CC69BF" box="[339,466,1694,1720]" captionStart="FIGURE 3" captionStartId="5.[151,250,1652,1676]" captionTargetBox="[165,1422,194,1621]" captionTargetId="figure-18@5.[151,1436,181,1629]" captionTargetPageId="5" captionText="FIGURE 3. Ceratorchestes (Ceratorchestes) duocornutus sp. nov., adult, male (A, B) and female (C, D): A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted); D—anogenital region. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13747448" httpUri="https://zenodo.org/record/13747448/files/figure.png" pageId="4" pageNumber="559">Figs 3A, C</figureCitation>
). Rostrum tripartite; all rostral protrusions triangular, slightly rounded apically, similar in size. Lateral tooth of prodorsum
<emphasis id="B935E442B82AC620FDE3C1C30B5569DC" box="[565,587,1730,1755]" italics="true" pageId="4" pageNumber="559">tp</emphasis>
strong. Lamella about 2/3 length of prodorsum; lamellar cusp with well-developed lateral tooth and oblique medial part; translamella absent; sublamella not observable. Rostral (3437) and lamellar (4549) setae setiform, barbed; rostral seta inserted medial to tutorial end and distanced from it; interlamellar seta (7579) slightly thickened, stiff, acuminate, barbed; bothridial seta (94101) setiform, shortly ciliated; exobothridial seta (2630) setiform, thin, roughened.
</paragraph>
<paragraph id="8BFE3850B82AC620FF11C0770BD268DF" blockId="4.[151,1437,1045,2008]" pageId="4" pageNumber="559">
<taxonomicName id="4C4143D3B82AC620FF11C077085D6888" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[199,323,1910,1935]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B935E442B82AC620FF11C077085D6888" box="[199,323,1910,1935]" italics="true" pageId="4" pageNumber="559">Notogaster</emphasis>
</taxonomicName>
(
<figureCitation id="137A24D5B82AC620FE84C07708D16897" box="[338,463,1910,1936]" captionStart="FIGURE 3" captionStartId="5.[151,250,1652,1676]" captionTargetBox="[165,1422,194,1621]" captionTargetId="figure-18@5.[151,1436,181,1629]" captionTargetPageId="5" captionText="FIGURE 3. Ceratorchestes (Ceratorchestes) duocornutus sp. nov., adult, male (A, B) and female (C, D): A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted); D—anogenital region. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13747448" httpUri="https://zenodo.org/record/13747448/files/figure.png" pageId="4" pageNumber="559">Figs 3A, C</figureCitation>
). Anterior notogastral margin convex medially. Notogastral seta
<emphasis id="B935E442B82AC620FB4AC0770DB66888" box="[1180,1192,1910,1935]" italics="true" pageId="4" pageNumber="559">c</emphasis>
(4) needleform; other notogastral setae (
<emphasis id="B935E442B82AC620FEB3C09B086568B3" box="[357,379,1946,1972]" italics="true" pageId="4" pageNumber="559">la</emphasis>
,
<emphasis id="B935E442B82AC620FE5CC09B08B868B3" box="[394,422,1946,1972]" italics="true" pageId="4" pageNumber="559">lm</emphasis>
,
<emphasis id="B935E442B82AC620FE63C09B08D568B3" box="[437,459,1946,1972]" italics="true" pageId="4" pageNumber="559">lp</emphasis>
,
<emphasis id="B935E442B82AC620FE0FC09B08F968B3" box="[473,487,1946,1972]" italics="true" pageId="4" pageNumber="559">h</emphasis>
<subScript id="17C53A15B82AC620FE31C0AD08F168BD" attach="left" box="[487,495,1964,1978]" fontSize="6" pageId="4" pageNumber="559">1</subScript>
<emphasis id="B935E442B82AC620FE28C09B0B1268B3" box="[510,524,1946,1972]" italics="true" pageId="4" pageNumber="559">h</emphasis>
<subScript id="17C53A15B82AC620FDDAC0AD0B0A68BD" attach="left" box="[524,532,1964,1978]" fontSize="6" pageId="4" pageNumber="559">3</subScript>
: 8694;
<emphasis id="B935E442B82AC620FDAFC09B0B9968B4" box="[633,647,1946,1971]" italics="true" pageId="4" pageNumber="559">p</emphasis>
<subScript id="17C53A15B82AC620FD51C0AD0B9168BD" attach="left" box="[647,655,1964,1978]" fontSize="6" pageId="4" pageNumber="559">1</subScript>
<emphasis id="B935E442B82AC620FD48C09B0BB268B4" box="[670,684,1946,1971]" italics="true" pageId="4" pageNumber="559">p</emphasis>
<subScript id="17C53A15B82AC620FD7AC0AD0BAA68BD" attach="left" box="[684,692,1964,1978]" fontSize="6" pageId="4" pageNumber="559">3</subScript>
: 7175) slightly thickened, stiff, acuminate, barbed. Opisthonotal gland opening and all lyrifissures well observable.
</paragraph>
<caption id="DF3E68D8B82BC621FF41C175080D69D2" ID-DOI="http://doi.org/10.5281/zenodo.13747448" ID-Zenodo-Dep="13747448" httpUri="https://zenodo.org/record/13747448/files/figure.png" pageId="5" pageNumber="560" startId="5.[151,250,1652,1676]" targetBox="[165,1422,194,1621]" targetPageId="5" targetType="figure">
<paragraph id="8BFE3850B82BC621FF41C175080D69D2" blockId="5.[151,1436,1652,1749]" pageId="5" pageNumber="560">
<emphasis id="B935E442B82BC621FF41C175080C698B" bold="true" box="[151,274,1652,1676]" pageId="5" pageNumber="560">FIGURE 3.</emphasis>
<taxonomicName id="4C4143D3B82BC621FECEC1750BFB698B" authorityName="Ermilov" authorityYear="2024" box="[280,741,1652,1677]" class="Arachnida" family="Ceratoppiidae" genus="Ceratorchestes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="duocornutus" status="sp. nov." subGenus="Ceratorchestes">
<emphasis id="B935E442B82BC621FECEC17508AE698B" box="[280,432,1652,1676]" italics="true" pageId="5" pageNumber="560">Ceratorchestes</emphasis>
(
<emphasis id="B935E442B82BC621FE69C1750B49698B" box="[447,599,1652,1676]" italics="true" pageId="5" pageNumber="560">Ceratorchestes</emphasis>
)
<emphasis id="B935E442B82BC621FDB0C1750BFB698B" box="[614,741,1652,1676]" italics="true" pageId="5" pageNumber="560">duocornutus</emphasis>
</taxonomicName>
<emphasis id="B935E442B82BC621FD3DC1740A25698A" bold="true" box="[747,827,1653,1677]" pageId="5" pageNumber="560">
<taxonomicNameLabel id="A2065939B82BC621FD3DC1740A25698A" box="[747,827,1653,1677]" pageId="5" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult, male (A, B) and female (C, D): A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted); D—anogenital region. Scale bar 100 μm.
</paragraph>
</caption>
<paragraph id="8BFE3850B82BC621FF11C0260C6D68AA" blockId="5.[151,1436,1831,2037]" pageId="5" pageNumber="560">
<taxonomicName id="4C4143D3B82BC621FF11C02608486846" authorityName="Krefft" authorityYear="1871" box="[199,342,1831,1857]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="5" pageNumber="560" phylum="Nematoda" rank="genus">
<emphasis id="B935E442B82BC621FF11C02608486846" box="[199,342,1831,1857]" italics="true" pageId="5" pageNumber="560">Gnathosoma</emphasis>
</taxonomicName>
. Similar to
<taxonomicName id="4C4143D3B82BC621FE08C0260BB86846" authorityName="Ermilov" authorityYear="2024" box="[478,678,1831,1857]" class="Arachnida" family="Ceratoppiidae" genus="Ceratorchestes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="tripartitus" status="sp. nov." subGenus="Ceratorchestes">
<emphasis id="B935E442B82BC621FE08C02608E96846" box="[478,503,1831,1857]" italics="true" pageId="5" pageNumber="560">C.</emphasis>
(
<emphasis id="B935E442B82BC621FDDFC0260B3C6846" box="[521,546,1831,1857]" italics="true" pageId="5" pageNumber="560">C.</emphasis>
)
<emphasis id="B935E442B82BC621FDE2C0290BB86846" box="[564,678,1832,1857]" italics="true" pageId="5" pageNumber="560">tripartitus</emphasis>
</taxonomicName>
<emphasis id="B935E442B82BC621FD7BC0260A166846" bold="true" box="[685,776,1831,1857]" pageId="5" pageNumber="560">
<taxonomicNameLabel id="A2065939B82BC621FD7BC0260A166846" box="[685,776,1831,1857]" pageId="5" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Subcapitulum size: 97105 × 8690; subcapitular setae
<emphasis id="B935E442B82BC621FA51C0290C8B6846" box="[1415,1429,1832,1857]" italics="true" pageId="5" pageNumber="560">a</emphasis>
,
<emphasis id="B935E442B82BC621FF41C04D09B56862" box="[151,171,1868,1893]" italics="true" pageId="5" pageNumber="560">m</emphasis>
(2226) setiform, roughened;
<emphasis id="B935E442B82BC621FDDDC04A0B076862" box="[523,537,1867,1893]" italics="true" pageId="5" pageNumber="560">h</emphasis>
(3237) slightly thicker, setiform, barbed; both adoral setae (7) setiform, thin, smooth. Palp length: 7579; setation: 0-2-1-3-9 (+solenidion); postpalpal seta (6) spiniform, smooth. Chelicera length: 8690; teeth of both digits small but well developed; setae (
<emphasis id="B935E442B82BC621FC5CC0920AAC68AA" box="[906,946,1939,1965]" italics="true" pageId="5" pageNumber="560">cha</emphasis>
: 15;
<emphasis id="B935E442B82BC621FC3AC0920D0A68AA" box="[1004,1044,1939,1965]" italics="true" pageId="5" pageNumber="560">chb</emphasis>
: 19) setiform, thin, roughened.
</paragraph>
<paragraph id="8BFE3850B82BC622FF11C0B60BB96FD2" blockId="5.[151,1436,1831,2037]" lastBlockId="6.[151,1436,150,573]" lastPageId="6" lastPageNumber="561" pageId="5" pageNumber="560">
<emphasis id="B935E442B82BC621FF11C0B60B8F68D6" box="[199,657,1975,2001]" italics="true" pageId="5" pageNumber="560">Epimeral and lateral podosomal regions</emphasis>
(
<figureCitation id="137A24D5B82BC621FD77C0B60A3E68D6" box="[673,800,1975,2001]" captionStart="FIGURE 3" captionStartId="5.[151,250,1652,1676]" captionTargetBox="[165,1422,194,1621]" captionTargetId="figure-18@5.[151,1436,181,1629]" captionTargetPageId="5" captionText="FIGURE 3. Ceratorchestes (Ceratorchestes) duocornutus sp. nov., adult, male (A, B) and female (C, D): A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted); D—anogenital region. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13747448" httpUri="https://zenodo.org/record/13747448/files/figure.png" pageId="5" pageNumber="560">Figs 3B, C</figureCitation>
). Epimeral formula: 3-1-3-5;
<emphasis id="B935E442B82BC621FBA6C0B60D9268D6" box="[1136,1164,1975,2001]" italics="true" pageId="5" pageNumber="560">1b</emphasis>
,
<emphasis id="B935E442B82BC621FB4CC0B60DAB68D6" box="[1178,1205,1975,2001]" italics="true" pageId="5" pageNumber="560">1c</emphasis>
,
<emphasis id="B935E442B82BC621FB12C0B60DFE68D6" box="[1220,1248,1975,2001]" italics="true" pageId="5" pageNumber="560">3b</emphasis>
(4549) slightly thickened, stiff, acuminate, barbed;
<emphasis id="B935E442B82BC621FDFAC0DA0B5868F2" box="[556,582,2011,2037]" italics="true" pageId="5" pageNumber="560">3c</emphasis>
(2630),
<emphasis id="B935E442B82BC621FD60C0DA0BCD68F2" box="[694,723,2011,2037]" italics="true" pageId="5" pageNumber="560">4a</emphasis>
,
<emphasis id="B935E442B82BC621FD37C0DA0BE068F2" box="[737,766,2011,2037]" italics="true" pageId="5" pageNumber="560">4b</emphasis>
,
<emphasis id="B935E442B82BC621FCDAC0DA0A3968F2" box="[780,807,2011,2037]" italics="true" pageId="5" pageNumber="560">4c</emphasis>
(2226) setiform, slightly barbed;
<emphasis id="B935E442B82BC621FB65C0DA0DD168F2" box="[1203,1231,2011,2037]" italics="true" pageId="5" pageNumber="560">4d</emphasis>
(3437) setiform, barbed;
<emphasis id="B935E442B828C622FF25C79608116FB6" box="[243,271,151,177]" italics="true" pageId="6" pageNumber="561">1a</emphasis>
,
<emphasis id="B935E442B828C622FEC8C79608276FB6" box="[286,313,151,177]" italics="true" pageId="6" pageNumber="561">2a</emphasis>
,
<emphasis id="B935E442B828C622FE9EC796087A6FB6" box="[328,356,151,177]" italics="true" pageId="6" pageNumber="561">3a</emphasis>
(1315),
<emphasis id="B935E442B828C622FE02C79608F06FB6" box="[468,494,151,177]" italics="true" pageId="6" pageNumber="561">4e</emphasis>
(2226) setiform, thin, roughened;
<emphasis id="B935E442B828C622FC54C7960A806FB6" box="[898,926,151,177]" italics="true" pageId="6" pageNumber="561">3a</emphasis>
noticeably distanced from
<emphasis id="B935E442B828C622FB04C7960DF06FB6" box="[1234,1262,151,177]" italics="true" pageId="6" pageNumber="561">2a</emphasis>
and inserted in the same oblique row with setae of epimere IV.
</paragraph>
<paragraph id="8BFE3850B828C622FF11C7DE0A7A6E62" blockId="6.[151,1436,150,573]" pageId="6" pageNumber="561">
<emphasis id="B935E442B828C622FF11C7DE088F6FFF" box="[199,401,223,249]" italics="true" pageId="6" pageNumber="561">Anogenital region</emphasis>
(
<figureCitation id="137A24D5B828C622FE77C7DE0B3F6FFE" box="[417,545,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1652,1676]" captionTargetBox="[165,1422,194,1621]" captionTargetId="figure-18@5.[151,1436,181,1629]" captionTargetPageId="5" captionText="FIGURE 3. Ceratorchestes (Ceratorchestes) duocornutus sp. nov., adult, male (A, B) and female (C, D): A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted); D—anogenital region. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13747448" httpUri="https://zenodo.org/record/13747448/files/figure.png" pageId="6" pageNumber="561">Figs 3BD</figureCitation>
). Anogenital formula: 6-1-2-3; genital setae (
<emphasis id="B935E442B828C622FBC8C7DE0D326FFF" box="[1054,1068,223,248]" italics="true" pageId="6" pageNumber="561">g</emphasis>
<subScript id="17C53A15B828C622FBFAC7F00D2A6FF8" attach="left" box="[1068,1076,241,255]" fontSize="6" pageId="6" pageNumber="561">1</subScript>
: 1519;
<emphasis id="B935E442B828C622FB4FC7DE0DB96FFF" box="[1177,1191,223,248]" italics="true" pageId="6" pageNumber="561">g</emphasis>
<subScript id="17C53A15B828C622FB71C7F00DB16FF8" attach="left" box="[1191,1199,241,255]" fontSize="6" pageId="6" pageNumber="561">2</subScript>
<emphasis id="B935E442B828C622FB6BC7DE0DD56FFF" box="[1213,1227,223,248]" italics="true" pageId="6" pageNumber="561">g</emphasis>
<subScript id="17C53A15B828C622FB1DC7F00DCD6FF8" attach="left" box="[1227,1235,241,255]" fontSize="6" pageId="6" pageNumber="561">6</subScript>
: 1115) setiform, roughened; aggenital (3034), anal (2830) setae and adanal setae
<emphasis id="B935E442B828C622FCAFC6020A8B6E1A" box="[889,917,259,285]" italics="true" pageId="6" pageNumber="561">ad</emphasis>
<subScript id="17C53A15B828C622FC43C6140A836E24" attach="left" box="[917,925,277,291]" fontSize="6" pageId="6" pageNumber="561">2</subScript>
(3437),
<emphasis id="B935E442B828C622FBD0C6020D3C6E1A" box="[1030,1058,259,285]" italics="true" pageId="6" pageNumber="561">ad</emphasis>
<subScript id="17C53A15B828C622FBF4C6140D346E24" attach="left" box="[1058,1066,277,291]" fontSize="6" pageId="6" pageNumber="561">3</subScript>
(3034) setiform, barbed; adanal seta
<emphasis id="B935E442B828C622FF1AC62609F66E46" box="[204,232,295,321]" italics="true" pageId="6" pageNumber="561">ad</emphasis>
<subScript id="17C53A15B828C622FF3EC63809EE6E40" attach="left" box="[232,240,313,327]" fontSize="6" pageId="6" pageNumber="561">1</subScript>
(5256) slightly thickened, stiff, acuminate, barbed. Genital and anal plates in females noticeably larger than in males. Adanal lyrifissure close and oblique to anal plate.
</paragraph>
<paragraph id="8BFE3850B828C622FF11C66E0BB86E8E" blockId="6.[151,1436,150,573]" box="[199,678,367,393]" pageId="6" pageNumber="561">
<emphasis id="B935E442B828C622FF11C66E09E26E8F" box="[199,252,367,392]" italics="true" pageId="6" pageNumber="561">Legs</emphasis>
. Similar to
<taxonomicName id="4C4143D3B828C622FE56C66E0B586E8F" authorityName="Ermilov" authorityYear="2024" box="[384,582,367,393]" class="Arachnida" family="Ceratoppiidae" genus="Ceratorchestes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="tripartitus" status="sp. nov." subGenus="Ceratorchestes">
<emphasis id="B935E442B828C622FE56C66E08846E8E" box="[384,410,367,393]" italics="true" pageId="6" pageNumber="561">C.</emphasis>
(
<emphasis id="B935E442B828C622FE7CC66E08DA6E8E" box="[426,452,367,393]" italics="true" pageId="6" pageNumber="561">C.</emphasis>
)
<emphasis id="B935E442B828C622FE02C66E0B586E8F" box="[468,582,367,392]" italics="true" pageId="6" pageNumber="561">tripartitus</emphasis>
</taxonomicName>
<emphasis id="B935E442B828C622FD9BC66E0BB86E8E" bold="true" box="[589,678,367,393]" pageId="6" pageNumber="561">
<taxonomicNameLabel id="A2065939B828C622FD9BC66E0BB86E8E" box="[589,678,367,393]" pageId="6" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35B6BDBB828C622FF11C6920D406EF2" pageId="6" pageNumber="561" type="discussion">
<paragraph id="8BFE3850B828C622FF11C6920D406EF2" blockId="6.[151,1436,150,573]" pageId="6" pageNumber="561">
<emphasis id="B935E442B828C622FF11C69208256EAA" bold="true" box="[199,315,403,429]" pageId="6" pageNumber="561">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="4C4143D3B828C622FD47C6920AED6EAA" authority="Mahunka, 1983" authorityName="Mahunka" authorityYear="1983" box="[657,1011,403,429]" class="Arachnida" family="Ceratoppiidae" genus="Ceratorchestes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="baloghi" subGenus="Ceratorchestes">
<emphasis id="B935E442B828C622FD47C6920BB56EAA" box="[657,683,403,429]" italics="true" pageId="6" pageNumber="561">C.</emphasis>
(
<emphasis id="B935E442B828C622FD6DC6920BCB6EAA" box="[699,725,403,429]" italics="true" pageId="6" pageNumber="561">C.</emphasis>
)
<emphasis id="B935E442B828C622FD30C6920A226EAA" box="[742,828,403,429]" italics="true" pageId="6" pageNumber="561">baloghi</emphasis>
<bibRefCitation id="EFD045A1B828C622FC95C6920AED6EAA" author="Mahunka, S." box="[835,1011,403,429]" pageId="6" pageNumber="561" pagination="269 - 298" refId="ref7900" refString="Mahunka, S. (1983) Neue und interessante Milben aus dem Genfer Museum XLV. Oribatida Americana 6: Mexico II (Acari). Revue suisse de Zoologie, 90, 269 - 298. https: // doi. org / 10.5962 / bhl. part. 81976" type="journal article" year="1983">Mahunka, 1983</bibRefCitation>
</taxonomicName>
(see also
<bibRefCitation id="EFD045A1B828C622FBB2C6920C116EAA" author="Mahunka, S." box="[1124,1295,403,429]" pageId="6" pageNumber="561" pagination="179 - 193" refId="ref7849" refString="Mahunka, S. (1982) Neue und interessante Milben aus dem Genfer Museum XLIV. Oribatida Americana 5: Costa Rica (Acari). Archives des Sciences, 35, 179 - 193. https: // doi. org / 10.5962 / bhl. part. 82450" type="journal article" year="1982">Mahunka 1982</bibRefCitation>
) and
<taxonomicName id="4C4143D3B828C622FA99C69208B16ED6" authority="Mahunka, 1982" authorityName="Mahunka" authorityYear="1982" class="Arachnida" family="Ceratoppiidae" genus="Ceratorchestes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="cornutus" subGenus="Ceratorchestes">
<emphasis id="B935E442B828C622FA99C6920C766EAA" box="[1359,1384,403,429]" italics="true" pageId="6" pageNumber="561">C.</emphasis>
(
<emphasis id="B935E442B828C622FAAFC6920C8D6EAA" box="[1401,1427,403,429]" italics="true" pageId="6" pageNumber="561">C.</emphasis>
)
<emphasis id="B935E442B828C622FF41C6B609E76ED7" box="[151,249,439,464]" italics="true" pageId="6" pageNumber="561">cornutus</emphasis>
<bibRefCitation id="EFD045A1B828C622FED6C6B608B16ED6" author="Mahunka, S." box="[256,431,439,465]" pageId="6" pageNumber="561" pagination="179 - 193" refId="ref7849" refString="Mahunka, S. (1982) Neue und interessante Milben aus dem Genfer Museum XLIV. Oribatida Americana 5: Costa Rica (Acari). Archives des Sciences, 35, 179 - 193. https: // doi. org / 10.5962 / bhl. part. 82450" type="journal article" year="1982">
Mahunka,
<quantity id="4CB995B5B828C622FEA1C6B608B16ED6" box="[375,431,439,465]" metricMagnitude="1" metricUnit="m" metricValue="5.03428" pageId="6" pageNumber="561" unit="in" value="1982.0">1982</quantity>
</bibRefCitation>
</taxonomicName>
in the absence of the translamella, but differs from both by the morphology of the rostrum (tripartite versus pointed) and lamellar cusp (with strong lateral tooth versus truncate).
</paragraph>
</subSubSection>
<subSubSection id="C35B6BDBB828C622FF11C6FE08F46D3A" pageId="6" pageNumber="561" type="etymology">
<paragraph id="8BFE3850B828C622FF11C6FE08F46D3A" blockId="6.[151,1436,150,573]" pageId="6" pageNumber="561">
<emphasis id="B935E442B828C622FF11C6FE08506D1E" bold="true" box="[199,334,511,537]" pageId="6" pageNumber="561">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C4143D3B828C622FDECC6FE0BD86D1E" authorityName="Ermilov" authorityYear="2024" box="[570,710,511,537]" class="Arachnida" family="Ceratoppiidae" genus="Ceratorchestes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="duocornutus">
<emphasis id="B935E442B828C622FDECC6FE0BD86D1E" box="[570,710,511,537]" italics="true" pageId="6" pageNumber="561">duocornutus</emphasis>
</taxonomicName>
refers to the presence of one pair of the well-developed teeth on lateral parts of the prodorsum.
</paragraph>
</subSubSection>
</treatment>
</document>