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<document id="9B6E02DDEA23772196CEB26B5FEECBC7" ID-DOI="10.11646/zootaxa.5297.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7993190" ID-ZooBank="454B72E3-2659-4F3D-9C9A-503DBABEE9CE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1685602563737" checkinUser="plazi" docAuthor="Molineri, Carlos, Rodríguez, José Sebastián, Leiva, Marta &amp; Márquez, Javier Andrés" docDate="2023" docId="03E22F20A7712220FF1CF25AFA46FCBC" docLanguage="en" docName="zootaxa.5297.2.4.pdf" docOrigin="Zootaxa 5297 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5297.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Progomphus Selys 1854" docType="key" docVersion="3" lastPageNumber="258" masterDocId="FFDB5758A7622233FF8BF37BFFDBFFEB" masterDocTitle="Diagnoses and key for the larvae of Progomphus Selys, 1854 from Argentina (Anisoptera: Gomphidae), with first larval descriptions for P. aberrans Belle, 1973 and P. kimminsi Belle, 1973" masterLastPageNumber="259" masterPageNumber="239" pageNumber="258" updateTime="1685723757853" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="9969819BF56B915553CA97D380820E8D">Diagnoses and key for the larvae of Progomphus Selys, 1854 from Argentina (Anisoptera: Gomphidae), with first larval descriptions for P. aberrans Belle, 1973 and P. kimminsi Belle, 1973</mods:title>
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<mods:namePart id="66442B81EBFE92C1034EDE7199A44A5C">Molineri, Carlos</mods:namePart>
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<mods:namePart id="270D656B2EDA51A1E19E5E9E6BD7C909">Rodríguez, José Sebastián</mods:namePart>
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<mods:affiliation id="39B221F3949522F7BE4CCF42FD48FF38">Instituto de Biodiversidad Neotropical, CONICET-UNT (Consejo Nacional de Investigaciones Científicas y Técnicas-Universidad Nacional de Tucumán), Facultad de Ciencias Naturales e Instituto M. Lillo, Tucumán, Argentina. &amp; josephum @ hotmail. com; https: // orcid. org / 0000 - 0002 - 8917 - 2882</mods:affiliation>
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<mods:namePart id="F754C9F9926042BABB83E33007652220">Leiva, Marta</mods:namePart>
<mods:affiliation id="11AD4AEBFF0928B7107DD8C777734B97">Facultad de Ciencias Forestales (FCF), Universidad Nacional de Santiago del Estero (UNSE), Avenida Belgrano (Sur) 1912, Santiago del Estero, Argentina.</mods:affiliation>
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Key to the larvae of
<taxonomicName id="4C4BE5B5A7712220FE0DF25AFDFAFED7" ID-CoL="6WHQ" authorityName="Selys" authorityYear="1854" box="[390,545,289,316]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B93F4224A7712220FE0DF25AFDFAFED7" bold="true" box="[390,545,289,316]" italics="true" pageId="19" pageNumber="258">Progomphus</emphasis>
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from
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(four species remain unknown in this stage:
<taxonomicName id="4C4BE5B5A7712220FB71F259FA4EFED0" authorityName="Ris" authorityYear="1911" box="[1274,1429,289,316]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="auropictus">
<emphasis id="B93F4224A7712220FB71F259FA4EFED0" bold="true" box="[1274,1429,289,316]" italics="true" pageId="19" pageNumber="258">P. auropictus</emphasis>
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,
<taxonomicName id="4C4BE5B5A7712220FF1CF23DFEC7FE8B" authorityName="Belle" authorityYear="1973" box="[151,284,325,352]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B93F4224A7712220FF1CF23DFEC7FE8B" bold="true" box="[151,284,325,352]" italics="true" pageId="19" pageNumber="258">P. australis</emphasis>
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,
<taxonomicName id="4C4BE5B5A7712220FEA0F23DFE1DFE8B" authorityName="Ris" authorityYear="1911" box="[299,454,325,352]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="basistictus">
<emphasis id="B93F4224A7712220FEA0F23DFE1DFE8B" bold="true" box="[299,454,325,352]" italics="true" pageId="19" pageNumber="258">P. basistictus</emphasis>
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and
<taxonomicName id="4C4BE5B5A7712220FD88F23DFDA1FE8B" box="[515,634,325,352]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="costalis">
<emphasis id="B93F4224A7712220FD88F23DFDA1FE8B" bold="true" box="[515,634,325,352]" italics="true" pageId="19" pageNumber="258">P. costalis</emphasis>
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).
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1 Last antennomere (the fourth) short, spherical (
<figureCitation id="137082B3A7712220FD14F2F1FD36FE4B" box="[671,749,394,416]" captionStart="FIGURE 6" captionStartId="9.[152,255,1959,1983]" captionTargetBox="[207,1379,181,1935]" captionTargetId="figure-18@9.[207,1380,181,1935]" captionTargetPageId="9" captionText="FIGURE 6. P. joergenseni Ris, last instar larval exuvia: a, dorsal habitus (antennomeres are numbered, ts=tibial spur); b, abdomen, lateral; c, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993208" httpUri="https://zenodo.org/record/7993208/files/figure.png" pageId="19" pageNumber="258">Figs. 6a</figureCitation>
,
<figureCitation id="137082B3A7712220FD7DF2F1FCC0FE4B" box="[758,795,394,416]" captionStart="FIGURE 11" captionStartId="15.[152,255,1646,1670]" captionTargetBox="[204,1384,181,1623]" captionTargetId="figure-18@15.[204,1384,181,1623]" captionTargetPageId="15" captionText="FIGURE 11. P. lepidus Ris, mature larva, female: a, dorsal habitus; b, antennae (antennomeres are numbered); c, abdomen lateral; d, abdominal sterna S710 (numbers indicate plates on S8, dotted lines indicate lateral sutures); e, anal pyramid, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993219" httpUri="https://zenodo.org/record/7993219/files/figure.png" pageId="19" pageNumber="258">11b</figureCitation>
,
<figureCitation id="137082B3A7712220FCAFF2F1FC91FE4B" box="[804,842,394,416]" captionStart="FIGURE 13" captionStartId="18.[152,255,1943,1967]" captionTargetBox="[231,1356,181,1919]" captionTargetId="figure-18@18.[231,1356,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. P. phyllochromus Ris, last instar male larva: a, dorsal habitus; b, head and pronotum, dorsal (antennomeres are numbered); c, detail of tibial spurs (ts), ventral; d, abdomen, lateral; e, same, ventral; f, anal pyramid, dorsal; g, same, lateral; h, abdominal sterna S810 (numbers indicate plates on S8)." figureDoi="http://doi.org/10.5281/zenodo.7993223" httpUri="https://zenodo.org/record/7993223/files/figure.png" pageId="19" pageNumber="258">13b</figureCitation>
), pro- and mesotibiae with distal spur (
<figureCitation id="137082B3A7712220FB33F2F1FADDFE4B" box="[1208,1286,394,416]" captionStart="FIGURE 6" captionStartId="9.[152,255,1959,1983]" captionTargetBox="[207,1379,181,1935]" captionTargetId="figure-18@9.[207,1380,181,1935]" captionTargetPageId="9" captionText="FIGURE 6. P. joergenseni Ris, last instar larval exuvia: a, dorsal habitus (antennomeres are numbered, ts=tibial spur); b, abdomen, lateral; c, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993208" httpUri="https://zenodo.org/record/7993208/files/figure.png" pageId="19" pageNumber="258">Figs. 6a</figureCitation>
,
<figureCitation id="137082B3A7712220FA84F2F1FAEFFE74" box="[1295,1332,394,415]" captionStart="FIGURE 13" captionStartId="18.[152,255,1943,1967]" captionTargetBox="[231,1356,181,1919]" captionTargetId="figure-18@18.[231,1356,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. P. phyllochromus Ris, last instar male larva: a, dorsal habitus; b, head and pronotum, dorsal (antennomeres are numbered); c, detail of tibial spurs (ts), ventral; d, abdomen, lateral; e, same, ventral; f, anal pyramid, dorsal; g, same, lateral; h, abdominal sterna S810 (numbers indicate plates on S8)." figureDoi="http://doi.org/10.5281/zenodo.7993223" httpUri="https://zenodo.org/record/7993223/files/figure.png" pageId="19" pageNumber="258">13a</figureCitation>
), abdomen with posterolateral spines on S69........................................................................ 2
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- Last antennomere long (at least longer than antennomere1-2 combined) and slender, cylindrical (
<figureCitation id="137082B3A7712220FB13F2BEFB31FE30" box="[1176,1258,453,475]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[250,1337,181,1919]" captionTargetId="figure-18@4.[250,1337,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. P. aberrans Belle, female last instar larval exuvia: a, dorsal habitus; b, antennae, lateral (numbers indicate antennomeres); c, fore leg with cheliform claws; d, abdomen, dorsal; e, abdomen, lateral; f, abdominal S810, ventral (g= gonapophyses); g, anal pyramid, dorsal; h, same, lateral; i, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993196" httpUri="https://zenodo.org/record/7993196/files/figure.png" pageId="19" pageNumber="258">Figs. 2b</figureCitation>
,
<figureCitation id="137082B3A7712220FB7DF2BEFACBFE30" box="[1270,1296,453,475]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="19" pageNumber="258">4b</figureCitation>
,
<figureCitation id="137082B3A7712220FA97F2BEFAECFE30" box="[1308,1335,453,475]" captionStart="FIGURE 9" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[209,1378,181,1919]" captionTargetId="figure-18@13.[209,1378,181,1919]" captionTargetPageId="13" captionText="FIGURE 9. P. kimminsi Belle, last instar larval exuvia: a, dorsal habitus; b, detail of head, dorsal; c, detail of abdominal dorsal hooks (arrows); d, abdomen, lateral; e, anal pyramid, dorsal; f, same, lateral; g, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993214" httpUri="https://zenodo.org/record/7993214/files/figure.png" pageId="19" pageNumber="258">9b</figureCitation>
), pro- and mesotibiae without distal spur (
<figureCitation id="137082B3A7712220FD9BF299FD85FE1C" box="[528,606,482,504]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[250,1337,181,1919]" captionTargetId="figure-18@4.[250,1337,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. P. aberrans Belle, female last instar larval exuvia: a, dorsal habitus; b, antennae, lateral (numbers indicate antennomeres); c, fore leg with cheliform claws; d, abdomen, dorsal; e, abdomen, lateral; f, abdominal S810, ventral (g= gonapophyses); g, anal pyramid, dorsal; h, same, lateral; i, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993196" httpUri="https://zenodo.org/record/7993196/files/figure.png" pageId="19" pageNumber="258">Figs. 2c</figureCitation>
,
<figureCitation id="137082B3A7712220FDE2F299FD5AFE1C" box="[617,641,482,503]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="19" pageNumber="258">4c</figureCitation>
), abdomen with posterolateral spines at least on S59 (
<figureCitation id="137082B3A7712220FB19F299FB07FE13" box="[1170,1244,482,504]" captionStart="FIGURE 1" captionStartId="3.[152,255,1882,1906]" captionTargetBox="[191,1410,191,1856]" captionTargetId="figure-18@3.[170,1416,181,1859]" captionTargetPageId="3" captionText="FIGURE 1. P. aberrans Belle, last instar larval exuvia: a, left mandible, occlusal view; b, right mandible with detail, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, maxilar palp, dorsal view; e, detail of ligula; f, same (other specimen); g, prementum, dorsal view (om= outer margin of palpal segment); h, abdominal lateral outline (sterna omitted); i, abdominal ventral outline." figureDoi="http://doi.org/10.5281/zenodo.7993194" httpUri="https://zenodo.org/record/7993194/files/figure.png" pageId="19" pageNumber="258">Figs. 1i</figureCitation>
,
<figureCitation id="137082B3A7712220FB6CF299FADAFE13" box="[1255,1281,482,504]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="19" pageNumber="258">4d</figureCitation>
,
<figureCitation id="137082B3A7712220FA80F299FAF9FE1C" box="[1291,1314,482,503]" captionStart="FIGURE 9" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[209,1378,181,1919]" captionTargetId="figure-18@13.[209,1378,181,1919]" captionTargetPageId="13" captionText="FIGURE 9. P. kimminsi Belle, last instar larval exuvia: a, dorsal habitus; b, detail of head, dorsal; c, detail of abdominal dorsal hooks (arrows); d, abdomen, lateral; e, anal pyramid, dorsal; f, same, lateral; g, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993214" httpUri="https://zenodo.org/record/7993214/files/figure.png" pageId="19" pageNumber="258">9a</figureCitation>
)........ 4
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<keyLead id="30BA3D03A7712220FF1CF17BFA47FDFE" box="[151,1436,512,534]" pageId="19" pageNumber="258">
2 Sternum of S8 formed by 3 plates (
<figureCitation id="137082B3A7712220FDB8F17BFDACFDFD" box="[563,631,512,534]" captionStart="FIGURE 6" captionStartId="9.[152,255,1959,1983]" captionTargetBox="[207,1379,181,1935]" captionTargetId="figure-18@9.[207,1380,181,1935]" captionTargetPageId="9" captionText="FIGURE 6. P. joergenseni Ris, last instar larval exuvia: a, dorsal habitus (antennomeres are numbered, ts=tibial spur); b, abdomen, lateral; c, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993208" httpUri="https://zenodo.org/record/7993208/files/figure.png" pageId="19" pageNumber="258">Fig. 6c</figureCitation>
).......................................................
<taxonomicName id="4C4BE5B5A7712220FA9DF17BFA47FDFE" authorityName="Ris" authorityYear="1908" box="[1302,1436,512,533]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="joergenseni">
<emphasis id="B93F4224A7712220FA9DF17BFA47FDFE" box="[1302,1436,512,533]" italics="true" pageId="19" pageNumber="258">P. joergenseni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF49E36A7712220FF1CF166FA47FDD9" blockId="19.[151,1437,394,856]" box="[151,1436,541,563]" pageId="19" pageNumber="258">
<keyLead id="30BA3D03A7712220FF1CF166FA47FDD9" box="[151,1436,541,563]" pageId="19" pageNumber="258">
- Sternum of S8 formed by 5 plates (
<figureCitation id="137082B3A7712220FDB8F166FD55FDD8" box="[563,654,541,563]" captionStart="FIGURE 11" captionStartId="15.[152,255,1646,1670]" captionTargetBox="[204,1384,181,1623]" captionTargetId="figure-18@15.[204,1384,181,1623]" captionTargetPageId="15" captionText="FIGURE 11. P. lepidus Ris, mature larva, female: a, dorsal habitus; b, antennae (antennomeres are numbered); c, abdomen lateral; d, abdominal sterna S710 (numbers indicate plates on S8, dotted lines indicate lateral sutures); e, anal pyramid, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993219" httpUri="https://zenodo.org/record/7993219/files/figure.png" pageId="19" pageNumber="258">Figs. 11d</figureCitation>
,
<figureCitation id="137082B3A7712220FD13F166FD65FDD8" box="[664,702,541,563]" captionStart="FIGURE 13" captionStartId="18.[152,255,1943,1967]" captionTargetBox="[231,1356,181,1919]" captionTargetId="figure-18@18.[231,1356,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. P. phyllochromus Ris, last instar male larva: a, dorsal habitus; b, head and pronotum, dorsal (antennomeres are numbered); c, detail of tibial spurs (ts), ventral; d, abdomen, lateral; e, same, ventral; f, anal pyramid, dorsal; g, same, lateral; h, abdominal sterna S810 (numbers indicate plates on S8)." figureDoi="http://doi.org/10.5281/zenodo.7993223" httpUri="https://zenodo.org/record/7993223/files/figure.png" pageId="19" pageNumber="258">13h</figureCitation>
)........................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BF8693A7712220FF1CF141FA47FD61" pageId="19" pageNumber="258">
<paragraph id="8BF49E36A7712220FF1CF141FA47FDA4" blockId="19.[151,1437,394,856]" box="[151,1436,570,592]" pageId="19" pageNumber="258">
<keyLead id="30BA3D03A7712220FF1CF141FA47FDA4" box="[151,1436,570,592]" pageId="19" pageNumber="258">
3 Third antennomere flat and wide, subcircular in shape (
<figureCitation id="137082B3A7712220FD7DF141FC93FDBB" box="[758,840,570,592]" captionStart="FIGURE 11" captionStartId="15.[152,255,1646,1670]" captionTargetBox="[204,1384,181,1623]" captionTargetId="figure-18@15.[204,1384,181,1623]" captionTargetPageId="15" captionText="FIGURE 11. P. lepidus Ris, mature larva, female: a, dorsal habitus; b, antennae (antennomeres are numbered); c, abdomen lateral; d, abdominal sterna S710 (numbers indicate plates on S8, dotted lines indicate lateral sutures); e, anal pyramid, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993219" httpUri="https://zenodo.org/record/7993219/files/figure.png" pageId="19" pageNumber="258">Fig. 11b</figureCitation>
), ligula with 14 marginal tubercles (
<figureCitation id="137082B3A7712220FB28F141FB29FDBB" box="[1187,1266,570,592]" captionStart="FIGURE 10" captionStartId="14.[152,255,1915,1939]" captionTargetBox="[196,1390,852,1890]" captionTargetId="figure-248@14.[193,1394,849,1892]" captionTargetPageId="14" captionText="FIGURE 10. P. lepidus Ris, last instar female larva: a, left mandible, occlusal view; b, right mandible, occlusal view; c, same, dorso-occlusal view; d, maxilla, dorsdal view; e, maxila, ventral view (arrow indicates first tooth); f, detail of ligula; g, prementum, ventral view with detail of marginal microspines; h, subgenital lobe from pharate female." figureDoi="http://doi.org/10.5281/zenodo.7993217" httpUri="https://zenodo.org/record/7993217/files/figure.png" pageId="19" pageNumber="258">Fig. 10f</figureCitation>
).....
<taxonomicName id="4C4BE5B5A7712220FAB4F140FA47FDA4" authorityName="Ris" authorityYear="1911" box="[1343,1436,570,592]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="lepidus">
<emphasis id="B93F4224A7712220FAB4F140FA47FDA4" box="[1343,1436,570,592]" italics="true" pageId="19" pageNumber="258">P. lepidus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF49E36A7712220FF1CF123FA47FD61" blockId="19.[151,1437,394,856]" pageId="19" pageNumber="258">
<keyLead id="30BA3D03A7712220FF1CF123FA47FD61" pageId="19" pageNumber="258">
- Third antennomer subcylindrical, elongated (
<figureCitation id="137082B3A7712220FD18F123FD3DFD85" box="[659,742,600,622]" captionStart="FIGURE 13" captionStartId="18.[152,255,1943,1967]" captionTargetBox="[231,1356,181,1919]" captionTargetId="figure-18@18.[231,1356,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. P. phyllochromus Ris, last instar male larva: a, dorsal habitus; b, head and pronotum, dorsal (antennomeres are numbered); c, detail of tibial spurs (ts), ventral; d, abdomen, lateral; e, same, ventral; f, anal pyramid, dorsal; g, same, lateral; h, abdominal sterna S810 (numbers indicate plates on S8)." figureDoi="http://doi.org/10.5281/zenodo.7993223" httpUri="https://zenodo.org/record/7993223/files/figure.png" pageId="19" pageNumber="258">Fig. 13b</figureCitation>
), ligula with only 2 median marginal tubercles (
<figureCitation id="137082B3A7712220FB38F123FAD8FD86" box="[1203,1283,600,622]" captionStart="FIGURE 12" captionStartId="17.[152,255,1943,1967]" captionTargetBox="[183,1414,240,1917]" captionTargetId="figure-49@17.[171,1417,237,1920]" captionTargetPageId="17" captionText="FIGURE 12. P. phyllochromus Ris, last instar larval exuvia: a, left mandible, occlusal view; b, right mandible, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, same, dorsal view; e, ligula. detail of flat marginal setae; f, prementum, ventral view; g, detail of labial palp, ventral view; h, prementum with detail, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7993221" httpUri="https://zenodo.org/record/7993221/files/figure.png" pageId="19" pageNumber="258">Fig. 12e</figureCitation>
)..................................................................................................
<taxonomicName id="4C4BE5B5A7712220FB7CF10DFA47FD61" authorityName="Ris" authorityYear="1918" box="[1271,1436,629,651]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="phyllochromus">
<emphasis id="B93F4224A7712220FB7CF10DFA47FD61" box="[1271,1436,629,651]" italics="true" pageId="19" pageNumber="258">P. phyllochromus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BF8693A7712220FF1CF1E9FA47FCEB" pageId="19" pageNumber="258">
<paragraph id="8BF49E36A7712220FF1CF1E9FA47FD2E" blockId="19.[151,1437,394,856]" pageId="19" pageNumber="258">
<keyLead id="30BA3D03A7712220FF1CF1E9FA47FD2E" pageId="19" pageNumber="258">
4 Anal pyramid short (epiproct subequal in length to S10,
<figureCitation id="137082B3A7712220FD74F1E9FCBDFD43" box="[767,870,658,680]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="19" pageNumber="258">Figs. 4de</figureCitation>
), posterolateral spine absent on S4 (
<figureCitation id="137082B3A7712220FB4BF1E9FADCFD43" box="[1216,1287,658,680]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[274,1313,181,1919]" captionTargetId="figure-18@6.[274,1313,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. P. complicatus Selys, female last instar larval exuvia: a, dorsal habitus; b, antennae, dorsal (number indicate 4th antennomere); c, “normal” fore tarsal claws, lateral; d, abdomen, ventral; e, abdomen, lateral." figureDoi="http://doi.org/10.5281/zenodo.7993202" httpUri="https://zenodo.org/record/7993202/files/figure.png" pageId="19" pageNumber="258">Fig. 4d</figureCitation>
)...................................................................................................
<taxonomicName id="4C4BE5B5A7712220FA9AF1CBFA47FD2E" authorityName="Selys" authorityYear="1854" box="[1297,1436,688,709]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="B93F4224A7712220FA9AF1CBFA47FD2E" box="[1297,1436,688,709]" italics="true" pageId="19" pageNumber="258">P. complicatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF49E36A7712220FF1CF1B6FA47FCEB" blockId="19.[151,1437,394,856]" pageId="19" pageNumber="258">
<keyLead id="30BA3D03A7712220FF1CF1B6FA47FCEB" pageId="19" pageNumber="258">
- Anal pyramid long and slender (epiproct as long as or longer than S9 and 10 combined,
<figureCitation id="137082B3A7712220FBCAF1B6FB11FD08" box="[1089,1226,717,739]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[250,1337,181,1919]" captionTargetId="figure-18@4.[250,1337,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. P. aberrans Belle, female last instar larval exuvia: a, dorsal habitus; b, antennae, lateral (numbers indicate antennomeres); c, fore leg with cheliform claws; d, abdomen, dorsal; e, abdomen, lateral; f, abdominal S810, ventral (g= gonapophyses); g, anal pyramid, dorsal; h, same, lateral; i, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993196" httpUri="https://zenodo.org/record/7993196/files/figure.png" pageId="19" pageNumber="258">
Figs. 2d,
<quantity id="4CB333D3A7712220FB16F1B6FB6EFD08" box="[1181,1205,717,739]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="19" pageNumber="258" unit="g" value="2.0">2g</quantity>
i
</figureCitation>
,
<figureCitation id="137082B3A7712220FB5EF1B6FADCFD08" box="[1237,1287,717,739]" captionStart="FIGURE 9" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[209,1378,181,1919]" captionTargetId="figure-18@13.[209,1378,181,1919]" captionTargetPageId="13" captionText="FIGURE 9. P. kimminsi Belle, last instar larval exuvia: a, dorsal habitus; b, detail of head, dorsal; c, detail of abdominal dorsal hooks (arrows); d, abdomen, lateral; e, anal pyramid, dorsal; f, same, lateral; g, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993214" httpUri="https://zenodo.org/record/7993214/files/figure.png" pageId="19" pageNumber="258">9dg</figureCitation>
), posterolateral spine present on S4 (
<figureCitation id="137082B3A7712220FE21F191FE2FFCEB" box="[426,500,746,768]" captionStart="FIGURE 1" captionStartId="3.[152,255,1882,1906]" captionTargetBox="[191,1410,191,1856]" captionTargetId="figure-18@3.[170,1416,181,1859]" captionTargetPageId="3" captionText="FIGURE 1. P. aberrans Belle, last instar larval exuvia: a, left mandible, occlusal view; b, right mandible with detail, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, maxilar palp, dorsal view; e, detail of ligula; f, same (other specimen); g, prementum, dorsal view (om= outer margin of palpal segment); h, abdominal lateral outline (sterna omitted); i, abdominal ventral outline." figureDoi="http://doi.org/10.5281/zenodo.7993194" httpUri="https://zenodo.org/record/7993194/files/figure.png" pageId="19" pageNumber="258">Figs. 1i</figureCitation>
,
<figureCitation id="137082B3A7712220FE74F191FDCCFD14" box="[511,535,746,767]" captionStart="FIGURE 8" captionStartId="12.[152,255,1222,1246]" captionTargetBox="[174,1414,184,1195]" captionTargetId="figure-18@12.[171,1417,181,1198]" captionTargetPageId="12" captionText="FIGURE 8. P. kimminsi Belle, last instar larval exuvia: a, lateral abdominal outline, detail of tubercles on segments 13; b, lateral abdominal outline; c, ventral abdominal outline (arrow indicate spine present on S4)." figureDoi="http://doi.org/10.5281/zenodo.7993212" httpUri="https://zenodo.org/record/7993212/files/figure.png" pageId="19" pageNumber="258">8c</figureCitation>
)......................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BF8693A7712220FF1CF07CFA46FCBC" pageId="19" pageNumber="258">
<paragraph id="8BF49E36A7712220FF1CF07CFA46FCD1" blockId="19.[151,1437,394,856]" pageId="19" pageNumber="258">
<keyLead id="30BA3D03A7712220FF1CF07CFA46FCD1" pageId="19" pageNumber="258">
5 Small abdominal dorsal tubercles present on S19, sometimes very reduced on S5 to 8 (
<figureCitation id="137082B3A7712220FBB3F073FB53FCF5" box="[1080,1160,776,798]" captionStart="FIGURE 1" captionStartId="3.[152,255,1882,1906]" captionTargetBox="[191,1410,191,1856]" captionTargetId="figure-18@3.[170,1416,181,1859]" captionTargetPageId="3" captionText="FIGURE 1. P. aberrans Belle, last instar larval exuvia: a, left mandible, occlusal view; b, right mandible with detail, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, maxilar palp, dorsal view; e, detail of ligula; f, same (other specimen); g, prementum, dorsal view (om= outer margin of palpal segment); h, abdominal lateral outline (sterna omitted); i, abdominal ventral outline." figureDoi="http://doi.org/10.5281/zenodo.7993194" httpUri="https://zenodo.org/record/7993194/files/figure.png" pageId="19" pageNumber="258">Figs. 1h</figureCitation>
,
<figureCitation id="137082B3A7712220FB1FF073FB1EFCF5" box="[1172,1221,776,798]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[250,1337,181,1919]" captionTargetId="figure-18@4.[250,1337,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. P. aberrans Belle, female last instar larval exuvia: a, dorsal habitus; b, antennae, lateral (numbers indicate antennomeres); c, fore leg with cheliform claws; d, abdomen, dorsal; e, abdomen, lateral; f, abdominal S810, ventral (g= gonapophyses); g, anal pyramid, dorsal; h, same, lateral; i, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993196" httpUri="https://zenodo.org/record/7993196/files/figure.png" pageId="19" pageNumber="258">2de</figureCitation>
), posterolateral spines on S29 (
<figureCitation id="137082B3A7712220FEB5F05EFE53FCD0" box="[318,392,805,827]" captionStart="FIGURE 1" captionStartId="3.[152,255,1882,1906]" captionTargetBox="[191,1410,191,1856]" captionTargetId="figure-18@3.[170,1416,181,1859]" captionTargetPageId="3" captionText="FIGURE 1. P. aberrans Belle, last instar larval exuvia: a, left mandible, occlusal view; b, right mandible with detail, occlusal view; c, maxilla, ventral view (arrow indicates first tooth); d, maxilar palp, dorsal view; e, detail of ligula; f, same (other specimen); g, prementum, dorsal view (om= outer margin of palpal segment); h, abdominal lateral outline (sterna omitted); i, abdominal ventral outline." figureDoi="http://doi.org/10.5281/zenodo.7993194" httpUri="https://zenodo.org/record/7993194/files/figure.png" pageId="19" pageNumber="258">Figs. 1i</figureCitation>
,
<figureCitation id="137082B3A7712220FE18F05EFE77FCD0" box="[403,428,805,827]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[250,1337,181,1919]" captionTargetId="figure-18@4.[250,1337,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. P. aberrans Belle, female last instar larval exuvia: a, dorsal habitus; b, antennae, lateral (numbers indicate antennomeres); c, fore leg with cheliform claws; d, abdomen, dorsal; e, abdomen, lateral; f, abdominal S810, ventral (g= gonapophyses); g, anal pyramid, dorsal; h, same, lateral; i, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993196" httpUri="https://zenodo.org/record/7993196/files/figure.png" pageId="19" pageNumber="258">2d</figureCitation>
)..........................................................................
<taxonomicName id="4C4BE5B5A7712220FAA5F05DFA46FCD1" authorityName="Belle" authorityYear="1973" box="[1326,1437,805,827]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="aberrans">
<emphasis id="B93F4224A7712220FAA5F05DFA46FCD1" box="[1326,1437,805,827]" italics="true" pageId="19" pageNumber="258">P. aberrans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF49E36A7712220FF1CF038FA46FCBC" blockId="19.[151,1437,394,856]" box="[151,1437,834,856]" pageId="19" pageNumber="258">
<keyLead id="30BA3D03A7712220FF1CF038FA46FCBC" box="[151,1437,834,856]" pageId="19" pageNumber="258">
- Small abdominal dorsal tubercles present on S13 (
<figureCitation id="137082B3A7712220FD5BF039FCECFCB3" box="[720,823,834,856]" captionStart="FIGURE 8" captionStartId="12.[152,255,1222,1246]" captionTargetBox="[174,1414,184,1195]" captionTargetId="figure-18@12.[171,1417,181,1198]" captionTargetPageId="12" captionText="FIGURE 8. P. kimminsi Belle, last instar larval exuvia: a, lateral abdominal outline, detail of tubercles on segments 13; b, lateral abdominal outline; c, ventral abdominal outline (arrow indicate spine present on S4)." figureDoi="http://doi.org/10.5281/zenodo.7993212" httpUri="https://zenodo.org/record/7993212/files/figure.png" pageId="19" pageNumber="258">Figs. 8ab</figureCitation>
,
<figureCitation id="137082B3A7712220FCCAF039FC81FCBC" box="[833,858,834,855]" captionStart="FIGURE 9" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[209,1378,181,1919]" captionTargetId="figure-18@13.[209,1378,181,1919]" captionTargetPageId="13" captionText="FIGURE 9. P. kimminsi Belle, last instar larval exuvia: a, dorsal habitus; b, detail of head, dorsal; c, detail of abdominal dorsal hooks (arrows); d, abdomen, lateral; e, anal pyramid, dorsal; f, same, lateral; g, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993214" httpUri="https://zenodo.org/record/7993214/files/figure.png" pageId="19" pageNumber="258">9c</figureCitation>
), posterolateral spines on S49 (
<figureCitation id="137082B3A7712220FB18F039FB3AFCBC" box="[1171,1249,834,856]" captionStart="FIGURE 8" captionStartId="12.[152,255,1222,1246]" captionTargetBox="[174,1414,184,1195]" captionTargetId="figure-18@12.[171,1417,181,1198]" captionTargetPageId="12" captionText="FIGURE 8. P. kimminsi Belle, last instar larval exuvia: a, lateral abdominal outline, detail of tubercles on segments 13; b, lateral abdominal outline; c, ventral abdominal outline (arrow indicate spine present on S4)." figureDoi="http://doi.org/10.5281/zenodo.7993212" httpUri="https://zenodo.org/record/7993212/files/figure.png" pageId="19" pageNumber="258">Figs. 8c</figureCitation>
,
<figureCitation id="137082B3A7712220FB67F039FAD8FCBC" box="[1260,1283,834,855]" captionStart="FIGURE 9" captionStartId="13.[152,255,1943,1967]" captionTargetBox="[209,1378,181,1919]" captionTargetId="figure-18@13.[209,1378,181,1919]" captionTargetPageId="13" captionText="FIGURE 9. P. kimminsi Belle, last instar larval exuvia: a, dorsal habitus; b, detail of head, dorsal; c, detail of abdominal dorsal hooks (arrows); d, abdomen, lateral; e, anal pyramid, dorsal; f, same, lateral; g, same, ventral." figureDoi="http://doi.org/10.5281/zenodo.7993214" httpUri="https://zenodo.org/record/7993214/files/figure.png" pageId="19" pageNumber="258">9a</figureCitation>
)...
<taxonomicName id="4C4BE5B5A7712220FAA5F038FA46FCBC" authorityName="Belle" authorityYear="1973" box="[1326,1437,834,856]" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="258" phylum="Arthropoda" rank="species" species="kimminsi">
<emphasis id="B93F4224A7712220FAA5F038FA46FCBC" box="[1326,1437,834,856]" italics="true" pageId="19" pageNumber="258">P. kimminsi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>