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<document id="B9A72B3D3ED31238E30DA37AC0F60E4E" ID-DOI="10.5281/zenodo.3740269" ID-GBIF-Dataset="34e4a5a6-881c-4a1a-b64d-6868e12df7fb" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3740269" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1586037971521" checkinUser="plazi" docAuthor="Bonaccorso, Frank" docDate="2019" docId="03D587F2FFC74C0CFF1632ECFD40EE5E" docLanguage="en" docName="hbmw_9_Emballorunidae.pdf.imd" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docStyle="DocumentStyle{}" docTitle="Saccolaimus mixtus Troughton 1925" docType="treatment" docVersion="19" lastPageNumber="357" masterDocId="FFECFF8AFFCF4C04FFA53577FFF8FFE9" masterDocTitle="Emballonuridae" masterLastPageNumber="373" masterPageNumber="350" pageNumber="357" updateTime="1698735528461" updateUser="plazi">
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<mods:title id="A52C2D97A2A1DE379D31A1CDB8D1D9C9">Emballonuridae</mods:title>
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<mods:namePart id="43DB8B7EC93AC47AEB49499EA3C2FDFE">Bonaccorso, Frank</mods:namePart>
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<mods:namePart id="DB4C8DC18AAD0A78B59646BB8CE99FCB">Wilson, Don E.</mods:namePart>
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<mods:namePart id="DFDE23B8FD5BE018BFFE0CB347215B56">Mittermeier, Russel A.</mods:namePart>
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<mods:title id="6C4AF39954D9D05C4B0CE113A843B1FD">Handbook of the Mammals of the World Volume 9 Bats</mods:title>
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<treatment id="03D587F2FFC74C0CFF1632ECFD40EE5E" ID-DOI="http://doi.org/10.5281/zenodo.3810727" ID-GBIF-Taxon="163435157" ID-Zenodo-Dep="3810727" LSID="urn:lsid:plazi:treatment:03D587F2FFC74C0CFF1632ECFD40EE5E" httpUri="http://treatment.plazi.org/id/03D587F2FFC74C0CFF1632ECFD40EE5E" lastPageNumber="357" pageId="8" pageNumber="357">
<subSubSection id="C366656FFFC74C0CFF1632ECFC21F831" box="[179,985,1947,2008]" pageId="8" pageNumber="357" type="vernacular_names">
<paragraph id="8BC336E4FFC74C0CFF1632ECFC21F831" blockId="8.[179,1549,1947,2008]" box="[179,985,1947,2008]" pageId="8" pageNumber="357">
<figureCitation id="13472A61FFC74C0CFF1632ECFF0EF831" box="[179,246,1947,2008]" captionStart="On" captionStartId="2.[173,216,4419,4446]" captionTargetBox="[177,3496,132,4697]" captionTargetPageId="1" captionText="On following pages: 4. Hamilton'sTomb Bat [Taphozous hamiltoni); 5. Hildegarde'sTomb Bat (Taphozous hildegardeae}; 6. Long-winged Sheath-tailed Bat (Taphozous longimanus); 7. Black-beardedTomb Bat (Taphozous melanopogoh); 8. Theobald'sTomb Bat (Taphozous theobaldi); 9. Indonesian Tomb Bat (Taphozous achates); 10. Arnhem Sheath-tailed Bat (Taphozous kapalgensis}; 11. Coastal Sheath-tailed Bat (Taphozous australis}; 12. Common Sheath-tailed Bat (Taphozous georgianus); 13. Hill's Sheath-tailed Bat (Taphozous hilli); 14. Troughton's Sheath-tailed Bat (Taphozous troughtoni); 15. Pel's Pouched Bat (Saccolaimus peli); 16. Bare-rumped Sheathtailed Bat (Saccolaimus saccolaimus); 17. Papuan Sheath-tailed Bat (Saccolaimus mixtus); 18. Yellow-bellied Sheath-tailed Bat (Saccolaimus flaviventris)." figureDoi="http://doi.org/10.5281/zenodo.3740271" httpUri="https://zenodo.org/record/3740271/files/figure.png" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF1632ECFF0EF831" bold="true" box="[179,246,1947,2008]" pageId="8" pageNumber="357">17</emphasis>
</figureCitation>
.
<emphasis id="B908EAF6FFC74C0CFEB132ECFC21F831" box="[276,985,1947,2008]" pageId="8" pageNumber="357">Papuan Sheath-tailed Bat</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFBB832ECF9F5F831" box="[1053,1549,1947,2008]" pageId="8" pageNumber="357" type="nomenclature">
<paragraph id="8BC336E4FFC74C0CFBB832ECF9F5F831" blockId="8.[179,1549,1947,2008]" box="[1053,1549,1947,2008]" pageId="8" pageNumber="357">
<taxonomicName id="4C7C4D67FFC74C0CFBB832ECF9F5F831" ID-CoL="4TX5D" authorityName="Troughton" authorityYear="1925" box="[1053,1549,1947,2008]" class="Mammalia" family="Emballonuridae" genus="Saccolaimus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="6" phylum="Chordata" rank="species" species="mixtus">
<emphasis id="B908EAF6FFC74C0CFBB832ECF9F5F831" bold="true" box="[1053,1549,1947,2008]" italics="true" pageId="8" pageNumber="357">Saccolaimus mixtus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFF0A3299FC80F7D3" pageId="8" pageNumber="357" type="vernacular_names">
<paragraph id="8BC336E4FFC74C0CFF0A3299FA7AF7EC" blockId="8.[175,1410,2026,2106]" box="[175,1410,2026,2053]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A3299FEECF7ED" bold="true" box="[175,276,2030,2052]" italics="true" pageId="8" pageNumber="357">French:</emphasis>
Taphien papou /
<emphasis id="B908EAF6FFC74C0CFE5F3299FD8DF7ED" bold="true" box="[506,629,2030,2052]" pageId="8" pageNumber="357">German:</emphasis>
Papua-Glattnasenfreischwanz /
<emphasis id="B908EAF6FFC74C0CFB803299FB59F7ED" bold="true" box="[1061,1185,2030,2052]" pageId="8" pageNumber="357">Spanish:</emphasis>
<emphasis id="B908EAF6FFC74C0CFB02329DFB06F7EC" box="[1191,1278,2026,2053]" italics="true" pageId="8" pageNumber="357">Tafozo</emphasis>
de Papua
</paragraph>
<paragraph id="8BC336E4FFC74C0CFF0A3D54FC80F7D3" blockId="8.[175,1410,2026,2106]" box="[175,888,2079,2106]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A3D54FE03F7D0" bold="true" box="[175,507,2083,2105]" pageId="8" pageNumber="357">Other common names:</emphasis>
Cape
<emphasis id="B908EAF6FFC74C0CFDF53D68FD71F7D3" box="[592,649,2079,2106]" italics="true" pageId="8" pageNumber="357">York</emphasis>
Sheath-tailed Bat
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFC403DF4FBB3F737" pageId="8" pageNumber="357" type="reference_group">
<paragraph id="8BC336E4FFC74C0CFC403DF4FBB3F737" blockId="8.[997,1791,2179,2746]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFC403DF4FB48F743" box="[997,1200,2179,2218]" pageId="8" pageNumber="357">Taxonomy</emphasis>
.
<taxonomicName id="4C7C4D67FFC74C0CFB6F3DF4FBBBF737" ID-CoL="4TX5D" authority="Troughton, 1925" authorityName="Troughton" authorityYear="1925" class="Mammalia" family="Emballonuridae" genus="Saccolaimus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="357" phylum="Chordata" rank="species" species="mixtus">
<emphasis id="B908EAF6FFC74C0CFB6F3DF4F9E3F743" box="[1226,1563,2179,2218]" italics="true" pageId="8" pageNumber="357">Saccolaimus mixtus</emphasis>
Troughton, 1925
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFBFD3DC0F922F737" box="[1112,1754,2231,2270]" pageId="8" pageNumber="357" type="materials_examined">
<paragraph id="8BC336E4FFC74C0CFBFD3DC0F922F737" blockId="8.[997,1791,2179,2746]" box="[1112,1754,2231,2270]" pageId="8" pageNumber="357">
<materialsCitation id="3B143CB9FFC74C0CFBFD3DC0F92CF737" ID-GBIF-Occurrence="2595781797" box="[1112,1748,2231,2270]" country="Papua New Guinea" location="Port Moresby" pageId="8" pageNumber="357" specimenCount="1" stateProvince="Port Moresby" typeStatus="type">
<collectingRegion id="49B8F806FFC74C0CFBFD3DC0FAACF737" box="[1112,1364,2231,2270]" country="Papua New Guinea" name="National Capital" pageId="8" pageNumber="357">Port Moresby</collectingRegion>
,
<collectingCountry id="F36B7674FFC74C0CFACD3DC0F92CF737" box="[1384,1748,2231,2270]" name="Papua New Guinea" pageId="8" pageNumber="357">Papua New Guinea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFC433D9BFA35F6FA" box="[998,1485,2284,2323]" pageId="8" pageNumber="357" type="discussion">
<paragraph id="8BC336E4FFC74C0CFC433D9BFA35F6FA" blockId="8.[997,1791,2179,2746]" box="[998,1485,2284,2323]" pageId="8" pageNumber="357">This species is monotypic.</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFC433C56FB47F5F5" pageId="8" pageNumber="357" type="distribution">
<caption id="DF03666CFFC74C0CFC433C56FB47F5F5" ID-DOI="http://doi.org/10.5281/zenodo.3747936" ID-Zenodo-Dep="3747936" httpUri="https://zenodo.org/record/3747936/files/figure.png" inLine="true" pageId="8" pageNumber="357" targetBox="[174,967,2182,2739]" targetPageId="8">
<paragraph id="8BC336E4FFC74C0CFC433C56FB47F5F5" blockId="8.[997,1791,2179,2746]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFC433C56FB2BF6A1" bold="true" box="[998,1235,2337,2376]" pageId="8" pageNumber="357">Distribution.</emphasis>
S Papua New Guinea (S coastal region in
<emphasis id="B908EAF6FFC74C0CFAEC3C21F959F694" box="[1353,1697,2390,2429]" italics="true" pageId="8" pageNumber="357">Western Province</emphasis>
and National Capital District) and N Australia (N Cape
<emphasis id="B908EAF6FFC74C0CFB2A3CB7FB11F60E" box="[1167,1257,2496,2535]" italics="true" pageId="8" pageNumber="357">York</emphasis>
Peninsula); possibly on Torres Strait Is.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFC403F5EF936F41F" pageId="8" pageNumber="357" type="description">
<paragraph id="8BC336E4FFC74C0CFC403F5EF936F41F" blockId="8.[997,1791,2179,2746]" lastBlockId="8.[171,1790,2756,4535]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFC403F5EFB43F5B9" bold="true" box="[997,1211,2601,2640]" pageId="8" pageNumber="357">Descriptive</emphasis>
notes. Head-body 72-79 mm, tail 22-29 mm, ear 17-20 mm, hindfoot 13—15 mm, forearm 61-68 mm; weight 21-27 g. Dorsum of the Papuan Sheath-tailed Bat is dark gray-brown, with some whitetipped hairs; venter is white or pale buff-brown. Gular pouch is well developed in males and poorly developed in females. Radio-metacarpal sacs are present and lined with white fur in both sexes. Rump is well haired. Flight membranes, face, ears, tail, forearm, and feet are black. Eyes large. Ears are
<emphasis id="B908EAF6FFC74C0CFBB73EEDFB5AF428" box="[1042,1186,2970,3009]" italics="true" pageId="8" pageNumber="357">broadly</emphasis>
rounded and barely rise above crown of head. Strong, long calcars support trailing edges of uropatagium in flight.
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFF0A3973FCC6F0D6" pageId="8" pageNumber="357" type="biology_ecology">
<paragraph id="8BC336E4FFC74C0CFF0A3973FDC6F37D" blockId="8.[171,1790,2756,4535]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A3973FEBDF3C2" bold="true" box="[175,325,3076,3115]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A3973FEC6F3C2" box="[175,318,3076,3115]" pageId="8" pageNumber="357">Habitat</emphasis>
.
</emphasis>
Open eucalypt woodlands, sclerophyll forests, and along riparian systems. The Papuan Sheath-tailed Bat is common in larger tracts of Eucalyptus
<emphasis id="B908EAF6FFC74C0CFA7D394FF979F3B6" box="[1496,1665,3128,3167]" italics="true" pageId="8" pageNumber="357">tetrodonta</emphasis>
(
<emphasis id="B908EAF6FFC74C0CF93C394FF90BF3B6" bold="true" box="[1689,1779,3128,3167]" pageId="8" pageNumber="357">Myrt</emphasis>
aceae) on Cape
<emphasis id="B908EAF6FFC74C0CFE7B391AFDC1F37D" box="[478,569,3181,3220]" italics="true" pageId="8" pageNumber="357">York</emphasis>
.
</paragraph>
<paragraph id="8BC336E4FFC74C0CFF0A39D5FDD5F317" blockId="8.[171,1790,2756,4535]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A39D5FDECF320" bold="true" box="[175,532,3234,3273]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A39D5FDF4F320" box="[175,524,3234,3273]" pageId="8" pageNumber="357">Food and Feeding</emphasis>
.
</emphasis>
The Papuan Sheath-tailed Bat forages for insects over and underneath tree canopies.
</paragraph>
<paragraph id="8BC336E4FFC74C0CFF15387CFD5CF2DB" blockId="8.[171,1790,2756,4535]" box="[176,676,3339,3378]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF15387CFEA5F2DB" box="[176,349,3339,3378]" pageId="8" pageNumber="357">Breeding</emphasis>
. No information.
</paragraph>
<paragraph id="8BC336E4FFC74C0CFF083837FA70F131" blockId="8.[171,1790,2756,4535]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF083837FE11F28E" bold="true" box="[173,489,3392,3431]" pageId="8" pageNumber="357">Activity patterns.</emphasis>
The Papuan Sheath-tailed Bat is crepuscular. It will roost in limestone caves in Papua New Guinea but more
<emphasis id="B908EAF6FFC74C0CFCDA3802FBBCF275" box="[895,1092,3445,3484]" italics="true" pageId="8" pageNumber="357">frequently</emphasis>
shelters in tree hollows. In Australia, it often roosts in dead snags of
<emphasis id="B908EAF6FFC74C0CFCA638DDFC10F238" box="[771,1000,3498,3537]" italics="true" pageId="8" pageNumber="357">
<taxonomicName id="4C7C4D67FFC74C0CFCA638DDFC1BF238" box="[771,995,3498,3537]" class="Mammalia" family="Emballonuridae" genus="Emballonura" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="357" phylum="Chordata" rank="species" species="tetrodonta">E. tetrodonta</taxonomicName>
.
</emphasis>
It emerges at dusk to forage on aerial insects above sclerophyll forest canopies or high over rivers and canyons. Low-frequency, long-duration echolocation calls of the Papuan Sheath-tailed Bat are
<emphasis id="B908EAF6FFC74C0CFA4F3B64F9C3F1D3" box="[1514,1595,3603,3642]" italics="true" pageId="8" pageNumber="357">very</emphasis>
similar to those of the Bare-rumped Sheath-tailed Bat (
<taxonomicName id="4C7C4D67FFC74C0CFC5B3B3FFB0DF186" authorityName="Temminck" authorityYear="1838" box="[1022,1269,3656,3695]" class="Mammalia" family="Emballonuridae" genus="Taphozous" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="357" phylum="Chordata" rank="species" species="saccolaimus">
<emphasis id="B908EAF6FFC74C0CFC5B3B3FFBEFF186" box="[1022,1047,3656,3695]" italics="true" pageId="8" pageNumber="357">S</emphasis>
.
<emphasis id="B908EAF6FFC74C0CFB8E3B3FFB0DF186" box="[1067,1269,3656,3695]" italics="true" pageId="8" pageNumber="357">saccolaimus</emphasis>
</taxonomicName>
), but they lack sharp downward
<emphasis id="B908EAF6FFC74C0CFEB03B0AFE2FF14D" box="[277,471,3709,3748]" italics="true" pageId="8" pageNumber="357">frequency</emphasis>
sweep near ends of calls of the latter. Multiharmonic call
<emphasis id="B908EAF6FFC74C0CF9B53B0AF942F14D" box="[1552,1722,3709,3748]" italics="true" pageId="8" pageNumber="357">pulses of</emphasis>
the Papuan Sheath-tailed Bat have characteristic frequency of 21 kHz.
</paragraph>
<paragraph id="8BC336E4FFC74C0CFF0A3B91FCC6F0D6" blockId="8.[171,1790,2756,4535]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A3B91FB69F0E4" bold="true" box="[175,1169,3814,3853]" pageId="8" pageNumber="357">Movements, Home range and Social organization.</emphasis>
The Papuan Sheath-tailed Bat roosts in groups of 2—7 individuals.
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFF0A3A38F95BEEF5" pageId="8" pageNumber="357" type="conservation">
<paragraph id="8BC336E4FFC74C0CFF0A3A38F95BEEF5" blockId="8.[171,1790,2756,4535]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A3A38FD84F09F" bold="true" box="[175,636,3919,3958]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0A3A38FD8DF09F" box="[175,629,3919,3958]" pageId="8" pageNumber="357">Status and Conservation</emphasis>
.
</emphasis>
Classified as Near Threatened on 77
<emphasis id="B908EAF6FFC74C0CFABA3A38FAC4F09F" box="[1311,1340,3919,3958]" italics="true" pageId="8" pageNumber="357">ze</emphasis>
IUCNRed
<emphasis id="B908EAF6FFC74C0CFA593A38F9BCF09F" box="[1532,1604,3919,3958]" italics="true" pageId="8" pageNumber="357">List.</emphasis>
Numbers ofreproductively mature Papuan Sheath-tailed Bats have declined in some regional populations. Threats include large-scale removal of tall forests for bauxite mining on Cape
<emphasis id="B908EAF6FFC74C0CFF0E3A9AFEFEEFFD" box="[171,262,4077,4116]" italics="true" pageId="8" pageNumber="357">York</emphasis>
, habitat modification in southern New Guinea, and frequent hot burning regimes in parts of Cape
<emphasis id="B908EAF6FFC74C0CFE462555FDC4EFA0" box="[483,572,4130,4169]" italics="true" pageId="8" pageNumber="357">York</emphasis>
affecting succession and longevity of large, old trees required for roost hollows. The Papuan Sheath-tailed Bat occurs in protected areas on Cape
<emphasis id="B908EAF6FFC74C0CF9002521F904EF94" box="[1701,1788,4182,4221]" italics="true" pageId="8" pageNumber="357">York</emphasis>
Peninsula and Tonda Wildlife Management Area in Papua New Guinea. Research on basic biology, in particular habitat and roosting preferences, is needed. Acoustic
<emphasis id="B908EAF6FFC74C0CF9DF25B7F903EF0E" box="[1658,1787,4288,4327]" italics="true" pageId="8" pageNumber="357">surveys</emphasis>
should be conducted to better define distribution especially in Papua New Guinea.
</paragraph>
</subSubSection>
<subSubSection id="C366656FFFC74C0CFF0B2446FD40EE5E" pageId="8" pageNumber="357" type="reference_group">
<paragraph id="8BC336E4FFC74C0CFF0B2446FD40EE5E" blockId="8.[171,1790,2756,4535]" pageId="8" pageNumber="357">
<emphasis id="B908EAF6FFC74C0CFF0B2446FE86EEA5" bold="true" box="[174,382,4401,4428]" pageId="8" pageNumber="357">Bibliography.</emphasis>
Armstrong &amp; Konishi (2013), Armstrong, Broken-Brow et
<emphasis id="B908EAF6FFC74C0CFB4A2446FAF6EEA5" box="[1263,1294,4401,4428]" italics="true" pageId="8" pageNumber="357">at.</emphasis>
(2017), Bonaccorso (1998), Flannery (1995a), Hall &amp; Richards (1979), Reardon
<emphasis id="B908EAF6FFC74C0CFCF92411FC77EE68" box="[860,911,4454,4481]" italics="true" pageId="8" pageNumber="357">eta</emphasis>
/. (2010), Strahan (1995),Tate (1941),Thomson
<emphasis id="B908EAF6FFC74C0CF9E02411F980EE68" box="[1605,1656,4454,4481]" italics="true" pageId="8" pageNumber="357">eta</emphasis>
/. (2008), Troughton (1925),
<emphasis id="B908EAF6FFC74C0CFE1024EBFDB3EE5E" box="[437,587,4508,4535]" italics="true" pageId="8" pageNumber="357">Waithman</emphasis>
(1979).
</paragraph>
</subSubSection>
</treatment>
</document>