treatments-xml/data/03/D5/87/03D587B2FFA31B4B29AC44ACFB00FE07.xml

<document id="41FE5FFF230D4EFE25D4FB72BAAB93E6" ID-DOI="10.24349/acarologia/20204407" ID-ISSN="2107-7207" ID-Zenodo-Dep="5757771" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630330299514" checkinUser="marcus" docAuthor="Mironov, Sergey V. &amp; Galloway, Terry D." docDate="2020" docId="03D587B2FFA31B4B29AC44ACFB00FE07" docLanguage="en" docName="Acarologia.2020.60.4.878-891.pdf" docOrigin="Acarologia 60 (4)" docSource="http://dx.doi.org/10.24349/acarologia/20204407" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Nycteridocaulus sulcatus Mironov &amp; Galloway 2020, sp. n." docType="treatment" docUuid="33E79AC0-047C-41F5-9610-7B62F20718F8" docUuidSource="ZooBank" docVersion="4" lastPageNumber="885" masterDocId="FFECFFCAFFA01B4328474668FFAEFFE8" masterDocTitle="Two new feather mites of the subfamily Proctophyllodinae (Acariformes: Proctophyllodidae) from the Yellow-bellied Flycatcher Empidonax flaviventris (Passeriformes: Tyrannidae) in Canada" masterLastPageNumber="891" masterPageNumber="878" pageNumber="880" updateTime="1699230057472" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B48F1C48CBEC07328DD161E59C19D322">Two new feather mites of the subfamily Proctophyllodinae (Acariformes: Proctophyllodidae) from the Yellow-bellied Flycatcher Empidonax flaviventris (Passeriformes: Tyrannidae) in Canada</mods:title>
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<mods:namePart id="C3F66CDAAC1A9872C528B5171B52CFB1">Mironov, Sergey V.</mods:namePart>
<mods:affiliation id="90EC0D8AD7B081AADB60FDA0229CC39E">Zoological Institute, Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint</mods:affiliation>
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<mods:namePart id="B751E5607D91AB987C81892E3D786316">Galloway, Terry D.</mods:namePart>
<mods:affiliation id="89012E46714F190EDC41A4BB8C5E42C1">Department of Entomology, University of Manitoba, Winnipeg, Manitoba, R 3 T 2 N 2, Canada. &amp; Zoological Institute, Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint</mods:affiliation>
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<mods:title id="DE619714C40C07FF9EF24968398E56F8">Acarologia</mods:title>
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<mods:date id="0E8C574A9228CC5C2E4B7BCB6DA754E0">2020</mods:date>
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<subSubSection id="C366652FFFA31B4029AC44ACFC48FCE7" pageId="3" pageNumber="880" type="nomenclature">
<paragraph id="8BC336A4FFA31B4029AC44ACFCD4FD36" blockId="3.[491,890,708,734]" box="[491,890,708,734]" pageId="3" pageNumber="880">
<heading id="D08B81C8FFA31B4029AC44ACFCD4FD36" bold="true" box="[491,890,708,734]" fontSize="11" level="2" pageId="3" pageNumber="880" reason="3">
<emphasis id="B908EAB6FFA31B4029AC44ACFCD4FD36" bold="true" box="[491,890,708,734]" pageId="3" pageNumber="880">
<taxonomicName id="4C7C4D27FFA31B4029AC44ACFC85FD36" authority="Mironov &amp; Galloway, 2020" authorityName="Mironov &amp; Galloway" authorityYear="2020" box="[491,811,708,734]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="880" phylum="Arthropoda" rank="species" species="sulcatus" status="sp. nov.">
<emphasis id="B908EAB6FFA31B4029AC44ACFC85FD36" bold="true" box="[491,811,708,734]" italics="true" pageId="3" pageNumber="880">Nycteridocaulus sulcatus</emphasis>
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<taxonomicNameLabel id="A23B57CDFFA31B402B7444ACFCD4FD36" box="[819,890,708,734]" pageId="3" pageNumber="880" rank="species">sp. n.</taxonomicNameLabel>
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Zoobank: 
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</paragraph>
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<subSubSection id="C366652FFFA31B402A544573FD3DFCDB" box="[531,659,795,819]" pageId="3" pageNumber="880" type="description">
<paragraph id="8BC336A4FFA31B402A544573FD3DFCDB" blockId="3.[491,1547,764,2028]" box="[531,659,795,819]" pageId="3" pageNumber="880">
<figureCitation id="13472A21FFA31B402A544573FD3DFCDB" box="[531,659,795,819]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="4.[119,186,1883,1904]" captionStartId-1="5.[119,186,1722,1743]" captionStartId-2="6.[119,186,1911,1932]" captionTargetBox-0="[96,1502,306,1804]" captionTargetBox-1="[95,1502,251,1634]" captionTargetBox-2="[114,1477,238,1839]" captionTargetId-0="figure-2@4.[95,1502,299,1830]" captionTargetId-1="figure-2@5.[95,1502,221,1669]" captionTargetId-2="figure-2@6.[95,1502,226,1858]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figure 1 Nycteridocaulus sulcatussp. n., male: A – dorsal view, B – ventral view." captionText-1="Figure 2 Nycteridocaulus sulcatussp. n., female: A – dorsal view, B – ventral view." captionText-2="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi-0="http://doi.org/10.5281/zenodo.5757773" figureDoi-1="http://doi.org/10.5281/zenodo.5757775" figureDoi-2="http://doi.org/10.5281/zenodo.5757777" httpUri-0="https://zenodo.org/record/5757773/files/figure.png" httpUri-1="https://zenodo.org/record/5757775/files/figure.png" httpUri-2="https://zenodo.org/record/5757777/files/figure.png" pageId="3" pageNumber="880">Figures 1–3</figureCitation>
</paragraph>
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<subSubSection id="C366652FFFA31B402A544556FCBDFBA0" pageId="3" pageNumber="880" type="materials_examined">
<paragraph id="8BC336A4FFA31B402A544556FA69FC90" blockId="3.[491,1547,764,2028]" pageId="3" pageNumber="880">
<emphasis id="B908EAB6FFA31B402A544556FD1FFCBE" bold="true" box="[531,689,830,854]" pageId="3" pageNumber="880">Type material</emphasis>
— Male 
<typeStatus id="54C78806FFA31B402B544556FCD6FCBE" box="[787,888,830,854]" pageId="3" pageNumber="880" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D7AFD2DFFA31B402BC54557FC78FCBE" box="[898,982,830,854]" pageId="3" pageNumber="880" type="male">7 males</specimenCount>
, 
<specimenCount id="9D7AFD2DFFA31B402BA74556FBFDFCBE" box="[992,1107,830,854]" pageId="3" pageNumber="880" type="female">12 females</specimenCount>
from 
<taxonomicName id="4C7C4D27FFA31B402CCB4556FA26FCBE" baseAuthorityName="Baird &amp; Baird" baseAuthorityYear="1843" box="[1164,1416,830,854]" class="Aves" family="Tyrannidae" genus="Empidonax" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="880" phylum="Chordata" rank="species" species="flaviventris">
<emphasis id="B908EAB6FFA31B402CCB4556FA26FCBE" box="[1164,1416,830,854]" italics="true" pageId="3" pageNumber="880">Empidonax flaviventris</emphasis>
</taxonomicName>
(
<taxonomicName id="4C7C4D27FFA31B402DC84556FD91FC90" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="880" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
, 
<taxonomicName id="4C7C4D27FFA31B402A0C4508FD62FC90" box="[587,716,864,888]" class="Aves" family="Tyrannidae" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="880" phylum="Chordata" rank="family">Tyrannidae</taxonomicName>
), (YBFL/430/CEN/09), (SM 2000), 
<collectingCountry id="F36B7634FFA31B402C0F4508FB12FC90" box="[1096,1212,864,888]" name="Canada" pageId="3" pageNumber="880">CANADA</collectingCountry>
, 
<collectingRegion id="49B8F846FFA31B402C8F4508FA9CFC90" box="[1224,1330,864,888]" country="Canada" name="Manitoba" pageId="3" pageNumber="880">Manitoba</collectingRegion>
, Winnipeg, 5
</paragraph>
<paragraph id="8BC336A4FFA31B4029AC45ECFBAEFC73" blockId="3.[491,1547,764,2028]" box="[491,1024,899,923]" pageId="3" pageNumber="880">June 2009, coll. T.D Galloway and J.A. McLeod.</paragraph>
<paragraph id="8BC336A4FFA31B402A5445CEFA1EFC55" blockId="3.[491,1547,764,2028]" box="[531,1456,933,958]" pageId="3" pageNumber="880">
<emphasis id="B908EAB6FFA31B402A5445CEFD5BFC56" bold="true" box="[531,757,934,958]" pageId="3" pageNumber="880">Additional material</emphasis>
— 
<specimenCount id="9D7AFD2DFFA31B402B6745CDFCDCFC55" box="[800,882,933,957]" pageId="3" pageNumber="880" type="male">4 males</specimenCount>
, 
<specimenCount id="9D7AFD2DFFA31B402B3945CDFC49FC55" box="[894,999,933,957]" pageId="3" pageNumber="880" type="female">5 females</specimenCount>
from 
<taxonomicName id="4C7C4D27FFA31B402C6845CEFB69FC55" baseAuthorityName="Baird &amp; Baird" baseAuthorityYear="1843" box="[1071,1223,933,957]" class="Aves" family="Tyrannidae" genus="Empidonax" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="880" phylum="Chordata" rank="species" species="flaviventris">
<emphasis id="B908EAB6FFA31B402C6845CEFB69FC55" box="[1071,1223,933,957]" italics="true" pageId="3" pageNumber="880">E. flaviventris</emphasis>
</taxonomicName>
, (OHM b854­14) (SM
</paragraph>
<paragraph id="8BC336A4FFA31B4029AC45A0FDFEFBEB" blockId="3.[491,1547,764,2028]" pageId="3" pageNumber="880">
2601), 
<collectingCountry id="F36B7634FFA31B402A7D45A0FD00FC08" box="[570,686,968,992]" name="Canada" pageId="3" pageNumber="880">CANADA</collectingCountry>
, 
<collectingRegion id="49B8F846FFA31B402AFC45A0FC8BFC08" box="[699,805,968,992]" country="Canada" name="Manitoba" pageId="3" pageNumber="880">Manitoba</collectingRegion>
, High Bluff, 
<date id="FFC21064FFA31B402B8245A0FBDAFC08" box="[965,1140,968,992]" pageId="3" pageNumber="880" value="2014-08-31">31 August 2014</date>
, coll. T.D Galloway and E.N. McNally.
</paragraph>
<paragraph id="8BC336A4FFA31B402A544266FCBDFBA0" blockId="3.[491,1547,764,2028]" pageId="3" pageNumber="880">
<emphasis id="B908EAB6FFA31B402A544266FD21FBCE" bold="true" box="[531,655,1038,1062]" pageId="3" pageNumber="880">Depository</emphasis>
— 
<materialsCitation id="3B143CF9FFA31B402AFD4265FCBDFBA0" collectionCode="CNC, UMMZ, WRME, ZISP" pageId="3" pageNumber="880" specimenCount="6" specimenCount-female="4" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="54C78806FFA31B402AFD4265FC8EFBCD" box="[698,800,1037,1061]" pageId="3" pageNumber="880" type="holotype">Holotype</typeStatus>
, 
<specimenCount id="9D7AFD2DFFA31B402B6C4265FCDAFBCD" box="[811,884,1037,1061]" pageId="3" pageNumber="880" type="male">2 male</specimenCount>
and 
<specimenCount id="9D7AFD2DFFA31B402BF74265FBA7FBCD" box="[944,1033,1037,1061]" pageId="3" pageNumber="880" type="female">4 female</specimenCount>
<typeStatus id="54C78806FFA31B402C564265FBD6FBCD" box="[1041,1144,1037,1061]" pageId="3" pageNumber="880" type="paratype">paratypes</typeStatus>
— 
<collectionCode id="ED6DAE61FFA31B402CD34265FB65FBCD" box="[1172,1227,1037,1061]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="880">CNC</collectionCode>
, remaining 
<typeStatus id="54C78806FFA31B402D174266FA1BFBCE" box="[1360,1461,1038,1062]" pageId="3" pageNumber="880" type="paratype">paratypes</typeStatus>
— 
<collectionCode id="ED6DAE61FFA31B4029AC4258FDEEFBAF" box="[491,576,1072,1095]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="3" pageNumber="880" type="University or college">UMMZ</collectionCode>
, 
<collectionCode id="ED6DAE61FFA31B402A0C4258FD30FBAF" box="[587,670,1072,1095]" pageId="3" pageNumber="880">WRME</collectionCode>
and 
<collectionCode id="ED6DAE61FFA31B402A9F4258FCA1FBA0" box="[728,783,1072,1096]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="880">ZISP</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C366652FFFA31B442A54423BFB94F92F" lastPageId="7" lastPageNumber="884" pageId="3" pageNumber="880" type="description">
<paragraph id="8BC336A4FFA31B402A54423BFAC4F930" blockId="3.[491,1547,764,2028]" pageId="3" pageNumber="880">
<emphasis id="B908EAB6FFA31B402A54423BFD59FB83" bold="true" box="[531,759,1107,1131]" pageId="3" pageNumber="880">Description — Male</emphasis>
(
<typeStatus id="54C78806FFA31B402B43423AFCC4FB82" box="[772,874,1106,1130]" pageId="3" pageNumber="880" type="holotype">holotype</typeStatus>
, range for 
<specimenCount id="9D7AFD2DFFA31B402BA6423BFBF4FB83" box="[993,1114,1107,1131]" pageId="3" pageNumber="880" type="generic" typeStatus="paratypes">7 paratypes</specimenCount>
in parentheses) (
<figureCitation id="13472A21FFA31B402D54423AFAD2FB82" box="[1299,1404,1106,1130]" captionStart="Figure 1" captionStartId="4.[119,186,1883,1904]" captionTargetBox="[96,1502,306,1804]" captionTargetId="figure-2@4.[95,1502,299,1830]" captionTargetPageId="4" captionText="Figure 1 Nycteridocaulus sulcatussp. n., male: A – dorsal view, B – ventral view." figureDoi="http://doi.org/10.5281/zenodo.5757773" httpUri="https://zenodo.org/record/5757773/files/figure.png" pageId="3" pageNumber="880">Figures 1</figureCitation>
, 
<figureCitation id="13472A21FFA31B402DCE423AFA3AFB82" box="[1417,1428,1106,1130]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="3" pageNumber="880">3</figureCitation>
A­D, G). Idiosoma, length × width, 290 (275–285) × 160 (150–165), length of hysterosoma 185 (180– 190). Prodorsal shield: anterolateral extensions with acute tips, lateral margins entire, posterior margin slightly concave, posterior corners almost rectangular, 82 (80–88) in length, 105 (100– 105) in width, posterior part with minute sparsely disposed lacunae. Setae 
<emphasis id="B908EAB6FFA31B402D5C42B5FA9DFB1C" box="[1307,1331,1245,1268]" italics="true" pageId="3" pageNumber="880">ve</emphasis>
rudimentary. Scapular setae 
<emphasis id="B908EAB6FFA31B402AC84368FD0BFAFF" box="[655,677,1280,1303]" italics="true" pageId="3" pageNumber="880">se</emphasis>
separated by 60 (57–63). Setae 
<emphasis id="B908EAB6FFA31B402C4B4297FB8AFAFF" box="[1036,1060,1279,1303]" italics="true" pageId="3" pageNumber="880">c2</emphasis>
and 
<emphasis id="B908EAB6FFA31B402C1B4368FBDBFAFF" box="[1116,1141,1280,1303]" italics="true" pageId="3" pageNumber="880">cp</emphasis>
on humeral shields. Setae 
<emphasis id="B908EAB6FFA31B402DE64297FA14FAFF" box="[1441,1466,1279,1303]" italics="true" pageId="3" pageNumber="880">c3</emphasis>
lanceolate, 13 (13–15) long, about 5 wide. Distance between prodorsal and hysteronotal shields 25 (20–25). Hysteronotal shield: 185 (175–185) in length and 105 (95–100) in width; anterior margin slightly concave, surface with sparsely disposed, small, circular lacunae. Opisthosoma slightly attenuate posteriorly. Opisthosomal lobes short, roughly rectangular, posterior margin almost straight; terminal lamellae short rectangular, 10 (10–13) in length and 18 (16–18) in width. Terminal cleft small triangular, with anterior end extending to level of setae 
<emphasis id="B908EAB6FFA31B402D2043A6FA2FFA0E" box="[1383,1409,1486,1510]" italics="true" pageId="3" pageNumber="880">h2</emphasis>
, length from anterior end to margins of lobes 15 (13–15), width at level of setae 
<emphasis id="B908EAB6FFA31B402CB64399FAB9F9E1" box="[1265,1303,1521,1545]" italics="true" pageId="3" pageNumber="880">ps1</emphasis>
6 (6–8). Supranal concavity long, parallel­sided, with strongly sclerotized margins, open posteriorly, 58 (52–58) long. Setae 
<emphasis id="B908EAB6FFA31B402A2B405EFD29F9A6" box="[620,647,1590,1614]" italics="true" pageId="3" pageNumber="880">h2</emphasis>
situated on truncate lateral extensions of opisthosomal lobes; setae 
<emphasis id="B908EAB6FFA31B402D1B405EFAD9F9A6" box="[1372,1399,1590,1614]" italics="true" pageId="3" pageNumber="880">h3</emphasis>
situated in posterolateral angles of lobes; setae 
<emphasis id="B908EAB6FFA31B402BCD4030FC01F998" box="[906,943,1624,1648]" italics="true" pageId="3" pageNumber="880">
ps 
<quantity id="4C849B41FFA31B402BE34030FC01F998" box="[932,943,1624,1648]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="880" unit="in" value="1.0">1</quantity>
</emphasis>
in posteromedial angles of lobes, slightly posterior to level of setae 
<emphasis id="B908EAB6FFA31B402ADB4013FD19F97B" box="[668,695,1659,1683]" italics="true" pageId="3" pageNumber="880">h3</emphasis>
; setae 
<emphasis id="B908EAB6FFA31B402B454013FCB3F97B" box="[770,797,1659,1683]" italics="true" pageId="3" pageNumber="880">h1</emphasis>
situated slightly posterior to setae 
<emphasis id="B908EAB6FFA31B402CD44013FB16F97B" box="[1171,1208,1659,1683]" italics="true" pageId="3" pageNumber="880">ps2</emphasis>
. Distance between dorsal setae: 
<emphasis id="B908EAB6FFA31B402A7340F5FDDAF95D" box="[564,628,1693,1717]" italics="true" pageId="3" pageNumber="880">c2:d2</emphasis>
60 (55–60), 
<emphasis id="B908EAB6FFA31B402AB940F5FC91F95D" box="[766,831,1693,1717]" italics="true" pageId="3" pageNumber="880">d2:e2</emphasis>
65 (62–68), 
<emphasis id="B908EAB6FFA31B402B8E40F5FBA8F95D" box="[969,1030,1693,1717]" italics="true" pageId="3" pageNumber="880">e2:h3</emphasis>
60 (52–60), 
<emphasis id="B908EAB6FFA31B402CD340F5FB7CF95D" box="[1172,1234,1693,1717]" italics="true" pageId="3" pageNumber="880">h2:h2</emphasis>
60 (57–63), 
<emphasis id="B908EAB6FFA31B402D2740F5FA30F95D" box="[1376,1438,1693,1717]" italics="true" pageId="3" pageNumber="880">h3:h3</emphasis>
50 (47–53), 
<emphasis id="B908EAB6FFA31B402A0B40A8FD31F930" box="[588,671,1728,1752]" italics="true" pageId="3" pageNumber="880">ps2:ps2</emphasis>
67 (65–70), 
<emphasis id="B908EAB6FFA31B402B6F40A8FCCBF930" box="[808,869,1728,1752]" italics="true" pageId="3" pageNumber="880">h1:h3</emphasis>
30 (27–30), 
<emphasis id="B908EAB6FFA31B402BA840A8FBE1F930" box="[1007,1103,1728,1752]" italics="true" pageId="3" pageNumber="880">d1:d220</emphasis>
(18–22), 
<emphasis id="B908EAB6FFA31B402CF140A8FB5CF930" box="[1206,1266,1728,1752]" italics="true" pageId="3" pageNumber="880">
e1: 
<geoCoordinate id="EE485063FFA31B402C9140A8FB5CF930" box="[1238,1266,1728,1752]" degrees="2" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="880" precision="925" value="2.4166667">e2</geoCoordinate>
</emphasis>
25 (23–28).
</paragraph>
<paragraph id="8BC336A4FFA31B462A54408AFA63F868" blockId="3.[491,1547,764,2028]" lastBlockId="5.[491,1547,1862,2024]" lastPageId="5" lastPageNumber="882" pageId="3" pageNumber="880">
Epimerites I free, close to each other, posterior tips divergent; epimerites I, II with narrow surface fields; epimerites IVa absent. Rudimentary sclerite rEpIIa barely distinct. Trochanters III flanked by sclerotized bands going from bases of epimerites IIIa. Epimerites IIIa with sclerotized areas around their inner tips with acute extensions directed anteromediad. Trochanters IV flanked by sclerotized bands stretching from bases of epimerites IV. Genital apparatus situated at level of trochanters IV. Genital arch shaped as recurved bow, 8 (7–8) in length and 45 (45–48) in width. Aedeagus stylet­like, 17 (17–18) in length, reaching level of setae 
<emphasis id="B908EAB6FFA31B402A6F41BDFD96F804" box="[552,568,2005,2028]" italics="true" pageId="3" pageNumber="880">g</emphasis>
(
<figureCitation id="13472A21FFA31B402A0241BCFD17F804" box="[581,697,2004,2028]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="3" pageNumber="880">Figure 3G</figureCitation>
). Genital papillae of each side on small oval plates at level of genital arch apex. Distance from genital arch apex to level of setae 
<emphasis id="B908EAB6FFA51B462C77412EFBE4F8B6" box="[1072,1098,1862,1886]" italics="true" pageId="5" pageNumber="882">h3</emphasis>
105 (100–105). Pregenital apodemes shaped as longitudinal sclerites of irregular form. Setae 
<emphasis id="B908EAB6FFA51B462C004100FBCAF868" box="[1095,1124,1896,1920]" italics="true" pageId="5" pageNumber="882">4a</emphasis>
on pregenital apodemes, setae 
<emphasis id="B908EAB6FFA51B462DF54100FA63F868" box="[1458,1485,1896,1920]" italics="true" pageId="5" pageNumber="882">4b</emphasis>
</paragraph>
<caption id="DF03662CFFA41B4728304133FC34F899" ID-DOI="http://doi.org/10.5281/zenodo.5757773" ID-Zenodo-Dep="5757773" box="[119,922,1883,1905]" httpUri="https://zenodo.org/record/5757773/files/figure.png" pageId="4" pageNumber="881" startId="4.[119,186,1883,1904]" targetBox="[96,1502,306,1804]" targetPageId="4">
<paragraph id="8BC336A4FFA41B4728304133FC34F899" blockId="4.[119,922,1883,1905]" box="[119,922,1883,1905]" pageId="4" pageNumber="881">
<emphasis id="B908EAB6FFA41B4728304133FF62F898" bold="true" box="[119,204,1883,1904]" pageId="4" pageNumber="881">Figure 1</emphasis>
<taxonomicName id="4C7C4D27FFA41B47289F4133FE61F899" authority="Mironov &amp; Galloway, 2020" authorityName="Mironov &amp; Galloway" authorityYear="2020" box="[216,463,1883,1905]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="881" phylum="Arthropoda" rank="species" species="sulcatus" status="sp. nov.">
<emphasis id="B908EAB6FFA41B47289F4133FE61F899" box="[216,463,1883,1905]" italics="true" pageId="4" pageNumber="881">Nycteridocaulus sulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57CDFFA41B4729884133FDA8F898" box="[463,518,1883,1904]" pageId="4" pageNumber="881" rank="species">sp. n.</taxonomicNameLabel>
, male: A – dorsal view, B – ventral view.
</paragraph>
</caption>
<caption id="DF03662CFFA51B46283040D2FC02F938" ID-DOI="http://doi.org/10.5281/zenodo.5757775" ID-Zenodo-Dep="5757775" box="[119,940,1722,1744]" httpUri="https://zenodo.org/record/5757775/files/figure.png" pageId="5" pageNumber="882" startId="5.[119,186,1722,1743]" targetBox="[95,1502,251,1634]" targetPageId="5">
<paragraph id="8BC336A4FFA51B46283040D2FC02F938" blockId="5.[119,940,1722,1744]" box="[119,940,1722,1744]" pageId="5" pageNumber="882">
<emphasis id="B908EAB6FFA51B46283040D2FF62F927" bold="true" box="[119,204,1722,1743]" pageId="5" pageNumber="882">Figure 2</emphasis>
<taxonomicName id="4C7C4D27FFA51B46289F40D2FE61F938" authority="Mironov &amp; Galloway, 2020" authorityName="Mironov &amp; Galloway" authorityYear="2020" box="[216,463,1722,1744]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="882" phylum="Arthropoda" rank="species" species="sulcatus" status="sp. nov.">
<emphasis id="B908EAB6FFA51B46289F40D2FE61F938" box="[216,463,1722,1744]" italics="true" pageId="5" pageNumber="882">Nycteridocaulus sulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57CDFFA51B46298840D2FDA8F927" box="[463,518,1722,1743]" pageId="5" pageNumber="882" rank="species">sp. n.</taxonomicNameLabel>
, female: A – dorsal view, B – ventral view.
</paragraph>
</caption>
<paragraph id="8BC336A4FFA51B4429AC41E4FA6CFEAA" blockId="5.[491,1547,1862,2024]" lastBlockId="7.[490,1573,229,2045]" lastPageId="7" lastPageNumber="884" pageId="5" pageNumber="882">
on sclerotized areas of epimerites IIIa. Adanal suckers 15 (14–15) in diameter, corolla with 11–12 denticles. Adanal shields represented by a pair of roughly L­shaped sclerites situated anterior to anal suckers and touching at midline, length of shields 25 (23–25), distance between their lateral margins 48 (44–48). Setae 
<emphasis id="B908EAB6FFA71B442BD4468DFC16FF15" box="[915,952,229,253]" italics="true" pageId="7" pageNumber="884">ps3</emphasis>
situated on transverse branches of adanal shields. Opisthoventral shields shaped as small, roughly triangular extensions at level of adanal suckers. Distance between ventral setae: 
<emphasis id="B908EAB6FFA71B442B0C4742FC25FEAA" box="[843,907,298,322]" italics="true" pageId="7" pageNumber="884">3a:4b</emphasis>
7 (5–8), 
<emphasis id="B908EAB6FFA71B442BA84742FB81FEAA" box="[1007,1071,298,322]" italics="true" pageId="7" pageNumber="884">4b:4a</emphasis>
27 (23–27), 
<emphasis id="B908EAB6FFA71B442CFC4742FB5EFEAA" box="[1211,1264,298,322]" italics="true" pageId="7" pageNumber="884">4a:g</emphasis>
35 (32–35), 
<emphasis id="B908EAB6FFA71B442D3C4743FA0AFEAA" box="[1403,1444,299,322]" italics="true" pageId="7" pageNumber="884">g:g</emphasis>
20
</paragraph>
<caption id="DF03662CFFA61B452830411FFAD1F852" ID-DOI="http://doi.org/10.5281/zenodo.5757777" ID-Zenodo-Dep="5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="6" pageNumber="883" startId="6.[119,186,1911,1932]" targetBox="[114,1477,238,1839]" targetPageId="6">
<paragraph id="8BC336A4FFA61B452830411FFA73F865" blockId="6.[119,1541,1911,1978]" box="[119,1501,1911,1933]" pageId="6" pageNumber="883">
<emphasis id="B908EAB6FFA61B452830411FFF63F864" bold="true" box="[119,205,1911,1932]" pageId="6" pageNumber="883">Figure 3</emphasis>
<taxonomicName id="4C7C4D27FFA61B45289E411FFE7EF865" authority="Mironov &amp; Galloway, 2020" authorityName="Mironov &amp; Galloway" authorityYear="2020" box="[217,464,1911,1933]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="883" phylum="Arthropoda" rank="species" species="sulcatus" status="sp. nov.">
<emphasis id="B908EAB6FFA61B45289E411FFE7EF865" box="[217,464,1911,1933]" italics="true" pageId="6" pageNumber="883">Nycteridocaulus sulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57CDFFA61B4529974110FDA6F865" box="[464,520,1912,1933]" pageId="6" pageNumber="883" rank="species">sp. n.</taxonomicNameLabel>
, details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus
</paragraph>
<paragraph id="8BC336A4FFA61B45283041E6FAD1F852" blockId="6.[119,1541,1911,1978]" pageId="6" pageNumber="883">IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella.</paragraph>
</caption>
<paragraph id="8BC336A4FFA71B4429AC4725FB1FFE8D" blockId="7.[490,1573,229,2045]" box="[491,1201,333,357]" pageId="7" pageNumber="884">
(19–21), 
<emphasis id="B908EAB6FFA71B442A0B4725FD07FE8D" box="[588,681,333,357]" italics="true" pageId="7" pageNumber="884">g:ps325</emphasis>
(23–25), 
<emphasis id="B908EAB6FFA71B442B574725FCCDFE8D" box="[784,867,333,357]" italics="true" pageId="7" pageNumber="884">ps3:ps3</emphasis>
27 (23–28), 
<emphasis id="B908EAB6FFA71B442BAB4725FB9AFE8D" box="[1004,1076,333,357]" italics="true" pageId="7" pageNumber="884">ps3:h3</emphasis>
65 (58–65).
</paragraph>
<paragraph id="8BC336A4FFA71B442A544718FA7CFDDC" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
Femora I, II with ventral crest. Solenidion 
<emphasis id="B908EAB6FFA71B442BAC4718FC56FE6F" box="[1003,1016,368,391]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
of genu I approximately 1.5 times longer than this segment and situated at its midlevel (
<figureCitation id="13472A21FFA71B442C4347FAFBD7FE42" box="[1028,1145,402,426]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="7" pageNumber="884">Figure 3A</figureCitation>
). Solenidion 
<emphasis id="B908EAB6FFA71B442CA547FBFB41FE42" box="[1250,1263,403,426]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
of genu III situated at midlevel of this segment. Legs III, IV subequal in size, legs IV with ambulacral discs extending beyond posterior margin of terminal lamellae. Tarsus IV 25 (22–25) in length, with longitudinal dorsal ridge and with triangular ventral process bearing 
<emphasis id="B908EAB6FFA71B442CBB4792FAA0FDF9" box="[1276,1294,506,529]" italics="true" pageId="7" pageNumber="884">w</emphasis>
setamodified ; setae 
<emphasis id="B908EAB6FFA71B442D874791FA7CFDF9" box="[1472,1490,505,529]" italics="true" pageId="7" pageNumber="884">d</emphasis>
and 
<emphasis id="B908EAB6FFA71B442A5E4475FD8BFDDC" box="[537,549,541,564]" italics="true" pageId="7" pageNumber="884">e</emphasis>
button­like, setae 
<emphasis id="B908EAB6FFA71B442AAD4474FD52FDDC" box="[746,764,540,564]" italics="true" pageId="7" pageNumber="884">d</emphasis>
situated at midlength of segment (
<figureCitation id="13472A21FFA71B442CCE4474FB52FDDC" box="[1161,1276,540,564]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="7" pageNumber="884">Figure 3D</figureCitation>
). Setae 
<emphasis id="B908EAB6FFA71B442D774474FAECFDDC" box="[1328,1346,540,564]" italics="true" pageId="7" pageNumber="884">d</emphasis>
and 
<emphasis id="B908EAB6FFA71B442D344474FAD4FDDC" box="[1395,1402,540,564]" italics="true" pageId="7" pageNumber="884">f</emphasis>
of tarsi
</paragraph>
<paragraph id="8BC336A4FFA71B4429AC4457FDECFD91" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
II, III longer, subequal in length. Length of solenidia: 
<emphasis id="B908EAB6FFA71B442C754457FB91FDBE" box="[1074,1087,575,598]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
I 35 (35–40), 
<emphasis id="B908EAB6FFA71B442C944457FB4EFDBE" box="[1235,1248,575,598]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
III 20 (20–25), 
<emphasis id="B908EAB6FFA71B442DC14457FA3BFDBE" box="[1414,1429,575,598]" italics="true" pageId="7" pageNumber="884">φ</emphasis>
IV 25 (25–30).
</paragraph>
<paragraph id="8BC336A4FFA71B442A5444EBFA74FD73" blockId="7.[490,1573,229,2045]" box="[531,1498,643,667]" pageId="7" pageNumber="884">
<emphasis id="B908EAB6FFA71B442A5444EBFDC8FD73" bold="true" box="[531,614,643,667]" pageId="7" pageNumber="884">Female</emphasis>
(range for 
<specimenCount id="9D7AFD2DFFA71B442AA144EBFCDFFD73" box="[742,881,643,667]" pageId="7" pageNumber="884" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
) (
<figureCitation id="13472A21FFA71B442BC044EBFC5BFD73" box="[903,1013,643,667]" captionStart="Figure 2" captionStartId="5.[119,186,1722,1743]" captionTargetBox="[95,1502,251,1634]" captionTargetId="figure-2@5.[95,1502,221,1669]" captionTargetPageId="5" captionText="Figure 2 Nycteridocaulus sulcatussp. n., female: A – dorsal view, B – ventral view." figureDoi="http://doi.org/10.5281/zenodo.5757775" httpUri="https://zenodo.org/record/5757775/files/figure.png" pageId="7" pageNumber="884">Figures 2</figureCitation>
, 
<figureCitation id="13472A21FFA71B442C4544EBFB92FD73" box="[1026,1084,643,667]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="7" pageNumber="884">3E, F</figureCitation>
). Idiosoma, length × width, 365–385
</paragraph>
<paragraph id="8BC336A4FFA71B4429AC44CEFA76FCEA" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
× 165–180, length of hysterosoma 250–270. Prodorsal shield: generally shaped as in male, posterior corners pointed, 
<quantity id="4C849B41FFA71B442B4244A0FCF6FD08" box="[773,856,712,736]" metricMagnitude="0" metricUnit="m" metricValue="2.3495" metricValueMax="2.413" metricValueMin="2.286" pageId="7" pageNumber="884" unit="in" value="92.5" valueMax="95.0" valueMin="90.0">90–95 in</quantity>
length and 
<quantity id="4C849B41FFA71B442BA444A0FBF4FD08" box="[995,1114,712,736]" metricMagnitude="0" metricUnit="m" metricValue="3.1115" metricValueMax="3.175" metricValueMin="3.048" pageId="7" pageNumber="884" unit="in" value="122.5" valueMax="125.0" valueMin="120.0">120–125 in</quantity>
width, surface with sparsely disposed minute circular lacunae. Setae 
<emphasis id="B908EAB6FFA71B442B7A4483FCFDFCEA" box="[829,851,747,770]" italics="true" pageId="7" pageNumber="884">se</emphasis>
separated by 75–78. Setae 
<emphasis id="B908EAB6FFA71B442CC44482FB32FCEA" box="[1155,1180,746,770]" italics="true" pageId="7" pageNumber="884">c2</emphasis>
and 
<emphasis id="B908EAB6FFA71B442C954483FB45FCEA" box="[1234,1259,747,770]" italics="true" pageId="7" pageNumber="884">cp</emphasis>
on humeral shields.
</paragraph>
<paragraph id="8BC336A4FFA71B4429AC4565FA13FB2A" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
Setae 
<emphasis id="B908EAB6FFA71B442A6B4565FDEBFCCD" box="[556,581,781,805]" italics="true" pageId="7" pageNumber="884">c3</emphasis>
lanceolate, 15–18 long, about 5 wide. Distance between prodorsal and hysteronotal shields 25–32. Anterior hysteronotal shield: 
<quantity id="4C849B41FFA71B442BA54547FBFEFCAF" box="[994,1104,815,839]" metricMagnitude="0" metricUnit="m" metricValue="4.7625" metricValueMax="4.826" metricValueMin="4.699" pageId="7" pageNumber="884" unit="in" value="187.5" valueMax="190.0" valueMin="185.0">185–190 in</quantity>
length and 
<quantity id="4C849B41FFA71B442C994547FAFBFCAF" box="[1246,1365,815,839]" metricMagnitude="0" metricUnit="m" metricValue="2.8575" metricValueMax="2.921" metricValueMin="2.7939999999999996" pageId="7" pageNumber="884" unit="in" value="112.5" valueMax="115.0" valueMin="110.0">110–115 in</quantity>
width, anterior margin slightly concave, posterior margin with small truncate median extension and a pair of shallow concavities, median area of posterior half with sparsely disposed small circular lacunae (
<figureCitation id="13472A21FFA71B442A0945FFFD1CFC47" box="[590,690,919,943]" captionStart="Figure 2" captionStartId="5.[119,186,1722,1743]" captionTargetBox="[95,1502,251,1634]" captionTargetId="figure-2@5.[95,1502,221,1669]" captionTargetPageId="5" captionText="Figure 2 Nycteridocaulus sulcatussp. n., female: A – dorsal view, B – ventral view." figureDoi="http://doi.org/10.5281/zenodo.5757775" httpUri="https://zenodo.org/record/5757775/files/figure.png" pageId="7" pageNumber="884">Fig. 2A</figureCitation>
). Lobar region: 
<quantity id="4C849B41FFA71B442B2845FFFC61FC47" box="[879,975,919,943]" metricMagnitude="0" metricUnit="m" metricValue="1.8669" metricValueMax="1.905" metricValueMin="1.8288" pageId="7" pageNumber="884" unit="in" value="73.5" valueMax="75.0" valueMin="72.0">72–75 in</quantity>
length and 
<quantity id="4C849B41FFA71B442C2345FFFB71FC47" box="[1124,1247,919,943]" metricMagnitude="0" metricUnit="m" metricValue="2.6035" metricValueMax="2.667" metricValueMin="2.54" pageId="7" pageNumber="884" unit="in" value="102.5" valueMax="105.0" valueMin="100.0">100–105 in</quantity>
width, lateral margins strongly convex, without noticeable extensions bearing setae 
<emphasis id="B908EAB6FFA71B442C3C45D1FB3BFC39" box="[1147,1173,953,977]" italics="true" pageId="7" pageNumber="884">h2</emphasis>
; lobar shield entire, with pair of narrow incisions in anterior margin and with small roughly rectangular extension between them. Terminal cleft narrow, almost parallel­sided, 
<quantity id="4C849B41FFA71B442C674596FBD0FBFE" box="[1056,1150,1022,1046]" metricMagnitude="0" metricUnit="m" metricValue="1.3335" metricValueMax="1.3969999999999998" metricValueMin="1.27" pageId="7" pageNumber="884" unit="in" value="52.5" valueMax="55.0" valueMin="50.0">50–55 in</quantity>
length, 
<quantity id="4C849B41FFA71B442CA74596FA96FBFE" box="[1248,1336,1022,1046]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" metricValueMax="5.588" metricValueMin="4.572" pageId="7" pageNumber="884" unit="in" value="20.0" valueMax="22.0" valueMin="18.0">18–22 in</quantity>
width in anterior part. Supranal concavity absent. Setae 
<emphasis id="B908EAB6FFA71B442BC94248FC06FBD0" box="[910,936,1056,1080]" italics="true" pageId="7" pageNumber="884">h2</emphasis>
lanceolate in basal part and with long filiform apex, 
<quantity id="4C849B41FFA71B4429AC422BFDCAFBB3" box="[491,612,1091,1115]" metricMagnitude="0" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.7939999999999996" pageId="7" pageNumber="884" unit="in" value="115.0" valueMax="120.0" valueMin="110.0">110–120 in</quantity>
length, 
<quantity id="4C849B41FFA71B442A8E422BFCA9FBB3" box="[713,775,1091,1115]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="7" pageNumber="884" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">6–8 in</quantity>
width; setae 
<emphasis id="B908EAB6FFA71B442BD4422BFC00FBB3" box="[915,942,1091,1115]" italics="true" pageId="7" pageNumber="884">h3</emphasis>
70– 
<quantity id="4C849B41FFA71B442B99422BFBBBFBB3" box="[990,1045,1091,1115]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="884" unit="in" value="60.0">60 in</quantity>
length, approximately 3/4 the length of terminal appendages. Setae 
<emphasis id="B908EAB6FFA71B442B59420DFC97FB95" box="[798,825,1125,1149]" italics="true" pageId="7" pageNumber="884">h1</emphasis>
inserted on striated tegument between anterior hysteronotal and lobar shields. Setae 
<emphasis id="B908EAB6FFA71B442AB442E0FCA0FB48" box="[755,782,1160,1184]" italics="true" pageId="7" pageNumber="884">h1</emphasis>
and 
<emphasis id="B908EAB6FFA71B442B0442E0FCF6FB48" box="[835,856,1160,1184]" italics="true" pageId="7" pageNumber="884">
f 
<quantity id="4C849B41FFA71B442B0842E0FCF6FB48" box="[847,856,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="884" unit="in" value="2.0">2</quantity>
</emphasis>
in trapezoidal arrangement. Setae 
<emphasis id="B908EAB6FFA71B442C8842E0FB5BFB48" box="[1231,1269,1160,1184]" italics="true" pageId="7" pageNumber="884">ps1</emphasis>
on inner margins of opisthosomal lobes, close to level of setae 
<emphasis id="B908EAB6FFA71B442BFE42C2FC7DFB2A" box="[953,979,1194,1218]" italics="true" pageId="7" pageNumber="884">h3</emphasis>
. Distance between dorsal setae: 
<emphasis id="B908EAB6FFA71B442D7B42C2FAD3FB2A" box="[1340,1405,1194,1218]" italics="true" pageId="7" pageNumber="884">c2:d2</emphasis>
78–82,
</paragraph>
<paragraph id="8BC336A4FFA71B4429AC42A5FA6EFB0D" blockId="7.[490,1573,229,2045]" box="[491,1472,1229,1253]" pageId="7" pageNumber="884">
<emphasis id="B908EAB6FFA71B4429AC42A5FD82FB0D" box="[491,556,1229,1253]" italics="true" pageId="7" pageNumber="884">d2:e2</emphasis>
95–105, 
<emphasis id="B908EAB6FFA71B442AC842A5FD62FB0D" box="[655,716,1229,1253]" italics="true" pageId="7" pageNumber="884">e2:h2</emphasis>
35–40, 
<emphasis id="B908EAB6FFA71B442B6142A5FCCAFB0D" box="[806,868,1229,1253]" italics="true" pageId="7" pageNumber="884">h2:h3</emphasis>
32–35, 
<emphasis id="B908EAB6FFA71B442BFA42A5FC55FB0D" box="[957,1019,1229,1253]" italics="true" pageId="7" pageNumber="884">h1:h2</emphasis>
35–37, 
<emphasis id="B908EAB6FFA71B442C1242A5FB35FB0D" box="[1109,1179,1229,1253]" italics="true" pageId="7" pageNumber="884">d1:d2</emphasis>
25–28, 
<emphasis id="B908EAB6FFA71B442CAA42A5FA87FB0D" box="[1261,1321,1229,1253]" italics="true" pageId="7" pageNumber="884">
e1: 
<geoCoordinate id="EE485063FFA71B442D4A42A5FA87FB0D" box="[1293,1321,1229,1253]" degrees="2" direction="east" minutes="45" orientation="longitude" pageId="7" pageNumber="884" precision="925" value="2.75">e2</geoCoordinate>
</emphasis>
45–50, 
<emphasis id="B908EAB6FFA71B442DC542A5FA6EFB0D" box="[1410,1472,1229,1253]" italics="true" pageId="7" pageNumber="884">h1:h1</emphasis>
</paragraph>
<paragraph id="8BC336A4FFA71B4429AC4287FCCAFAEF" blockId="7.[490,1573,229,2045]" box="[491,868,1263,1287]" pageId="7" pageNumber="884">
25–33, 
<emphasis id="B908EAB6FFA71B442A7C4287FDD4FAEF" box="[571,634,1263,1287]" italics="true" pageId="7" pageNumber="884">h2:h2</emphasis>
80–84, 
<emphasis id="B908EAB6FFA71B442A974287FCB6FAEF" box="[720,792,1263,1287]" italics="true" pageId="7" pageNumber="884">h2:ps1</emphasis>
22–30.
</paragraph>
<paragraph id="8BC336A4FFA71B442A54437AFA72FA87" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
Epimerites I as in male; epimerites I, II with narrow surface fields. Epimerites IVa large, roughly triangular, with heavily sclerotized margins. Epigynum horseshoe­shaped, thick, lateral margins without extensions, tips acute, not extending to level of setae 
<emphasis id="B908EAB6FFA71B442C8B4330FB77FA87" box="[1228,1241,1368,1391]" italics="true" pageId="7" pageNumber="884">g</emphasis>
, 
<quantity id="4C849B41FFA71B442CA2433FFA94FA87" box="[1253,1338,1367,1391]" metricMagnitude="0" metricUnit="m" metricValue="1.0795" metricValueMax="1.143" metricValueMin="1.016" pageId="7" pageNumber="884" unit="in" value="42.5" valueMax="45.0" valueMin="40.0">40–45 in</quantity>
length, 55–67
</paragraph>
<paragraph id="8BC336A4FFA71B4429AC4311FAFBF9F3" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
in width. Genital papillae of each side on small, ovate sclerotized plate. Setae 
<emphasis id="B908EAB6FFA71B442D704311FAF5FA79" box="[1335,1371,1401,1425]" italics="true" pageId="7" pageNumber="884">ps2</emphasis>
situated at midlevel of anal opening. Translobar apodemes not fused to each other anterior to terminal cleft. Copulatory opening situated on anterior end of terminal cleft. Head of spermatheca short; proximal part of primary spermaduct with ampuliform enlargement 20–25 long (
<figureCitation id="13472A21FFA71B442D234389FA7AFA11" box="[1380,1492,1505,1529]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="7" pageNumber="884">Figure 3F</figureCitation>
). Distance between pseudanal setae: 
<emphasis id="B908EAB6FFA71B442B23406BFC19F9F3" box="[868,951,1539,1563]" italics="true" pageId="7" pageNumber="884">ps2:ps2</emphasis>
43–45, 
<emphasis id="B908EAB6FFA71B442C4A406BFBCEF9F3" box="[1037,1120,1539,1563]" italics="true" pageId="7" pageNumber="884">ps3:ps3</emphasis>
19–21, 
<emphasis id="B908EAB6FFA71B442CF1406BFAA7F9F3" box="[1206,1289,1539,1563]" italics="true" pageId="7" pageNumber="884">ps2:ps3</emphasis>
13–15.
</paragraph>
<paragraph id="8BC336A4FFA71B442A54404EFB94F92F" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
Legs I, II as male. Legs IV with ambulacral disc extending beyond level of setae 
<emphasis id="B908EAB6FFA71B442DDE404DFA1AF9D5" box="[1433,1460,1573,1597]" italics="true" pageId="7" pageNumber="884">h2</emphasis>
. Solenidion 
<emphasis id="B908EAB6FFA71B442A214021FDDDF988" box="[614,627,1609,1632]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
of genu I about 1.5 times longer than this segment and situated at its midlevel. Solenidion 
<emphasis id="B908EAB6FFA71B442A204003FDDAF96A" box="[615,628,1643,1666]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
of genu III in proximal half of this segment. Solenidion 
<emphasis id="B908EAB6FFA71B442CAB4003FB55F96A" box="[1260,1275,1643,1666]" italics="true" pageId="7" pageNumber="884">φ</emphasis>
of tibia IV equal to corresponding tarsus (
<figureCitation id="13472A21FFA71B442B4640E5FCDAF94D" box="[769,884,1677,1701]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="7" pageNumber="884">Figure 3E</figureCitation>
). Setae 
<emphasis id="B908EAB6FFA71B442BFD40E5FC4CF94D" box="[954,994,1677,1701]" italics="true" pageId="7" pageNumber="884">d, f</emphasis>
of tarsi II–IV subequal in length. Length of solenidia: 
<emphasis id="B908EAB6FFA71B442A1B40D8FDC7F92F" box="[604,617,1712,1735]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
I 35–40, 
<emphasis id="B908EAB6FFA71B442A8E40D8FD78F92F" box="[713,726,1712,1735]" italics="true" pageId="7" pageNumber="884">σ</emphasis>
III 23–25, 
<emphasis id="B908EAB6FFA71B442B0F40D8FCF9F92F" box="[840,855,1712,1735]" italics="true" pageId="7" pageNumber="884">φ</emphasis>
III 50–55, 
<emphasis id="B908EAB6FFA71B442B8040D8FC78F92F" box="[967,982,1712,1735]" italics="true" pageId="7" pageNumber="884">φ</emphasis>
IV 30–35.
</paragraph>
</subSubSection>
<subSubSection id="C366652FFFA71B4B2A5440BAFB5AFE42" lastPageId="8" lastPageNumber="885" pageId="7" pageNumber="884" type="diagnosis">
<paragraph id="8BC336A4FFA71B442A5440BAFA18F8E4" blockId="7.[490,1573,229,2045]" pageId="7" pageNumber="884">
<emphasis id="B908EAB6FFA71B442A5440BAFCA6F902" bold="true" box="[531,776,1746,1770]" pageId="7" pageNumber="884">Differential diagnosis</emphasis>
— The new species, 
<taxonomicName id="4C7C4D27FFA71B442BBF40BAFAA0F902" authority="Mironov &amp; Galloway, 2020" authorityName="Mironov &amp; Galloway" authorityYear="2020" box="[1016,1294,1746,1770]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="884" phylum="Arthropoda" rank="species" species="sulcatus" status="sp. nov.">
<emphasis id="B908EAB6FFA71B442BBF40BAFAA0F902" box="[1016,1294,1746,1770]" italics="true" pageId="7" pageNumber="884">Nycteridocaulus sulcatus</emphasis>
</taxonomicName>
<emphasis id="B908EAB6FFA71B442D4840BAFAFAF902" bold="true" box="[1295,1364,1746,1770]" pageId="7" pageNumber="884">
<taxonomicNameLabel id="A23B57CDFFA71B442D4840BAFAFAF902" box="[1295,1364,1746,1770]" pageId="7" pageNumber="884" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, is close to 
<taxonomicName id="4C7C4D27FFA71B442A4E409DFC2CF8E4" authority="Hernandes, 2014" authorityName="Hernandes" authorityYear="2014" box="[521,898,1780,1804]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="884" phylum="Arthropoda" rank="species" species="guaratubensis">
<emphasis id="B908EAB6FFA71B442A4E409DFD68F8E4" box="[521,710,1780,1804]" italics="true" pageId="7" pageNumber="884">N. guaratubensis</emphasis>
Hernandes, 2014
</taxonomicName>
from the Restinga Tyrannulet, 
<taxonomicName id="4C7C4D27FFA71B442C9F409CFA18F8E4" authorityName="Willis &amp; Oniki" authorityYear="1992" box="[1240,1462,1780,1804]" class="Aves" family="Tyrannidae" genus="Phylloscartes" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="884" phylum="Chordata" rank="species" species="kronei">
<emphasis id="B908EAB6FFA71B442C9F409CFA18F8E4" box="[1240,1462,1780,1804]" italics="true" pageId="7" pageNumber="884">Phylloscartes kronei</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC336A4FFA71B4B29AD417FFB5AFE42" blockId="7.[490,1573,229,2045]" lastBlockId="8.[491,1557,229,495]" lastPageId="8" lastPageNumber="885" pageId="7" pageNumber="884">
Willis &amp; Oniki (Tyrannides: 
<taxonomicName id="4C7C4D27FFA71B442B75417FFC1DF8C7" box="[818,947,1815,1839]" class="Aves" family="Tyrannidae" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="884" phylum="Chordata" rank="family">Tyrannidae</taxonomicName>
), in having minute lacunae on the hysteronotal shield in both sexes, short and truncate opisthosomal lobes with nearly rectangular terminal lamellae and the adanal shields roughly L­shaped in males, and the primary spermaduct with ampuliform enlargement in females. 
<taxonomicName id="4C7C4D27FFA71B442B334116FB2BF87E" authorityName="Mironov &amp; Galloway" authorityYear="2020" box="[884,1157,1918,1942]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="884" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B908EAB6FFA71B442B334116FB2BF87E" box="[884,1157,1918,1942]" italics="true" pageId="7" pageNumber="884">Nycteridocaulus sulcatus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7C4D27FFA71B442D4E4117FA6AF87E" authorityName="Hernandes" authorityYear="2014" box="[1289,1476,1918,1942]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="884" phylum="Arthropoda" rank="species" species="guaratubensis">
<emphasis id="B908EAB6FFA71B442D4E4117FA6AF87E" box="[1289,1476,1918,1942]" italics="true" pageId="7" pageNumber="884">N. guaratubensis</emphasis>
</taxonomicName>
in the following features: in males, the supranal concavity is parallel­sided and open posteriorly, pregenital apodemes are present, and tarsus IV has a longitudinal dorsal ridge (
<figureCitation id="13472A21FFA71B442D3D41ABFA54F833" box="[1402,1530,1987,2011]" captionStart="Figure 1" captionStartId="4.[119,186,1883,1904]" captionTargetBox="[96,1502,306,1804]" captionTargetId="figure-2@4.[95,1502,299,1830]" captionTargetPageId="4" captionText="Figure 1 Nycteridocaulus sulcatussp. n., male: A – dorsal view, B – ventral view." figureDoi="http://doi.org/10.5281/zenodo.5757773" httpUri="https://zenodo.org/record/5757773/files/figure.png" pageId="7" pageNumber="884">Figures 1A</figureCitation>
, 
<figureCitation id="13472A21FFA71B4429AC418DFDA2F815" box="[491,524,2021,2045]" captionStart="Figure 3" captionStartId="6.[119,186,1911,1932]" captionTargetBox="[114,1477,238,1839]" captionTargetId="figure-2@6.[95,1502,226,1858]" captionTargetPageId="6" captionText="Figure 3 Nycteridocaulus sulcatussp. n., details: A–C – legs I–III of male, respectively, D – tibia and tarsus IV of male, E – tibia and tarsus IV of female, F – spermatheca and spermaducts, G – opisthosoma of male, ventral view. Abbreviations: ad– adanal shield, ga– genital arch, hs– head of spermatheca, pd– primary spermaduct, pg – pregenital apodemes, sd – secondary spermaducts, tl– terminal lamella." figureDoi="http://doi.org/10.5281/zenodo.5757777" httpUri="https://zenodo.org/record/5757777/files/figure.png" pageId="7" pageNumber="884">3G</figureCitation>
); in females, the posterior half of the hysteronotal shield bears sparsely disposed small circular lacunae (
<figureCitation id="13472A21FFA81B4B2AE4468DFCB4FF15" box="[675,794,229,253]" captionStart="Figure 2" captionStartId="5.[119,186,1722,1743]" captionTargetBox="[95,1502,251,1634]" captionTargetId="figure-2@5.[95,1502,221,1669]" captionTargetPageId="5" captionText="Figure 2 Nycteridocaulus sulcatussp. n., female: A – dorsal view, B – ventral view." figureDoi="http://doi.org/10.5281/zenodo.5757775" httpUri="https://zenodo.org/record/5757775/files/figure.png" pageId="8" pageNumber="885">Figure 2A</figureCitation>
), and setae 
<emphasis id="B908EAB6FFA81B4B2BD7468DFC05FF15" box="[912,939,229,253]" italics="true" pageId="8" pageNumber="885">h3</emphasis>
are 75–85 long, about 3/4 the length of the terminal appendages. In males of 
<taxonomicName id="4C7C4D27FFA81B4B2ABA4761FC1AFEC8" authorityName="Hernandes" authorityYear="2014" box="[765,948,264,288]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="885" phylum="Arthropoda" rank="species" species="guaratubensis">
<emphasis id="B908EAB6FFA81B4B2ABA4761FC1AFEC8" box="[765,948,264,288]" italics="true" pageId="8" pageNumber="885">N. guaratubensis</emphasis>
</taxonomicName>
, the supranal concavity is closed posteriorly and shaped as a long teardrop, pregenital apodemes are absent, and tarsus IV does not have a dorsal ridge; in females, the entire surface of the hysteronotal shield bears small circular lacunae and the posterior part has a pair of longitudinal rows consisting of 3 to 4 large ovate lacunae, and setae 
<emphasis id="B908EAB6FFA81B4B2A6047FAFDECFE42" box="[551,578,402,426]" italics="true" pageId="8" pageNumber="885">h3</emphasis>
are 66–72 long, about 1/2 the length of terminal appendages.
</paragraph>
</subSubSection>
<subSubSection id="C366652FFFA81B4B2A5447DDFB00FE07" pageId="8" pageNumber="885" type="etymology">
<paragraph id="8BC336A4FFA81B4B2A5447DDFB00FE07" blockId="8.[491,1557,229,495]" pageId="8" pageNumber="885">
<emphasis id="B908EAB6FFA81B4B2A5447DDFD20FE25" bold="true" box="[531,654,437,461]" pageId="8" pageNumber="885">Etymology</emphasis>
— The specific epithet 
<taxonomicName id="4C7C4D27FFA81B4B2BDD47DCFC5DFE24" authorityName="Mironov &amp; Galloway" authorityYear="2020" box="[922,1011,436,460]" class="Arachnida" family="Proctophyllodidae" genus="Nycteridocaulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="885" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B908EAB6FFA81B4B2BDD47DCFC5DFE24" box="[922,1011,436,460]" italics="true" pageId="8" pageNumber="885">sulcatus</emphasis>
</taxonomicName>
(from 
<emphasis id="B908EAB6FFA81B4B2C0747DCFB2CFE24" box="[1088,1154,436,460]" italics="true" pageId="8" pageNumber="885">sulcus</emphasis>
, groove, L.) is an adjective referring to the distinct groove­like supranal concavity in males.
</paragraph>
</subSubSection>
</treatment>
</document>