277 lines
36 KiB
XML
277 lines
36 KiB
XML
<document id="AD34869027D5A06548BCDF9C3EEEF6CD" ID-DOI="10.37828/em.2021.49.7" ID-ISSN="2336-9744" ID-Zenodo-Dep="13250302" ID-ZooBank="urn:lsid:zoobank.org:pub:602033D9-4D4E-402A-8828-E9184FA55C6C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722975773888" checkinUser="felipe" docAuthor="Mikhailov, Yuri, Simanchuk, Elena, Yunussova, Gulnara & Kazkenova, Gulzhan" docDate="2021" docId="03D58787220FFFBED18CFF460377FE18" docLanguage="en" docName="EcolMontenegrina.49.79-87.pdf" docOrigin="Ecologica Montenegrina 49" docSource="http://dx.doi.org/10.37828/em.2021.49.7" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Cryptocephalus (Asionus) ergenensis subsp. kalbensis Mikhailov 2021, ssp. nov." docType="treatment" docVersion="2" lastPageNumber="83" masterDocId="FFECFFFF220BFFBBD11BFF820137FFD2" masterDocTitle="A new subspecies of Сryptocephalus ergenensis Morawitz, 1863 (Coleoptera, Chrysomelidae) from Kazakhstan and significance of colour pattern polymorphism examination" masterLastPageNumber="87" masterPageNumber="79" pageNumber="83" updateTime="1722985653207" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="98BBB2CBAF7DB5E3219FB59AB590CAF5">A new subspecies of Сryptocephalus ergenensis Morawitz, 1863 (Coleoptera, Chrysomelidae) from Kazakhstan and significance of colour pattern polymorphism examination</mods:title>
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<mods:roleTerm id="1A4A5358C8BCCB7E08C3C277108A903A">Author</mods:roleTerm>
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<mods:namePart id="03D5D52FC13C2C04E3285830FC3B31F6">Mikhailov, Yuri</mods:namePart>
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<mods:affiliation id="D1E6C31B8931010F96A35D0332EBFC8B">Ural State Forest Engineering University, Sibirsky trakt 37, Yekaterinburg, 620100, Russia</mods:affiliation>
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<mods:nameIdentifier id="657D2022BA9FA9EE4EDDF719279B7FDB" type="email">yuemikhailov@gmail.com</mods:nameIdentifier>
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<mods:namePart id="960D4888627128810435F594182B37AB">Simanchuk, Elena</mods:namePart>
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<mods:affiliation id="0FBA3E8D63F96C86C6924306D4D030D3">A. Baitursynov Kostanay Regional University, Baitursynov street 47, Kostanay, 110000, Kazakhstan * Corresponding Author. E-mail: yuemikhailov @ gmail. com</mods:affiliation>
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<mods:namePart id="4230FB89679ACB88DB468BB6534139B4">Yunussova, Gulnara</mods:namePart>
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<mods:affiliation id="D465A0994221E8BC083D6EE21F5C6C8E">A. Baitursynov Kostanay Regional University, Baitursynov street 47, Kostanay, 110000, Kazakhstan * Corresponding Author. E-mail: yuemikhailov @ gmail. com</mods:affiliation>
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<mods:namePart id="CE656DD076648E6463566E84E8A0D740">Kazkenova, Gulzhan</mods:namePart>
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<mods:affiliation id="6C20400B8567977C3063E7B0B373ABB0">A. Baitursynov Kostanay Regional University, Baitursynov street 47, Kostanay, 110000, Kazakhstan * Corresponding Author. E-mail: yuemikhailov @ gmail. com</mods:affiliation>
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<mods:title id="361AB4B4F6219E9CDFC5B74A792931D7">Ecologica Montenegrina</mods:title>
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<mods:date id="C62A35EAE95D4861F2928C935FDDE87F">2021</mods:date>
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<mods:number id="6BC0E54765B81BC84523F35EE55E608D">49</mods:number>
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<treatment id="03D58787220FFFBED18CFF460377FE18" LSID="urn:lsid:plazi:treatment:03D58787220FFFBED18CFF460377FE18" httpUri="http://treatment.plazi.org/id/03D58787220FFFBED18CFF460377FE18" lastPageId="5" lastPageNumber="83" pageId="4" pageNumber="83">
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<subSubSection id="C366651A220FFFBFD18CFF460566FF0C" box="[151,1105,195,222]" pageId="4" pageNumber="83" type="nomenclature">
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<paragraph id="8BC33691220FFFBFD18CFF460566FF0C" blockId="4.[151,1105,195,222]" box="[151,1105,195,222]" pageId="4" pageNumber="83">
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<heading id="D08B81FD220FFFBFD18CFF460566FF0C" box="[151,1105,195,222]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="83" reason="2">
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<taxonomicName id="4C7C4D12220FFFBFD18CFF460262FF0C" authority="Mikhailov" authorityName="Mikhailov" authorityYear="2021" box="[151,853,195,222]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="83" phylum="Arthropoda" rank="subSpecies" species="ergenensis" status="subsp. nov." subGenus="Asionus" subSpecies="kalbensis">
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<emphasis id="B908EA83220FFFBFD18CFF4603E6FF0C" bold="true" box="[151,721,196,222]" pageId="4" pageNumber="83">
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<emphasis id="B908EA83220FFFBFD18CFF460063FF0C" bold="true" box="[151,340,196,222]" italics="true" pageId="4" pageNumber="83">Cryptocephalus</emphasis>
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(
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||
<emphasis id="B908EA83220FFFBFD07EFF4600F0FF0C" bold="true" box="[357,455,196,222]" italics="true" pageId="4" pageNumber="83">Asionus</emphasis>
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)
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||
<emphasis id="B908EA83220FFFBFD0C3FF4603E6FF0C" bold="true" box="[472,721,196,222]" italics="true" pageId="4" pageNumber="83">ergenensis kalbensis</emphasis>
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</emphasis>
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||
Mikhailov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23B57F8220FFFBFD27AFF4602FEFF0C" box="[865,969,196,222]" pageId="4" pageNumber="83" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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(
|
||
<figureCitation id="13472A14220FFFBFD2C3FF410570FF0C" box="[984,1095,195,222]" captionStart="Figures 1–10" captionStartId="2.[151,237,1310,1334]" captionTargetBox="[231,1400,210,1257]" captionTargetId="figure-20@2.[199,1432,189,1292]" captionTargetPageId="2" captionText="Figures 1–10. Cryptocephalus (Asionus) spp., habitus. 1-4 – С. ergenensis ergenensis Morawitz: 1, 2 – female, SW Russia, Astrakhan region, env. of Baskunchak lake; 3 – male, West Kazakhstan, Atyrau region, Deukara; 4 – male, South Kazakhstan, Kyzylorda; 5 – C. lateralis Suffrian, female, NW Kazakhstan, Aktobe region, Martuk; 6-8 – С. ergenensis kalbensis ssp. n.: 6, 7 – female, paratype, East Kazakhstan, south spurs of Kalbinsky mt. range, 12 km SW Samarskoye; 8 – male, holotype, East Kazakhstan, 13 km SW Samarskoye; 9, 10 – С. apicalis Gebler, female, East Kazakhstan, west spurs of Narymsky mt. range, Kaiyndy. 2, 7 and 9 – lateral view, others – dorsal view. Scale bar – 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13250306" httpUri="https://zenodo.org/record/13250306/files/figure.png" pageId="4" pageNumber="83">Figs. 6–8</figureCitation>
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)
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C366651A220FFFBFD18CFE850234FDE2" pageId="4" pageNumber="83" type="materials_examined">
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<paragraph id="8BC33691220FFFBFD18CFE850234FDE2" blockId="4.[151,1436,263,560]" pageId="4" pageNumber="83">
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<emphasis id="B908EA83220FFFBFD18CFE850063FEF3" bold="true" box="[151,340,263,289]" pageId="4" pageNumber="83">Type material:</emphasis>
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<materialsCitation id="3B143CCC220FFFBFD045FE8505EFFE55" collectingDate="2019-06-02" collectionCode="ISEA" collectorName="A. A. Gurina & R. Yu. Dudko" country="Kazakhstan" county="Dudko" elevation="575" latitude="48.97167" location="Aksenovka river" longLatPrecision="110" longitude="83.191666" municipality="Kalbinsky mt. range" pageId="4" pageNumber="83" specimenCount="1" stateProvince="Shyghys Qazaqstan Oblysy" typeStatus="holotype">
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<typeStatus id="54C78833220FFFBFD045FE8500F9FEF0" box="[350,462,263,290]" pageId="4" pageNumber="83" type="holotype">Holotype</typeStatus>
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, male (
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<collectionCode id="ED6DAE54220FFFBFD33CFE850359FEF0" box="[551,622,263,290]" pageId="4" pageNumber="83">ISEA</collectionCode>
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), with labels: 1) Вост.
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<collectingCountry id="F36B7601220FFFBFD29CFE85053EFEF0" box="[903,1033,263,290]" name="Kazakhstan" pageId="4" pageNumber="83">КаЗахстан</collectingCountry>
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||
, Ю отроги Калбинского хр., 13 км ЮЗ с. Самарское, дол. р. Аксеновки, 550-600 м,
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||
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,
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<geoCoordinate id="EE485056220FFFBFD2D3FEAA057FFE91" box="[968,1096,296,323]" degrees="83" direction="east" minutes="11.5" orientation="longitude" pageId="4" pageNumber="83" precision="92" value="83.191666">83°11.5' E</geoCoordinate>
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, 1-
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<date id="FFC21051220FFFBFD56BFEAA05DFFE91" box="[1136,1256,296,323]" pageId="4" pageNumber="83" value="2019-06-02">
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<collectingDate id="EF86E9B9220FFFBFD56BFEAA05DFFE91" box="[1136,1256,296,323]" pageId="4" pageNumber="83" value="2019-06-02">2.06.2019</collectingDate>
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</date>
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, А.А. Гурина, Р.Ю.
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<collectingCounty id="62A24E1D220FFFBFD1C7FEC9001CFEB7" box="[220,299,331,357]" pageId="4" pageNumber="83">Дудко</collectingCounty>
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[
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<collectingRegion id="49B8F873220FFFBFD023FEC90336FEB7" box="[312,513,330,357]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="4" pageNumber="83">East Kazakhstan</collectingRegion>
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, south spurs of
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<collectingMunicipality id="6BA7ACEB220FFFBFD3A2FEC8029DFEB7" box="[697,938,330,357]" pageId="4" pageNumber="83">Kalbinsky mt. range</collectingMunicipality>
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,
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<locationDeviation id="9A712FB3220FFFBFD2ADFEC805FAFEB7" box="[950,1229,330,357]" location="Samarskoye" pageId="4" pageNumber="83">
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SW Samarskoye
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</locationDeviation>
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,
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<location id="8EA3604A220FFFBFD5C2FEC804ABFEB7" LSID="urn:lsid:plazi:treatment:03D58787220FFFBED18CFF460377FE18:8EA3604A220FFFBFD5C2FEC804ABFEB7" box="[1241,1436,330,357]" country="Kazakhstan" county="Dudko" latitude="48.97167" longLatPrecision="110" longitude="83.191666" municipality="Kalbinsky mt. range" name="Aksenovka river" pageId="4" pageNumber="83" stateProvince="Shyghys Qazaqstan Oblysy">Aksenovka river</location>
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valley,
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<quantity id="4C849B74220FFFBFD1E8FEEE004CFE55" box="[243,379,364,391]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="4" pageNumber="83" unit="m" value="575.0" valueMax="600.0" valueMin="550.0">
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<elevation id="0051D1A2220FFFBFD1E8FEEE004CFE55" box="[243,379,364,391]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="4" pageNumber="83" unit="m" value="575.0" valueMax="600.0" valueMin="550.0">550-600 m</elevation>
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</quantity>
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,
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,
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<geoCoordinate id="EE485056220FFFBFD33FFEEE0390FE55" box="[548,679,364,391]" degrees="83" direction="east" minutes="11.5" orientation="longitude" pageId="4" pageNumber="83" precision="92" value="83.191666">83°11.5' E</geoCoordinate>
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, 1-
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<date id="FFC21051220FFFBFD3C9FEEE027DFE55" box="[722,842,364,391]" pageId="4" pageNumber="83" value="2019-06-02">
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<collectingDate id="EF86E9B9220FFFBFD3C9FEEE027DFE55" box="[722,842,364,391]" pageId="4" pageNumber="83" value="2019-06-02">2.06.2019</collectingDate>
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,
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<collectorName id="26895347220FFFBFD247FEEE02C0FE55" box="[860,1015,364,391]" pageId="4" pageNumber="83">A.A. Gurina</collectorName>
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,
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<collectorName id="26895347220FFFBFD513FEEF0587FE55" box="[1032,1200,364,391]" pageId="4" pageNumber="83">R.Yu. Dudko</collectorName>
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], 2
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)
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<typeStatus id="54C78833220FFFBFD5F5FEEE04ABFE55" box="[1262,1436,364,391]" pageId="4" pageNumber="83" type="holotype">HOLOTYPUS</typeStatus>
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<emphasis id="B908EA83220FFFBFD18CFE0C0066FE7B" box="[151,337,398,425]" italics="true" pageId="4" pageNumber="83">Сryptocephalus</emphasis>
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<taxonomicName id="4C7C4D12220FFFBFD044FE0C036CFE7A" authorityName="Morawitz" authorityYear="1863" box="[351,603,398,425]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="83" phylum="Arthropoda" rank="species" species="ergenensis" subGenus="Asionus">
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(
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||
<emphasis id="B908EA83220FFFBFD072FE0C00F1FE7A" box="[361,454,398,424]" italics="true" pageId="4" pageNumber="83">Asionus</emphasis>
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)
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<emphasis id="B908EA83220FFFBFD0C6FE0D036CFE7A" box="[477,603,399,424]" italics="true" pageId="4" pageNumber="83">ergenensis</emphasis>
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</taxonomicName>
|
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<emphasis id="B908EA83220FFFBFD372FE0C03E1FE7B" box="[617,726,398,425]" italics="true" pageId="4" pageNumber="83">kalbensis</emphasis>
|
||
ssp.n. Yu. Mikhailov design. 2020.
|
||
<materialsCitation id="3B143CCC220FFFBFD5B3FE0D03B9FE18" collectingDate="2019-06-02" collectionCode="YMC" collectorName="A. A. Gurina & R. Yu. Dudko" country="Kazakhstan" county="Dudko" elevation="575" latitude="48.97167" location="Aksenovka river" longLatPrecision="110" longitude="83.191666" municipality="Kalbinsky mt. range" pageId="4" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="Shyghys Qazaqstan Oblysy" typeStatus="paratype">
|
||
<typeStatus id="54C78833220FFFBFD5B3FE0D0429FE7B" box="[1192,1310,399,425]" pageId="4" pageNumber="83" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D7AFD18220FFFBFD42BFE0C04ABFE7B" box="[1328,1436,398,425]" count="1" pageId="4" pageNumber="83" type="female">1 female</specimenCount>
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||
(
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||
<collectionCode id="ED6DAE54220FFFBFD184FE2D01D2FE18" box="[159,229,431,458]" pageId="4" pageNumber="83">YMC</collectionCode>
|
||
) collected together with holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B143CCC220FFFBFD3BAFE2D03C8FDE2" collectingDate="2019-06-02" collectionCode="YMC" collectorName="A. A. Gurina & R. Yu. Dudko" country="Kazakhstan" elevation="575" latitude="48.974167" location="Kalbinsky mt. range" longLatPrecision="11" longitude="83.19767" municipality="Dudko" pageId="4" pageNumber="83" specimenCount="2" specimenCount-female="2" stateProvince="Shyghys Qazaqstan Oblysy" typeStatus="paratype">
|
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<specimenCount id="9D7AFD18220FFFBFD3BAFE2D022FFE18" box="[673,792,431,458]" count="2" pageId="4" pageNumber="83" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED6DAE54220FFFBFD235FE2D0243FE18" box="[814,884,431,458]" pageId="4" pageNumber="83">YMC</collectionCode>
|
||
) with labels: Вост.
|
||
<collectingCountry id="F36B7601220FFFBFD59CFE2D043EFE18" box="[1159,1289,431,458]" name="Kazakhstan" pageId="4" pageNumber="83">КаЗахстан</collectingCountry>
|
||
, Ю отроги Калбинского хр., 12 км ЮЗ с. Самарское, 575 м,
|
||
<geoCoordinate id="EE485056220FFFBFD3E7FE5302BCFE3E" box="[764,907,465,492]" degrees="48" direction="north" minutes="58.45" orientation="latitude" pageId="4" pageNumber="83" precision="9" value="48.974167">48°58.45' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE485056220FFFBFD28CFE530514FE3E" box="[919,1059,465,492]" degrees="83" direction="east" minutes="11.86" orientation="longitude" pageId="4" pageNumber="83" precision="9" value="83.19767">83°11.86' E</geoCoordinate>
|
||
,
|
||
<date id="FFC21051220FFFBFD52BFE53059FFE3E" box="[1072,1192,465,492]" pageId="4" pageNumber="83" value="2019-06-02">
|
||
<collectingDate id="EF86E9B9220FFFBFD52BFE53059FFE3E" box="[1072,1192,465,492]" pageId="4" pageNumber="83" value="2019-06-02">2.06.2019</collectingDate>
|
||
</date>
|
||
, А.А. Гурина, Р.Ю.
|
||
<collectingMunicipality id="6BA7ACEB220FFFBFD18CFE7601D1FDDC" box="[151,230,500,526]" pageId="4" pageNumber="83">Дудко</collectingMunicipality>
|
||
[
|
||
<collectingRegion id="49B8F873220FFFBFD1EEFE7600F7FDDC" box="[245,448,499,526]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="4" pageNumber="83">East Kazakhstan</collectingRegion>
|
||
, south spurs of
|
||
<location id="8EA3604A220FFFBFD365FE710244FDDC" LSID="urn:lsid:plazi:treatment:03D58787220FFFBED18CFF460377FE18:8EA3604A220FFFBFD365FE710244FDDC" box="[638,883,499,526]" country="Kazakhstan" latitude="48.974167" longLatPrecision="11" longitude="83.19767" municipality="Dudko" name="Kalbinsky mt. range" pageId="4" pageNumber="83" stateProvince="Shyghys Qazaqstan Oblysy">Kalbinsky mt. range</location>
|
||
,
|
||
<locationDeviation id="9A712FB3220FFFBFD29BFE7105AAFDDC" box="[896,1181,499,526]" location="Samarskoye" pageId="4" pageNumber="83">
|
||
<quantity id="4C849B74220FFFBFD29BFE7102FCFDDC" box="[896,971,499,526]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="83" unit="km" value="12.0">12 km</quantity>
|
||
SW Samarskoye
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4C849B74220FFFBFD5B2FE7105CFFDDC" box="[1193,1272,499,526]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="4" pageNumber="83" unit="m" value="575.0">
|
||
<elevation id="0051D1A2220FFFBFD5B2FE7105CFFDDC" box="[1193,1272,499,526]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="4" pageNumber="83" unit="m" value="575.0">575 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE485056220FFFBFD41EFE7104A0FDDC" box="[1285,1431,499,526]" degrees="48" direction="north" minutes="58.45" orientation="latitude" pageId="4" pageNumber="83" precision="9" value="48.974167">48°58.45' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE485056220FFFBFD18CFD970015FDE2" box="[151,290,533,560]" degrees="83" direction="east" minutes="11.86" orientation="longitude" pageId="4" pageNumber="83" precision="9" value="83.19767">83°11.86' E</geoCoordinate>
|
||
,
|
||
<date id="FFC21051220FFFBFD034FD970090FDE2" box="[303,423,533,560]" pageId="4" pageNumber="83" value="2019-06-02">
|
||
<collectingDate id="EF86E9B9220FFFBFD034FD970090FDE2" box="[303,423,533,560]" pageId="4" pageNumber="83" value="2019-06-02">2.06.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26895347220FFFBFD0AFFD97037EFDE2" box="[436,585,533,560]" pageId="4" pageNumber="83">A.A. Gurina</collectorName>
|
||
,
|
||
<collectorName id="26895347220FFFBFD34FFD9403CEFDE2" box="[596,761,533,560]" pageId="4" pageNumber="83">R.Yu. Dudko</collectorName>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651A220FFFBED18CFDDB0377FE18" lastPageId="5" lastPageNumber="84" pageId="4" pageNumber="83" type="description">
|
||
<paragraph id="8BC33691220FFFBFD18CFDDB02D5FD47" blockId="4.[151,1437,600,1909]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908EA83220FFFBFD18CFDDB00F5FDA1" bold="true" box="[151,450,601,627]" pageId="4" pageNumber="83">
|
||
Description of
|
||
<typeStatus id="54C78833220FFFBFD04CFDDB00F5FDA1" box="[343,450,601,627]" pageId="4" pageNumber="83" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(male). Measurements. Body length
|
||
<quantity id="4C849B74220FFFBFD292FDDA02DEFDA1" box="[905,1001,600,627]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="83" unit="mm" value="4.4">4.4 mm</quantity>
|
||
, width of elytra at humeri
|
||
<quantity id="4C849B74220FFFBFD42DFDDA04A0FDA1" box="[1334,1431,600,627]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="83" unit="mm" value="2.3">2.3 mm</quantity>
|
||
; length of elytra
|
||
<quantity id="4C849B74220FFFBFD049FDF80098FD47" box="[338,431,634,661]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="83" unit="mm" value="3.1">3.1 mm</quantity>
|
||
; length of pronotum
|
||
<quantity id="4C849B74220FFFBFD3BFFDF803C9FD47" box="[676,766,634,661]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="83" unit="mm" value="1.4">1.4 mm</quantity>
|
||
and width
|
||
<quantity id="4C849B74220FFFBFD29AFDF802E9FD47" box="[897,990,634,661]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="83" unit="mm" value="2.1">2.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC33691220FFFBFD1EDFD1E027EFC71" blockId="4.[151,1437,600,1909]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908EA83220FFFBFD1EDFD1E0001FD65" box="[246,310,668,695]" italics="true" pageId="4" pageNumber="83">Head</emphasis>
|
||
. Shiny, blackish green with small yellow maculae on genae. Surface of frons densely covered with small punctures and longitudinal wrinkles, with short white setae. Labrum blackish green; short, transverse, almost impunctate, anterior angles rounded, anterior margin shallowly convex. Mandibles black. Terminal palpomere of labial palpi acute. Eyes medium-sized, evenly convex, upper lobes distant, distance between them 1.2 times eye length in lateral view. Antennae 0.7× body length. Antennomere I from above, antennomeres II to IV entirely, base of antennomere V rufous; antennomeres III and IV elongate, narrow, ratio ratio of their length 0.9: 1.1; antennomere V similar to IV, but less widened at apex; antennomeres VI to IX distinctly widened; antennomere XI with pointed tip.
|
||
</paragraph>
|
||
<paragraph id="8BC33691220FFFBFD1EDFC2B0290FAE5" blockId="4.[151,1437,600,1909]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908EA83220FFFBFD1EDFC2B007CFC16" box="[246,331,937,964]" italics="true" pageId="4" pageNumber="83">Thorax</emphasis>
|
||
. Pronotum blackish green, shiny and covered with dense oblong middle sized punctures, its anterior and posterior angles and two small narrow transverse maculae before scutellum yellow (
|
||
<figureCitation id="13472A14220FFFBFD45AFC4904BAFC34" box="[1345,1421,971,998]" captionStart="Figures 1–10" captionStartId="2.[151,237,1310,1334]" captionTargetBox="[231,1400,210,1257]" captionTargetId="figure-20@2.[199,1432,189,1292]" captionTargetPageId="2" captionText="Figures 1–10. Cryptocephalus (Asionus) spp., habitus. 1-4 – С. ergenensis ergenensis Morawitz: 1, 2 – female, SW Russia, Astrakhan region, env. of Baskunchak lake; 3 – male, West Kazakhstan, Atyrau region, Deukara; 4 – male, South Kazakhstan, Kyzylorda; 5 – C. lateralis Suffrian, female, NW Kazakhstan, Aktobe region, Martuk; 6-8 – С. ergenensis kalbensis ssp. n.: 6, 7 – female, paratype, East Kazakhstan, south spurs of Kalbinsky mt. range, 12 km SW Samarskoye; 8 – male, holotype, East Kazakhstan, 13 km SW Samarskoye; 9, 10 – С. apicalis Gebler, female, East Kazakhstan, west spurs of Narymsky mt. range, Kaiyndy. 2, 7 and 9 – lateral view, others – dorsal view. Scale bar – 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13250306" httpUri="https://zenodo.org/record/13250306/files/figure.png" pageId="4" pageNumber="83">Fig. 8</figureCitation>
|
||
). Pronotum transverse, 1.5 times broader than long, widest at basal third, strongly convex. Basal margin feebly sinuose, evenly covered with small homogenous teeth without toothless sections; posterior angles triangularly extended, bearing setiferous pore and long seta; median lobe of basal margin emarginate and slightly raised; anterior and lateral margins narrowly bordered, not visible in dorsal view; anterior angles rectangular. Thorax ventrally black with white setae. Scutellum triangular, elongate, with narrowly rounded apex, covered with very small sparse punctures. Elytra subcylindrical, widened in middle in dorsal view, 1.35 times as long as wide at humeral part; glabrous, lustrous, blackish green with epipleura and broad lateral band yellow; punctures large, dense and entirely confused, intervals slightly convex. Punctures along anterior edge and on apex of elytra smaller and sparser, broad yellow lateral band almost devoid of punctures. Epipleuron reaching 4/5 of length of elytra, impunctate.
|
||
</paragraph>
|
||
<paragraph id="8BC33691220FFFBFD1EDFAC2028CFA32" blockId="4.[151,1437,600,1909]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908EA83220FFFBFD1EDFAC20019FA8B" box="[246,302,1344,1369]" italics="true" pageId="4" pageNumber="83">Legs</emphasis>
|
||
blackish green with large yellow maculae at apices of middle and hind femora, small yellow maculae at apices of fore femora; fore tibiae as long as posterior tibiae, straight. Protarsomere I elongate and slightly broadened while mesotarsomere I broad and almost twice as broader as II. Ratio of length of protarsomeres: 1: 0.6: 0.6: 0.8; metatarsomeres: 1: 0.5: 0.5: 0.8; metatarsomere V narrow, more than half its length projecting out of lobes of metatarsomere III, claws simple.
|
||
</paragraph>
|
||
<paragraph id="8BC33691220FFFBFD1EDFA6501C9F9B5" blockId="4.[151,1437,600,1909]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908EA83220FFFBFD1EDFA650052F9D0" box="[246,357,1511,1538]" italics="true" pageId="4" pageNumber="83">Abdomen</emphasis>
|
||
. Ventrites blackish green, ventrites and pygidium with short white setae; last ventrite with flat longitudinal median area and weak transverse depression on it, its base with small laterally compressed denticle in a middle; pygidium black with apical margin yellow. Aedeagus as in nominotypical subspecies (
|
||
<figureCitation id="13472A14220FFFBFD184F9CE01C7F9B5" box="[159,240,1612,1639]" captionStart="Figures 11–14" captionStartId="3.[151,237,1906,1930]" captionTargetBox="[201,1325,669,1807]" captionTargetId="figure-202@3.[151,1436,627,1866]" captionTargetPageId="3" captionText="Figures 11–14. Aedeagi of Cryptocephalus (Asionus) spp in lateral and dorsal view. 11 – С. ergenensis ergenensis, West Kazakhstan, Atyrau region, Deukara; 12 – С. ergenensis kalbensis ssp. n., type locality; 13 – С. apicalis, East Kazakhstan, Kalbinsky mt. range, near Nizhniy Tainty; 14 – С. lateralis. Scale bar – 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13250308" httpUri="https://zenodo.org/record/13250308/files/figure.png" pageId="4" pageNumber="83">Fig 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC33691220FFFBFD1EDF9EC0068F95B" blockId="4.[151,1437,600,1909]" box="[246,351,1646,1673]" pageId="4" pageNumber="83">Females.</paragraph>
|
||
<paragraph id="8BC33691220FFFBFD1EDF9120393F8A7" blockId="4.[151,1437,600,1909]" pageId="4" pageNumber="83">
|
||
Body length 5.0–
|
||
<quantity id="4C849B74220FFFBFD0D1F912031DF979" box="[458,554,1680,1707]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="83" unit="mm" value="5.2">5.2 mm</quantity>
|
||
, width
|
||
<quantity id="4C849B74220FFFBFD391F9120228F979" box="[650,799,1680,1707]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="4" pageNumber="83" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
|
||
. Colour pattern similar to
|
||
<typeStatus id="54C78833220FFFBFD573F91205F8F979" box="[1128,1231,1680,1707]" pageId="4" pageNumber="83" type="holotype">holotype</typeStatus>
|
||
but rarely small yellow maculae may appear at sides of front near eyes and on sides of clypeus; maculae at basal angles of pronotum larger and rectangular, narrow band appear along fore margin (
|
||
<figureCitation id="13472A14220FFFBFD50CF95105B0F93C" box="[1047,1159,1747,1774]" captionStart="Figures 1–10" captionStartId="2.[151,237,1310,1334]" captionTargetBox="[231,1400,210,1257]" captionTargetId="figure-20@2.[199,1432,189,1292]" captionTargetPageId="2" captionText="Figures 1–10. Cryptocephalus (Asionus) spp., habitus. 1-4 – С. ergenensis ergenensis Morawitz: 1, 2 – female, SW Russia, Astrakhan region, env. of Baskunchak lake; 3 – male, West Kazakhstan, Atyrau region, Deukara; 4 – male, South Kazakhstan, Kyzylorda; 5 – C. lateralis Suffrian, female, NW Kazakhstan, Aktobe region, Martuk; 6-8 – С. ergenensis kalbensis ssp. n.: 6, 7 – female, paratype, East Kazakhstan, south spurs of Kalbinsky mt. range, 12 km SW Samarskoye; 8 – male, holotype, East Kazakhstan, 13 km SW Samarskoye; 9, 10 – С. apicalis Gebler, female, East Kazakhstan, west spurs of Narymsky mt. range, Kaiyndy. 2, 7 and 9 – lateral view, others – dorsal view. Scale bar – 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13250306" httpUri="https://zenodo.org/record/13250306/files/figure.png" pageId="4" pageNumber="83">Figs. 6-7</figureCitation>
|
||
). Apices of all femora with broad yellow bands, all tibiae and tarsi rufous, only middle of inner surface of fore tibiae and apices of tarsomeres dark. Pronotum 1.6 times as broader as long, strongly convex, shining, covered with dense oblong middle sized punctures. Scutellum covered with very small sparse punctures. Elytra 2.2 times as longer as pronotum, puncturation as in male.
|
||
</paragraph>
|
||
<paragraph id="8BC33691220FFFBFD18CF81D0005F86B" blockId="4.[151,1436,1951,2044]" box="[151,306,1951,1977]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908EA83220FFFBFD18CF81D0005F86B" bold="true" box="[151,306,1951,1977]" pageId="4" pageNumber="83">Comparison</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC33691220FFFBED18CF83D0377FE18" blockId="4.[151,1436,1951,2044]" lastBlockId="5.[151,1436,195,458]" lastPageId="5" lastPageNumber="84" pageId="4" pageNumber="83">
|
||
Nominotypical subspecies has pronotum and elytra with contrasting colouration (
|
||
<figureCitation id="13472A14220FFFBFD583F83D043BF808" box="[1176,1292,1983,2010]" captionStart="Figures 1–10" captionStartId="2.[151,237,1310,1334]" captionTargetBox="[231,1400,210,1257]" captionTargetId="figure-20@2.[199,1432,189,1292]" captionTargetPageId="2" captionText="Figures 1–10. Cryptocephalus (Asionus) spp., habitus. 1-4 – С. ergenensis ergenensis Morawitz: 1, 2 – female, SW Russia, Astrakhan region, env. of Baskunchak lake; 3 – male, West Kazakhstan, Atyrau region, Deukara; 4 – male, South Kazakhstan, Kyzylorda; 5 – C. lateralis Suffrian, female, NW Kazakhstan, Aktobe region, Martuk; 6-8 – С. ergenensis kalbensis ssp. n.: 6, 7 – female, paratype, East Kazakhstan, south spurs of Kalbinsky mt. range, 12 km SW Samarskoye; 8 – male, holotype, East Kazakhstan, 13 km SW Samarskoye; 9, 10 – С. apicalis Gebler, female, East Kazakhstan, west spurs of Narymsky mt. range, Kaiyndy. 2, 7 and 9 – lateral view, others – dorsal view. Scale bar – 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13250306" httpUri="https://zenodo.org/record/13250306/files/figure.png" pageId="4" pageNumber="83">Figs. 1-4</figureCitation>
|
||
), pronotum covered with small and sparse punctures, scutellum impunctate, elytra with sparse middle sized punctures. New subspecies has predominantly blackish green dorsum with broad lateral yellow emargination of the elytra, dense oblong middle sized punctures on the pronotum, small sparse punctures on the scutellum and dense large punctures on the elytra.
|
||
<emphasis id="B908EA83220EFFBED35CFE8A03FFFEF0" box="[583,712,263,290]" italics="true" pageId="5" pageNumber="84">
|
||
С.
|
||
<taxonomicName id="4C7C4D12220EFFBED370FE8503FFFEF0" authorityName="Gebler" authorityYear="1830" box="[619,712,263,290]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="apicalis">apicalis</taxonomicName>
|
||
</emphasis>
|
||
, which lives in sympatry with the new subspecies, differs in aedeagus shape (
|
||
<figureCitation id="13472A14220EFFBED044FEAA0081FE91" box="[351,438,296,323]" captionStart="Figures 11–14" captionStartId="3.[151,237,1906,1930]" captionTargetBox="[201,1325,669,1807]" captionTargetId="figure-202@3.[151,1436,627,1866]" captionTargetPageId="3" captionText="Figures 11–14. Aedeagi of Cryptocephalus (Asionus) spp in lateral and dorsal view. 11 – С. ergenensis ergenensis, West Kazakhstan, Atyrau region, Deukara; 12 – С. ergenensis kalbensis ssp. n., type locality; 13 – С. apicalis, East Kazakhstan, Kalbinsky mt. range, near Nizhniy Tainty; 14 – С. lateralis. Scale bar – 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13250308" httpUri="https://zenodo.org/record/13250308/files/figure.png" pageId="5" pageNumber="84">Fig. 13</figureCitation>
|
||
) and has yellow lateral emargination of the elytra clearly interrupted behind middle (
|
||
<figureCitation id="13472A14220EFFBED1BBFEC801CDFEB7" box="[160,250,330,357]" captionStart="Figures 1–10" captionStartId="2.[151,237,1310,1334]" captionTargetBox="[231,1400,210,1257]" captionTargetId="figure-20@2.[199,1432,189,1292]" captionTargetPageId="2" captionText="Figures 1–10. Cryptocephalus (Asionus) spp., habitus. 1-4 – С. ergenensis ergenensis Morawitz: 1, 2 – female, SW Russia, Astrakhan region, env. of Baskunchak lake; 3 – male, West Kazakhstan, Atyrau region, Deukara; 4 – male, South Kazakhstan, Kyzylorda; 5 – C. lateralis Suffrian, female, NW Kazakhstan, Aktobe region, Martuk; 6-8 – С. ergenensis kalbensis ssp. n.: 6, 7 – female, paratype, East Kazakhstan, south spurs of Kalbinsky mt. range, 12 km SW Samarskoye; 8 – male, holotype, East Kazakhstan, 13 km SW Samarskoye; 9, 10 – С. apicalis Gebler, female, East Kazakhstan, west spurs of Narymsky mt. range, Kaiyndy. 2, 7 and 9 – lateral view, others – dorsal view. Scale bar – 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13250306" httpUri="https://zenodo.org/record/13250306/files/figure.png" pageId="5" pageNumber="84">Fig. 10</figureCitation>
|
||
). From smaller
|
||
<emphasis id="B908EA83220EFFBED0A2FEC90308FEB7" box="[441,575,330,357]" italics="true" pageId="5" pageNumber="84">
|
||
С.
|
||
<taxonomicName id="4C7C4D12220EFFBED0C6FEC80308FEB7" authorityName="Suffrian" authorityYear="1853" box="[477,575,330,357]" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="lateralis">lateralis</taxonomicName>
|
||
</emphasis>
|
||
easily differs by larger body size. From
|
||
<emphasis id="B908EA83220EFFBED530FEC90433FEB7" box="[1067,1284,330,357]" italics="true" pageId="5" pageNumber="84">С. pseudolateralis</emphasis>
|
||
, known also from
|
||
<collectingRegion id="49B8F873220EFFBED1C1FEEF0097FE55" box="[218,416,364,391]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="5" pageNumber="84">East Kazakhstan</collectingRegion>
|
||
and having similar colour pattern differs in small laterally compressed denticle in a middle of last ventrite of males and aedeagus shape. The key below helps to distinguish the new taxon among the respective species group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |