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<document id="F0C1413EF534D3B8F81798DE613D0066" ID-CLB-Dataset="107002" ID-DOI="10.11646/zootaxa.5168.5.1" ID-GBIF-Dataset="b0039631-f341-4018-b9b6-657063012307" ID-ISSN="1175-5326" ID-Zenodo-Dep="6911501" ID-ZooBank="121EA9DC-F17B-46FD-8987-F879BC6DAA99" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658902027587" checkinUser="plazi" docAuthor="Cunha, Eduardo Victor De Paiva &amp; Fernandes, José Antônio Marin" docDate="2022" docId="03C787C3FFDCFFF3C2ABFC14FA57FDB3" docLanguage="en" docName="zootaxa.5168.5.1.pdf" docOrigin="Zootaxa 5168 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypoxys triangularis Nunes et al. 2020" docType="treatment" docVersion="5" lastPageNumber="504" masterDocId="FFFEFFBBFFDDFFF0C23CFFB1FFCBFF99" masterDocTitle="New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot &amp; Serville, 1843 (Hemiptera: Pentatomidae: Edessinae)" masterLastPageNumber="520" masterPageNumber="501" pageNumber="502" updateTime="1699362273591" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7665DBB1D42EB2DD038F0504EDC7189E">New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot &amp; Serville, 1843 (Hemiptera: Pentatomidae: Edessinae)</mods:title>
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<mods:namePart id="B1E00326BA2F6F15AD51D365D4CCDDCE">Cunha, Eduardo Victor De Paiva</mods:namePart>
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<mods:affiliation id="D6BC8976ECFD64FE59B795399C23E8F1">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1, Belém, Pará, Brazil 66075 - 110. &amp; eduardovdpc @ gmail. com; https: // orcid. org / 0000 - 0002 - 2386 - 6734</mods:affiliation>
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<mods:namePart id="A1BAB18441F35C1A7B98AE842054908A">Fernandes, José Antônio Marin</mods:namePart>
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<mods:affiliation id="C578CA1CA09A4E53720CCE0DA74961F7">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1, Belém, Pará, Brazil 66075 - 110. &amp; joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
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<emphasis id="B91AEAC7FFDCFFF1C2ABFC14FE5DFC26" bold="true" box="[151,406,932,959]" italics="true" pageId="1" pageNumber="502">Hypoxys triangularis</emphasis>
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Group
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<paragraph id="8BD136D5FFDCFFF1C2ABFC79FEEBFC7A" blockId="1.[151,497,932,995]" box="[151,288,968,995]" pageId="1" pageNumber="502">
(
<figureCitation id="13552A50FFDCFFF1C2A3FC79FED3FC7A" box="[159,280,968,995]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="4.[151,250,1905,1929]" captionStartId-1="6.[151,250,1949,1973]" captionStartId-10="16.[151,250,1899,1923]" captionStartId-11="17.[151,250,1988,2012]" captionStartId-2="7.[151,250,1427,1451]" captionStartId-3="9.[151,250,1895,1919]" captionStartId-4="10.[151,250,890,914]" captionStartId-5="11.[151,250,1824,1848]" captionStartId-6="12.[151,250,1114,1138]" captionStartId-7="13.[151,250,1859,1883]" captionStartId-8="14.[151,250,1838,1862]" captionStartId-9="15.[151,250,1953,1977]" captionTargetBox-0="[182,1399,216,1846]" captionTargetBox-1="[171,1432,202,1893]" captionTargetBox-10="[321,1260,659,1832]" captionTargetBox-11="[162,1430,319,1961]" captionTargetBox-2="[164,1409,212,1354]" captionTargetBox-3="[201,1400,209,1833]" captionTargetBox-4="[166,1415,197,851]" captionTargetBox-5="[229,1359,182,1800]" captionTargetBox-6="[179,1370,196,1079]" captionTargetBox-7="[250,1339,204,1800]" captionTargetBox-8="[247,1354,212,1787]" captionTargetBox-9="[365,1220,179,1908]" captionTargetId-0="figure-25@4.[151,1435,181,1881]" captionTargetId-1="figure-25@6.[151,1435,181,1926]" captionTargetId-10="figure-155@16.[151,1435,622,1874]" captionTargetId-11="figure-81@17.[151,1435,307,1971]" captionTargetId-2="figure-17@7.[151,1435,181,1403]" captionTargetId-3="figure-17@9.[151,1435,181,1871]" captionTargetId-4="figure-25@10.[151,1435,181,866]" captionTargetId-5="figure-17@11.[228,1359,181,1800]" captionTargetId-6="figure-25@12.[160,1427,181,1090]" captionTargetId-7="figure-17@13.[199,1389,181,1835]" captionTargetId-8="figure-25@14.[199,1389,181,1814]" captionTargetId-9="figure-44@15.[179,1412,151,1930]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-10="16" captionTargetPageId-11="17" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." captionText-1="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." captionText-10="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." captionText-11="FIGURE 12. Distributional map of the species H. triangularis group." captionText-2="FIGURE 3. Hypoxys saltensis sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Scale bars = 1 mm." captionText-3="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." captionText-4="FIGURE 5. Hypoxys ocanensis sp. n. Female genital plates: A—ventral view, B—oblique view. Scale bars = 1 mm." captionText-5="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." captionText-6="FIGURE 7. Phallus A—H. triangularis; B—H. subrastratus; C—Hypoxys myrtacivorus sp. n. Acronyms: ph—phallotheca; vs—vesica. Scale bars = 1mm." captionText-7="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." captionText-8="FIGURE 9. Female genital plates in oblique view, mediolateral crest swollen on valvifers VIII indicated by the arrow: A— Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." captionText-9="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi-0="http://doi.org/10.5281/zenodo.6911505" figureDoi-1="http://doi.org/10.5281/zenodo.6911507" figureDoi-10="http://doi.org/10.5281/zenodo.6911536" figureDoi-11="http://doi.org/10.5281/zenodo.6911540" figureDoi-2="http://doi.org/10.5281/zenodo.6911509" figureDoi-3="http://doi.org/10.5281/zenodo.6911514" figureDoi-4="http://doi.org/10.5281/zenodo.6911516" figureDoi-5="http://doi.org/10.5281/zenodo.6911520" figureDoi-6="http://doi.org/10.5281/zenodo.6911522" figureDoi-7="http://doi.org/10.5281/zenodo.6911526" figureDoi-8="http://doi.org/10.5281/zenodo.6911528" figureDoi-9="http://doi.org/10.5281/zenodo.6911532" httpUri-0="https://zenodo.org/record/6911505/files/figure.png" httpUri-1="https://zenodo.org/record/6911507/files/figure.png" httpUri-10="https://zenodo.org/record/6911536/files/figure.png" httpUri-11="https://zenodo.org/record/6911540/files/figure.png" httpUri-2="https://zenodo.org/record/6911509/files/figure.png" httpUri-3="https://zenodo.org/record/6911514/files/figure.png" httpUri-4="https://zenodo.org/record/6911516/files/figure.png" httpUri-5="https://zenodo.org/record/6911520/files/figure.png" httpUri-6="https://zenodo.org/record/6911522/files/figure.png" httpUri-7="https://zenodo.org/record/6911526/files/figure.png" httpUri-8="https://zenodo.org/record/6911528/files/figure.png" httpUri-9="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Figs. 1–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374655EFFDCFFF1C2ABFBA1FD61FABF" pageId="1" pageNumber="502" type="diagnosis">
<paragraph id="8BD136D5FFDCFFF1C2ABFBA1FD61FABF" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="502">
<emphasis id="B91AEAC7FFDCFFF1C2ABFBA1FED9FBB3" bold="true" box="[151,274,1040,1066]" pageId="1" pageNumber="502">Diagnosis.</emphasis>
Body uniformly green with corium reddish-brown (
<figureCitation id="13552A50FFDCFFF1C102FBA1FC40FBB2" box="[830,907,1040,1067]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Fig. 10</figureCitation>
) (green with yellow spots in 
<emphasis id="B91AEAC7FFDCFFF1C683FBA0FAB2FBB3" box="[1215,1401,1040,1066]" italics="true" pageId="1" pageNumber="502">
<taxonomicName id="4C6E4D56FFDCFFF1C683FBA0FA97FBB3" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[1215,1372,1040,1066]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="species" species="triangularis">H. triangularis</taxonomicName>
—
</emphasis>
see 
<bibRefCitation id="EFFF4B24FFDCFFF1C2ABFB84FE95FBD6" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[151,350,1076,1103]" pageId="1" pageNumber="502" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDCFFF1C2D9FB84FEDEFBD7" box="[229,277,1076,1102]" italics="true" pageId="1" pageNumber="502">et al</emphasis>
. 2020
</bibRefCitation>
: Fig. 19E–F), and a median large yellow spot on the ventral surface. Species of the 
<taxonomicName id="4C6E4D56FFDCFFF1C6CBFB84FA57FBD7" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[1271,1436,1076,1102]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDCFFF1C6CBFB84FA57FBD7" box="[1271,1436,1076,1102]" italics="true" pageId="1" pageNumber="502">H. triangularis</emphasis>
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group also lack any transversal black stripe on pronotum and ventral surface, like in the other species groups of 
<taxonomicName id="4C6E4D56FFDCFFF1C2ABFBCCFF3EFB0F" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[151,245,1149,1174]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDCFFF1C2ABFBCCFF3EFB0F" box="[151,245,1149,1174]" italics="true" pageId="1" pageNumber="502">Hypoxys</emphasis>
</taxonomicName>
, except 
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<emphasis id="B91AEAC7FFDCFFF1C364FBCCFE34FB0F" box="[344,511,1148,1174]" italics="true" pageId="1" pageNumber="502">H. triangularis</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFFF4B24FFDCFFF1C07CFBCCFCDBFB0E" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[576,784,1148,1175]" pageId="1" pageNumber="502" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDCFFF1C0ADFBCCFD0FFB0F" box="[657,708,1148,1174]" italics="true" pageId="1" pageNumber="502">et al</emphasis>
. 2020
</bibRefCitation>
: Fig. 19E). Trichobothria with base yellowish (
<figureCitation id="13552A50FFDCFFF1C717FBCDFA5BFB0E" box="[1323,1424,1148,1175]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Fig. 10F</figureCitation>
). Male genitalia. Paramere (Pa) usually ax-shaped (
<figureCitation id="13552A50FFDCFFF1C0FAFB11FCD7FB22" box="[710,796,1184,1211]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="1" pageNumber="502">Fig. 1B</figureCitation>
). Inner wall of pygophore laterally swollen (
<figureCitation id="13552A50FFDCFFF1C729FB11FA53FB22" box="[1301,1432,1184,1211]" captionStart="FIGURE 3" captionStartId="7.[151,250,1427,1451]" captionTargetBox="[164,1409,212,1354]" captionTargetId="figure-17@7.[151,1435,181,1403]" captionTargetPageId="7" captionText="FIGURE 3. Hypoxys saltensis sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911509" httpUri="https://zenodo.org/record/6911509/files/figure.png" pageId="1" pageNumber="502">Figs. 3B–C</figureCitation>
, 
<figureCitation id="13552A50FFDCFFF1C2ABFB75FF69FB47" box="[151,162,1220,1246]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="1" pageNumber="502">8</figureCitation>
—black arrow). Ventral rim (Vr) with expansions slightly developed and rounded (
<figureCitation id="13552A50FFDCFFF1C601FB75FB5FFB46" box="[1085,1172,1220,1247]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="1" pageNumber="502">Fig. 4C</figureCitation>
). Female genitalia (
<figureCitation id="13552A50FFDCFFF1C74CFB75FF6CFA9A" captionStart="FIGURE 9" captionStartId="14.[151,250,1838,1862]" captionTargetBox="[247,1354,212,1787]" captionTargetId="figure-25@14.[199,1389,181,1814]" captionTargetPageId="14" captionText="FIGURE 9. Female genital plates in oblique view, mediolateral crest swollen on valvifers VIII indicated by the arrow: A— Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911528" httpUri="https://zenodo.org/record/6911528/files/figure.png" pageId="1" pageNumber="502">Fig. 9</figureCitation>
). Valvifers VIII (Vf8) with mediolateral crest swollen (black arrow); outer lateral angle of the posterior margin (red arrow) rounded or ending in an acute projection.
</paragraph>
</subSubSection>
<subSubSection id="C374655EFFDCFFF1C2FBFA81FAA4F86F" pageId="1" pageNumber="502" type="description">
<paragraph id="8BD136D5FFDCFFF1C2FBFA81FDBAFAF6" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="502">
<emphasis id="B91AEAC7FFDCFFF1C2FBFA81FEBEFAD3" bold="true" box="[199,373,1328,1354]" pageId="1" pageNumber="502">Redescription.</emphasis>
Head. Mandibular plates without spot (
<figureCitation id="13552A50FFDCFFF1C10CFA81FC46FAD2" box="[816,909,1328,1355]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Figs. 10</figureCitation>
, 
<figureCitation id="13552A50FFDCFFF1C1A5FA80FC7DFAD2" box="[921,950,1329,1355]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="1" pageNumber="502">11</figureCitation>
). Antennae yellow with setae more densely distributed on antennomeres III, IV and V.
</paragraph>
<paragraph id="8BD136D5FFDCFFF1C2FBFAC9FBE2F9F2" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="502">
Thorax.Anterolateral margin of pronotum usually not punctured, except 
<taxonomicName id="4C6E4D56FFDCFFF1C1D1FAC8FB51FA0B" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Bergroth" baseAuthorityYear="1891" box="[1005,1178,1400,1426]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="species" species="subrastratus">
<emphasis id="B91AEAC7FFDCFFF1C1D1FAC8FB51FA0B" box="[1005,1178,1400,1426]" italics="true" pageId="1" pageNumber="502">H. subrastratus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFFF4B24FFDCFFF1C6EFFAC8FA53FA0A" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[1235,1432,1400,1427]" pageId="1" pageNumber="502" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDCFFF1C71CFAC8FA9BFA0B" box="[1312,1360,1400,1426]" italics="true" pageId="1" pageNumber="502">et al</emphasis>
. 2020
</bibRefCitation>
: Fig. 19I) and 
<taxonomicName id="4C6E4D56FFDCFFF1C30AFA2CFE71FA2F" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[310,442,1436,1462]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="species" species="saltensis" status="sp. nov.">
<emphasis id="B91AEAC7FFDCFFF1C30AFA2CFE71FA2F" box="[310,442,1436,1462]" italics="true" pageId="1" pageNumber="502">H. saltensis</emphasis>
</taxonomicName>
<emphasis id="B91AEAC7FFDCFFF1C3FFFA2CFDCEFA2E" bold="true" box="[451,517,1437,1463]" pageId="1" pageNumber="502">
<taxonomicNameLabel id="A22957BCFFDCFFF1C3FFFA2CFDCEFA2E" box="[451,517,1437,1463]" pageId="1" pageNumber="502" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13552A50FFDCFFF1C029FA2DFDB6FA2E" box="[533,637,1436,1463]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Fig. 10E</figureCitation>
). Apex of humeral angle with margin brown (
<figureCitation id="13552A50FFDCFFF1C6B0FA2DFB27FA2E" box="[1164,1260,1436,1463]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Figs. 10</figureCitation>
, 
<figureCitation id="13552A50FFDCFFF1C6C6FA2CFADCFA2E" box="[1274,1303,1437,1463]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="1" pageNumber="502">11</figureCitation>
). Pronotum and scutellum usually with brown punctures (
<figureCitation id="13552A50FFDCFFF1C09CFA71FCC1FA42" box="[672,778,1472,1499]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Fig. 10A</figureCitation>
). Anterior margin of scutellum without yellow stripe, like the one found in some species of 
<taxonomicName id="4C6E4D56FFDCFFF1C019FA54FD05FA66" baseAuthorityName="Fabricius" baseAuthorityYear="1803" box="[549,718,1508,1535]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B91AEAC7FFDCFFF1C019FA54FD05FA66" bold="true" box="[549,718,1508,1535]" italics="true" pageId="1" pageNumber="502">H. quadridens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6E4D56FFDCFFF1C137FA54FC0FFA66" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[779,964,1508,1535]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="B91AEAC7FFDCFFF1C137FA54FC0FFA66" bold="true" box="[779,964,1508,1535]" italics="true" pageId="1" pageNumber="502">H. oxyacanthus</emphasis>
</taxonomicName>
groups. Costal margin of corium with a black spot reaching approximately 1/3 of corial extension (
<figureCitation id="13552A50FFDCFFF1C178F9B9FC6EF9BA" box="[836,933,1544,1571]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Figs. 10</figureCitation>
, 
<figureCitation id="13552A50FFDCFFF1C188F9B8FC1AF9BA" box="[948,977,1545,1571]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="1" pageNumber="502">11</figureCitation>
). Propleura with concolorous punctures concentrated on posterior half. Evaporatorium slightly whitish (
<figureCitation id="13552A50FFDCFFF1C15AF99DFC19F9DE" box="[870,978,1580,1607]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Figs. 10F</figureCitation>
, 
<figureCitation id="13552A50FFDCFFF1C1E2F99CFBDAF9DE" box="[990,1041,1581,1607]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="1" pageNumber="502">11A</figureCitation>
). Metasternal process with arms of anterior bifurcation receiving fourth and most of third rostral segment (
<figureCitation id="13552A50FFDCFFF1C188F9E1FBD7F9F2" box="[948,1052,1616,1643]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Fig. 10A</figureCitation>
).
</paragraph>
<paragraph id="8BD136D5FFDCFFF1C2FBF9C5FE5AF94E" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="502">
Abdomen. Dorsal surface with brown spots. Connexivum with two barely excavated areas (
<figureCitation id="13552A50FFDCFFF1C6C5F9C5FAAAF916" box="[1273,1377,1652,1679]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Fig. 10C</figureCitation>
) and usually not associated with dark spot (except 
<taxonomicName id="4C6E4D56FFDCFFF1C0A1F928FC8DF92B" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[669,838,1688,1714]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="502" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDCFFF1C0A1F928FC8DF92B" box="[669,838,1688,1714]" italics="true" pageId="1" pageNumber="502">H. triangularis</emphasis>
</taxonomicName>
). Pseudosutures with or without adjacent dark green spot (
<figureCitation id="13552A50FFDCFFF1C2E9F90DFE8DF94E" box="[213,326,1724,1751]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="1" pageNumber="502">Figs. 10B</figureCitation>
, 
<figureCitation id="13552A50FFDCFFF1C36DF90CFE48F94E" box="[337,387,1725,1751]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="1" pageNumber="502">11D</figureCitation>
).
</paragraph>
<paragraph id="8BD136D5FFDCFFF1C2FBF951FAE1F8FE" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="502">
Male. Pygophore dorsally with brown to reddish brown spot occupying about 1/2 of the length of the pygophore; distal part of dorsal surface coarse (
<figureCitation id="13552A50FFDCFFF1C027F8B5FDB9F886" box="[539,626,1796,1823]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="1" pageNumber="502">Fig. 1A</figureCitation>
). Superior process (Pc) of genital cup brown and positioned near to dorsal rim (
<figureCitation id="13552A50FFDCFFF1C2F0F899FEE8F8DA" box="[204,291,1832,1859]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="1" pageNumber="502">Fig. 2C</figureCitation>
). Paramere directed posterolaterally (
<figureCitation id="13552A50FFDCFFF1C081F899FCCBF8DA" box="[701,768,1832,1859]" captionStart="FIGURE 9" captionStartId="14.[151,250,1838,1862]" captionTargetBox="[247,1354,212,1787]" captionTargetId="figure-25@14.[199,1389,181,1814]" captionTargetPageId="14" captionText="FIGURE 9. Female genital plates in oblique view, mediolateral crest swollen on valvifers VIII indicated by the arrow: A— Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911528" httpUri="https://zenodo.org/record/6911528/files/figure.png" pageId="1" pageNumber="502">Fig. 9</figureCitation>
); base with a swollen and rounded protuberance (
<figureCitation id="13552A50FFDCFFF1C723F899FAA9F8DA" box="[1311,1378,1832,1859]" captionStart="FIGURE 9" captionStartId="14.[151,250,1838,1862]" captionTargetBox="[247,1354,212,1787]" captionTargetId="figure-25@14.[199,1389,181,1814]" captionTargetPageId="14" captionText="FIGURE 9. Female genital plates in oblique view, mediolateral crest swollen on valvifers VIII indicated by the arrow: A— Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911528" httpUri="https://zenodo.org/record/6911528/files/figure.png" pageId="1" pageNumber="502">Fig. 9</figureCitation>
, blue arrow). Proctiger (Proc) with posterior face triangular, elliptical or pentagonal, and bent ventrally (
<figureCitation id="13552A50FFDCFFF1C6E4F8FDFAD0F8FE" box="[1240,1307,1868,1895]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="1" pageNumber="502">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BD136D5FFDCFFF1C2FBF8C1FAA4F86F" blockId="1.[151,1437,1040,2039]" pageId="1" pageNumber="502">
Female. Valvifers VIII and Valvifers IX (Vf9) convex (
<figureCitation id="13552A50FFDCFFF1C121F8C1FCB9F812" box="[797,882,1904,1931]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="1" pageNumber="502">Fig. 2E</figureCitation>
). Laterotergites VIII (La8) with margin black (
<figureCitation id="13552A50FFDCFFF1C753F8C1FF6CF836" captionStart="FIGURE 9" captionStartId="14.[151,250,1838,1862]" captionTargetBox="[247,1354,212,1787]" captionTargetId="figure-25@14.[199,1389,181,1814]" captionTargetPageId="14" captionText="FIGURE 9. Female genital plates in oblique view, mediolateral crest swollen on valvifers VIII indicated by the arrow: A— Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911528" httpUri="https://zenodo.org/record/6911528/files/figure.png" pageId="1" pageNumber="502">Fig. 9</figureCitation>
). Laterotergites IX (La9) with outer lateral margin sinuous, well projected beyond sclerite uniting laterotergites VIII; almost reaching level of laterotergites VIII (
<figureCitation id="13552A50FFDCFFF1C0EEF809FCE0F84A" box="[722,811,1976,2003]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="1" pageNumber="502">Fig. 4E</figureCitation>
). Capsula seminalis (Cs) with two thick projections of equal lengths (
<figureCitation id="13552A50FFDCFFF1C307F86DFE5EF86F" box="[315,405,2012,2038]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="1" pageNumber="502">Fig. 6D</figureCitation>
); projections of capsula seminalis of the same width as pars intermedialis (Pi) (
<figureCitation id="13552A50FFDCFFF1C731F86DFAA9F86F" box="[1293,1378,2012,2038]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="1" pageNumber="502">Fig. 6F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374655EFFDFFFF2C2FBFF27FC1FFB70" pageId="2" pageNumber="503" type="discussion">
<paragraph id="8BD136D5FFDFFFF2C2FBFF27FC1FFB70" blockId="2.[151,1437,150,1329]" pageId="2" pageNumber="503">
<emphasis id="B91AEAC7FFDFFFF2C2FBFF27FE84FF29" bold="true" box="[199,335,150,176]" pageId="2" pageNumber="503">Comments.</emphasis>
Five new species are described into 
<taxonomicName id="4C6E4D56FFDFFFF2C0D7FF26FC53FF28" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[747,920,150,177]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C0D7FF26FC53FF28" bold="true" box="[747,920,150,177]" italics="true" pageId="2" pageNumber="503">H. triangularis</emphasis>
</taxonomicName>
group. Therefore, the new composition of the group is: 
<taxonomicName id="4C6E4D56FFDFFFF2C33FFF0AFE39FF4C" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[259,498,187,213]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C33FFF0AFE39FF4C" box="[259,498,187,213]" italics="true" pageId="2" pageNumber="503">Hypoxys triangularis</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C03DFF0AFDA0FF4D" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[513,619,187,212]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="capito">
<emphasis id="B91AEAC7FFDFFFF2C03DFF0AFDA0FF4D" box="[513,619,187,212]" italics="true" pageId="2" pageNumber="503">H. capito</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C047FF0AFCE1FF4C" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Bergroth" baseAuthorityYear="1891" box="[635,810,187,213]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="subrastratus">
<emphasis id="B91AEAC7FFDFFFF2C047FF0AFCE1FF4C" box="[635,810,187,213]" italics="true" pageId="2" pageNumber="503">H. subrastratus</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C106FF0AFC3AFF4D" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[826,1009,187,212]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="myrtacivorus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C106FF0AFC3AFF4D" box="[826,1009,187,212]" italics="true" pageId="2" pageNumber="503">H. myrtacivorus</emphasis>
</taxonomicName>
<emphasis id="B91AEAC7FFDFFFF2C1C5FF0AFBF1FF4C" bold="true" box="[1017,1082,187,213]" pageId="2" pageNumber="503">
<taxonomicNameLabel id="A22957BCFFDFFFF2C1C5FF0AFBF1FF4C" box="[1017,1082,187,213]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C675FF0AFB07FF4C" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1097,1228,187,213]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="saltensis" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C675FF0AFB07FF4C" box="[1097,1228,187,213]" italics="true" pageId="2" pageNumber="503">H. saltensis</emphasis>
</taxonomicName>
<emphasis id="B91AEAC7FFDFFFF2C6E9FF0AFADEFF4C" bold="true" box="[1237,1301,187,213]" pageId="2" pageNumber="503">
<taxonomicNameLabel id="A22957BCFFDFFFF2C6E9FF0AFADEFF4C" box="[1237,1301,187,213]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C719FF0AFA57FF4D" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1317,1436,187,212]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C719FF0AFA57FF4D" box="[1317,1436,187,212]" italics="true" pageId="2" pageNumber="503">H. striatus</emphasis>
</taxonomicName>
<emphasis id="B91AEAC7FFDFFFF2C2ABFF6EFF1CFF60" bold="true" box="[151,215,223,249]" pageId="2" pageNumber="503">
<taxonomicNameLabel id="A22957BCFFDFFFF2C2ABFF6EFF1CFF60" box="[151,215,223,249]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C2DAFF6EFEA3FF61" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[230,360,223,248]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="curvatus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C2DAFF6EFEA3FF61" box="[230,360,223,248]" italics="true" pageId="2" pageNumber="503">H. curvatus</emphasis>
</taxonomicName>
<emphasis id="B91AEAC7FFDFFFF2C34CFF6EFE7BFF60" bold="true" box="[368,432,223,249]" pageId="2" pageNumber="503">
<taxonomicNameLabel id="A22957BCFFDFFFF2C34CFF6EFE7BFF60" box="[368,432,223,249]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C6E4D56FFDFFFF2C3DBFF6EFDBCFF61" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[487,631,223,248]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="ocanensis" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C3DBFF6EFDBCFF61" box="[487,631,223,248]" italics="true" pageId="2" pageNumber="503">H. ocanensis</emphasis>
</taxonomicName>
<emphasis id="B91AEAC7FFDFFFF2C043FF6EFD74FF60" bold="true" box="[639,703,223,249]" pageId="2" pageNumber="503">
<taxonomicNameLabel id="A22957BCFFDFFFF2C043FF6EFD74FF60" box="[639,703,223,249]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
The 
<taxonomicName id="4C6E4D56FFDFFFF2C0C4FF6EFC6EFF60" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[760,933,222,249]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C0C4FF6EFC6EFF60" bold="true" box="[760,933,222,249]" italics="true" pageId="2" pageNumber="503">H. triangularis</emphasis>
</taxonomicName>
group has species associated with cultivated plants of the family 
<taxonomicName id="4C6E4D56FFDFFFF2C3BBFEB2FDF5FE84" authority="Juss." authorityName="Juss." box="[391,574,259,285]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="503" phylum="Tracheophyta" rank="family">Myrtaceae Juss.</taxonomicName>
(Angiosperma; 
<taxonomicName id="4C6E4D56FFDFFFF2C0C6FEB2FCA8FE84" box="[762,867,259,285]" class="Magnoliopsida" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="503" phylum="Tracheophyta" rank="order">Myrtales</taxonomicName>
): 
<taxonomicName id="4C6E4D56FFDFFFF2C146FEB2FBEEFE84" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[890,1061,259,285]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C146FEB2FBEEFE84" box="[890,1061,259,285]" italics="true" pageId="2" pageNumber="503">H. triangularis</emphasis>
</taxonomicName>
associated the 
<taxonomicName id="4C6E4D56FFDFFFF2C6E6FEB2FA57FE84" box="[1242,1436,259,285]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="uniflora">
<emphasis id="B91AEAC7FFDFFFF2C6E6FEB2FA57FE84" box="[1242,1436,259,285]" italics="true" pageId="2" pageNumber="503">Eugenia uniflora</emphasis>
</taxonomicName>
L. (brazilian-cherry) and 
<taxonomicName id="4C6E4D56FFDFFFF2C382FE96FD2BFED8" authority="(guava)" baseAuthorityName="Guava" box="[446,736,295,321]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="519" phylum="Tracheophyta" rank="species" species="guajava">
<emphasis id="B91AEAC7FFDFFFF2C382FE96FDB4FED9" box="[446,639,295,321]" italics="true" pageId="2" pageNumber="503">Psidium guajava</emphasis>
(guava)
</taxonomicName>
(
<bibRefCitation id="EFFF4B24FFDFFFF2C0CFFE96FC74FED8" author="Lima, M. S. &amp; Fernandes, J. A. M. &amp; Lima, I. M. M." box="[755,959,295,321]" pageId="2" pageNumber="503" pagination="674 - 675" refId="ref10430" refString="Lima, M. S., Fernandes, J. A. M. &amp; Lima, I. M. M. (2010) Record of Edessa scabriventris Stal (Hemiptera: Pentatomidae) Associated to Eugenia uniflora (Brazilian-Cherry) and Psidium guajava (Guava) (Myrtaceae), in North-Northeastern Brazil. Neotropical Entomology 39 (4), 674 - 675. https: // doi. org / 10.1590 / S 1519 - 566 X 2010000400032" type="journal article" year="2010">
<collectingRegion id="49AAF837FFDFFFF2C0CFFE96FCFBFED8" box="[755,816,295,321]" country="Peru" name="Lima" pageId="2" pageNumber="503">Lima</collectingRegion>
<emphasis id="B91AEAC7FFDFFFF2C107FE96FCBCFED8" box="[827,887,295,321]" italics="true" pageId="2" pageNumber="503">et al.</emphasis>
2010
</bibRefCitation>
); 
<taxonomicName id="4C6E4D56FFDFFFF2C1EAFE96FB43FED8" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Bergroth" baseAuthorityYear="1891" box="[982,1160,295,321]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="subrastratus">
<emphasis id="B91AEAC7FFDFFFF2C1EAFE96FB43FED8" box="[982,1160,295,321]" italics="true" pageId="2" pageNumber="503">H. subrastratus</emphasis>
</taxonomicName>
associated the 
<taxonomicName id="4C6E4D56FFDFFFF2C703FE96FE0BFEFC" authority="(brazilian-cherry)" baseAuthorityName="Brazilian-Cherry" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="uniflora">
<emphasis id="B91AEAC7FFDFFFF2C703FE96FF3AFEFC" italics="true" pageId="2" pageNumber="503">Eugenia uniflora</emphasis>
(brazilian-cherry)
</taxonomicName>
(data base Inaturalist; Fernandes pers. Comm.); 
<taxonomicName id="4C6E4D56FFDFFFF2C1CFFEFAFB67FEFD" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1011,1196,331,356]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="myrtacivorus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C1CFFEFAFB67FEFD" box="[1011,1196,331,356]" italics="true" pageId="2" pageNumber="503">H. myrtacivorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957BCFFDFFFF2C689FEFAFB3FFEFC" box="[1205,1268,331,357]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
associated the 
<taxonomicName id="4C6E4D56FFDFFFF2C2ABFEDEFDECFE10" authority="(Kunth) McVaugh" authorityName="McVaugh" baseAuthorityName="Kunth" box="[151,551,367,393]" class="Magnoliopsida" family="Myrtaceae" genus="Myrciaria" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="503" phylum="Tracheophyta" rank="species" species="dubia">
<emphasis id="B91AEAC7FFDFFFF2C2ABFEDEFE9BFE10" box="[151,336,367,393]" italics="true" pageId="2" pageNumber="503">Myrciaria dubia</emphasis>
(Kunth) McVaugh
</taxonomicName>
(camu-camu) (
<bibRefCitation id="EFFF4B24FFDFFFF2C0E8FEDEFC71FE10" author="Panduro, M. P. &amp; Ruiz, R. R. &amp; Salgado, E. R. &amp; Vasquez, C. D. &amp; Vallejo, J. V. &amp; Coral, A. G. &amp; Sanchez, H. I. &amp; Ucariegue, A. L. &amp; Peramas, R. F. &amp; Vizcarra, R. V. &amp; Bensimon, C. L." box="[724,954,367,393]" pageId="2" pageNumber="503" refId="ref10819" refString="Panduro, M. P., Ruiz, R. R., Salgado, E. R., Vasquez, C. D., Vallejo, J. V., Coral, A. G., Sanchez, H. I., Ucariegue, A. L., Peramas, R. F., Vizcarra, R. V. &amp; Bensimon, C. L. (2001) Sistema de produccion de camu camu en restinga. Proyecto Bioexport camu camu, Peru. Proyecto Bioexport camu camu, Iquitos, 141 pp." type="book" year="2001">
Panduro 
<emphasis id="B91AEAC7FFDFFFF2C100FEDEFCBDFE10" box="[828,886,367,393]" italics="true" pageId="2" pageNumber="503">et al.</emphasis>
2001
</bibRefCitation>
), 
<taxonomicName id="4C6E4D56FFDFFFF2C1F3FEDEFB76FE10" authorityName="Reinw. ex Blume" authorityYear="1850" box="[975,1213,367,393]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="malaccensis">
<emphasis id="B91AEAC7FFDFFFF2C1F3FEDEFB76FE10" box="[975,1213,367,393]" italics="true" pageId="2" pageNumber="503">Eugenia malaccensis</emphasis>
</taxonomicName>
L. (red jambo) and 
<taxonomicName id="4C6E4D56FFDFFFF2C2ABFE22FE93FE34" box="[151,344,403,429]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="uniflora">
<emphasis id="B91AEAC7FFDFFFF2C2ABFE22FE93FE34" box="[151,344,403,429]" italics="true" pageId="2" pageNumber="503">Eugenia uniflora</emphasis>
</taxonomicName>
L. (see comments of 
<taxonomicName id="4C6E4D56FFDFFFF2C066FE22FCD8FE35" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[602,787,403,428]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="myrtacivorus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C066FE22FCD8FE35" box="[602,787,403,428]" italics="true" pageId="2" pageNumber="503">H. myrtacivorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957BCFFDFFFF2C121FE22FC96FE34" box="[797,861,403,429]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
). Parameres of this species group are also almost straight in dorsal view and never strongly curved laterally, characteristic shared with 
<taxonomicName id="4C6E4D56FFDFFFF2C672FE06FAEBFE48" authorityName="Walker" authorityYear="1868" box="[1102,1312,438,465]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="balteatus">
<emphasis id="B91AEAC7FFDFFFF2C672FE06FAEBFE48" bold="true" box="[1102,1312,438,465]" italics="true" pageId="2" pageNumber="503">Hypoxys balteatus</emphasis>
</taxonomicName>
group (see in 
<bibRefCitation id="EFFF4B24FFDFFFF2C28BFE6AFE7DFE6C" author="Mendonca, M. T. S. &amp; Nunes, B. M. &amp; Fernandes, J. A. M." box="[183,438,475,501]" pageId="2" pageNumber="503" pagination="289 - 318" refId="ref10514" refString="Mendonca, M. T. S., Nunes, B. M. &amp; Fernandes, J. A. M. (2021) Description of fifteen new species of the Hypoxys balteatus species group (Hemiptera: Heteroptera: Pentatomidae). Acta Entomologica Musei Nationalis Pragae, 61 (1), 289 - 318. https: // doi. org / 10.37520 / aemnp. 2021.017" type="journal article" year="2021">
Mendonça 
<emphasis id="B91AEAC7FFDFFFF2C30AFE6AFEA1FE6C" box="[310,362,475,501]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
. 2021
</bibRefCitation>
). The internal genitalia of some Edessinae genera were described, but never compared or discussed in a comprehensive way (
<taxonomicName id="4C6E4D56FFDFFFF2C066FE4EFC7CFD80" authority="Stal, 1870" authorityName="Stal" authorityYear="1870" box="[602,951,510,537]" class="Insecta" family="Pentatomidae" genus="Pantochlora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="vivida">
<emphasis id="B91AEAC7FFDFFFF2C066FE4EFCFEFD80" box="[602,821,511,537]" italics="true" pageId="2" pageNumber="503">Pantochlora vivida</emphasis>
Stål, 1870
</taxonomicName>
, ilustrated— 
<bibRefCitation id="EFFF4B24FFDFFFF2C675FE4EFB25FD80" author="Kumar, R." box="[1097,1262,511,537]" pageId="2" pageNumber="503" pagination="681 - 695" refId="ref10374" refString="Kumar, R. (1969) Morphology and Relationships of the Pentatomoidea (Heteroptera). III. Natalicolinae and Some Tessaratomidae of Uncertain Position, Annals of the Entomological Society of America, 62 (4), 681 - 695. https: // doi. org / 10.1093 / aesa / 62.4.681" type="journal article" year="1969">Kumar (1969)</bibRefCitation>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C6C2FE4EFEF8FDA4" authority="Laporte, 1832" authorityName="Laporte" authorityYear="1832" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C6C2FE4EFA56FD80" box="[1278,1437,511,537]" italics="true" pageId="2" pageNumber="503">Brachystethus</emphasis>
Laporte, 1832
</taxonomicName>
— 
<bibRefCitation id="EFFF4B24FFDFFFF2C36CFD92FDB4FDA4" author="Grazia, J. &amp; Barcellos, A." box="[336,639,547,573]" pageId="2" pageNumber="503" pagination="413 - 446" refId="ref10258" refString="Grazia, J. &amp; Barcellos, A. (2003) Revision of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Serie Zoologia, 93 (4), 413 - 446." type="journal article" year="2003">Grazia &amp; Barcellos (2003)</bibRefCitation>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C0B7FD92FBE6FDA4" authority="Silva &amp; Fernandes, 2013" authorityName="Silva &amp; Fernandes" authorityYear="2013" box="[651,1069,547,573]" class="Insecta" family="Pentatomidae" genus="Paraedessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C0B7FD92FCC5FDA4" box="[651,782,547,573]" italics="true" pageId="2" pageNumber="503">Paraedessa</emphasis>
<bibRefCitation id="EFFF4B24FFDFFFF2C129FD92FBE6FDA4" author="Silva, V. J. &amp; Nunes, B. M. &amp; Fernandes, J. A. M." box="[789,1069,547,573]" pageId="2" pageNumber="503" pagination="395 - 416" refId="ref11032" refString="Silva, V. J., Nunes, B. M. &amp; Fernandes, J. A. M. (2013) Paraedessa, a new genus of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 3716 (3), 395 - 416. https: // doi. org / 10.11646 / zootaxa. 3716.3.4" type="journal article" year="2013">Silva &amp; Fernandes, 2013</bibRefCitation>
</taxonomicName>
— 
<bibRefCitation id="EFFF4B24FFDFFFF2C676FD92FAD3FDA4" author="Silva, V. J. &amp; Nunes, B. M. &amp; Fernandes, J. A. M." box="[1098,1304,547,573]" pageId="2" pageNumber="503" pagination="395 - 416" refId="ref11032" refString="Silva, V. J., Nunes, B. M. &amp; Fernandes, J. A. M. (2013) Paraedessa, a new genus of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 3716 (3), 395 - 416. https: // doi. org / 10.11646 / zootaxa. 3716.3.4" type="journal article" year="2013">
Silva 
<emphasis id="B91AEAC7FFDFFFF2C6B6FD92FB77FDA4" box="[1162,1212,547,573]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
. (2013)
</bibRefCitation>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C718FD92FF05FDF8" authority="Say, 1832" authorityName="Say" authorityYear="1832" class="Magnoliopsida" family="Salicaceae" genus="Ascra" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="503" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C718FD92FAAEFDA4" box="[1316,1381,547,573]" italics="true" pageId="2" pageNumber="503">Ascra</emphasis>
Say, 1832
</taxonomicName>
— 
<bibRefCitation id="EFFF4B24FFDFFFF2C2D7FDF6FE05FDF8" author="Santos, B. T. S. &amp; Silva, V. J. &amp; Fernandes, J. A. M." box="[235,462,582,609]" pageId="2" pageNumber="503" pagination="445 - 470" refId="ref10958" refString="Santos, B. T. S., Silva, V. J. &amp; Fernandes, J. A. M. (2015) Revision of Ascra with proposition of the bifida species group and description of two new species (Hemiptera: Pentatomidae: Edessinae). Zootaxa, 4034 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4034.3.2" type="journal article" year="2015">
Santos 
<emphasis id="B91AEAC7FFDFFFF2C302FDF6FEBAFDF8" box="[318,369,583,609]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
. (2015)
</bibRefCitation>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C3E7FDF6FC7AFDF8" authority="Almeida &amp; Fernandes, 2018" authorityName="Almeida &amp; Fernandes" authorityYear="2018" box="[475,945,583,609]" class="Insecta" family="Pentatomidae" genus="Plagaedessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C3E7FDF6FDA2FDF8" box="[475,617,583,609]" italics="true" pageId="2" pageNumber="503">Plagaedessa</emphasis>
<bibRefCitation id="EFFF4B24FFDFFFF2C04CFDF6FC7AFDF8" author="Almeida, F. R. A. &amp; Nunes, B. M. &amp; Fernandes, J. A. M." box="[624,945,583,609]" pageId="2" pageNumber="503" pagination="254 - 268" refId="ref9806" refString="Almeida, F. R. A., Nunes, B. M. &amp; Fernandes, J. A. M. (2018) A new genus and new species of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 4377 (2), 254 - 268. https: // doi. org / 10.11646 / zootaxa. 4377.2.6" type="journal article" year="2018">Almeida &amp; Fernandes, 2018</bibRefCitation>
</taxonomicName>
— 
<bibRefCitation id="EFFF4B24FFDFFFF2C1F1FDF6FB0DFDF8" author="Almeida, F. R. A. &amp; Nunes, B. M. &amp; Fernandes, J. A. M." box="[973,1222,583,609]" pageId="2" pageNumber="503" pagination="254 - 268" refId="ref9806" refString="Almeida, F. R. A., Nunes, B. M. &amp; Fernandes, J. A. M. (2018) A new genus and new species of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 4377 (2), 254 - 268. https: // doi. org / 10.11646 / zootaxa. 4377.2.6" type="journal article" year="2018">
Almeida 
<emphasis id="B91AEAC7FFDFFFF2C60BFDF6FBA2FDF8" box="[1079,1129,583,609]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
. (2018)
</bibRefCitation>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C6E8FDF6FEF9FD1C" authority="Amyot &amp; Serville, 1843" authorityName="Amyot &amp; Serville" authorityYear="1843" class="Insecta" genus="Pygoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C6E8FDF6FAE1FDF8" box="[1236,1322,583,609]" italics="true" pageId="2" pageNumber="503">Pygoda</emphasis>
Amyot &amp; Serville, 1843
</taxonomicName>
— 
<bibRefCitation id="EFFF4B24FFDFFFF2C373FDDAFD90FD1C" author="Fernandes, J. A. M. &amp; Nascimento, A. T. S. &amp; Nunes, B. M." box="[335,603,619,645]" pageId="2" pageNumber="503" pagination="205 - 232" refId="ref10109" refString="Fernandes, J. A. M., Nascimento, A. T. S. &amp; Nunes, B. M. (2018) Revision of Pygoda Amyot &amp; Serville, 1843 stat. rest. (Heteroptera: Pentatomidae: Edessinae) with description of four new species. Zootaxa, 4461 (2), 205 - 232. https: // doi. org / 10.11646 / zootaxa. 4461.2.3" type="journal article" year="2018">
Fernandes 
<emphasis id="B91AEAC7FFDFFFF2C3F0FDDAFE35FD1C" box="[460,510,619,645]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
. (2018)
</bibRefCitation>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C05BFDDAFBDDFD1C" authority="Eger, 2021" authorityName="Eger" authorityYear="2021" box="[615,1046,619,645]" class="Insecta" family="Pentatomidae" genus="Graziaedessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="anastrephae">
<emphasis id="B91AEAC7FFDFFFF2C05BFDDAFC5BFD1C" box="[615,912,619,645]" italics="true" pageId="2" pageNumber="503">Graziaedessa anastrephae</emphasis>
<bibRefCitation id="EFFF4B24FFDFFFF2C1A5FDDAFBDDFD1C" author="Eger, J. E." box="[921,1046,619,645]" pageId="2" pageNumber="503" pagination="643 - 648" refId="ref9967" refString="Eger, J. E. (2021) Graziaedessa anastrephae (Heteroptera: Pentatomidae: Edessinae) a new genus and species collected in Multilure ® fruit fly traps baited with ammonium acetate and putrescine. Zootaxa, 4958 (1), 643 - 648. https: // doi. org / 10.11646 / zootaxa. 4958.1.38" type="journal article" year="2021">Eger, 2021</bibRefCitation>
</taxonomicName>
, only the 
<emphasis id="B91AEAC7FFDFFFF2C6BAFDDAFB13FD1C" box="[1158,1240,619,645]" italics="true" pageId="2" pageNumber="503">phallus</emphasis>
described— 
<bibRefCitation id="EFFF4B24FFDFFFF2C754FDDAFF2EFD30" author="Eger, J. E." pageId="2" pageNumber="503" pagination="643 - 648" refId="ref9967" refString="Eger, J. E. (2021) Graziaedessa anastrephae (Heteroptera: Pentatomidae: Edessinae) a new genus and species collected in Multilure ® fruit fly traps baited with ammonium acetate and putrescine. Zootaxa, 4958 (1), 643 - 648. https: // doi. org / 10.11646 / zootaxa. 4958.1.38" type="journal article" year="2021">Eger (2021)</bibRefCitation>
; 
<taxonomicName id="4C6E4D56FFDFFFF2C2CFFD3FFE46FD30" authorityName="Dallas" authorityYear="1851" box="[243,397,654,681]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="ovina">
<emphasis id="B91AEAC7FFDFFFF2C2CFFD3FFE46FD30" bold="true" box="[243,397,654,681]" italics="true" pageId="2" pageNumber="503">Edessa ovina</emphasis>
</taxonomicName>
group— 
<bibRefCitation id="EFFF4B24FFDFFFF2C3C8FD3EFCE1FD30" author="Fernandes, J. A. M. &amp; Silva, V. J." box="[500,810,655,681]" pageId="2" pageNumber="503" pagination="628 - 642" refId="ref10185" refString="Fernandes, J. A. M. &amp; Silva, V. J. (2021) A new species group to Edessa, the E. ovina group, with description of a new species (Heteroptera: Pentatomidae: Edessinae) from Brazil. Iheringia, Serie Zoologia, 4958 (1), 628 - 642. https: // doi. org / 10.11646 / zootaxa. 4958.1.37" type="journal article" year="2021">Fernandes &amp; Silva (2021))</bibRefCitation>
. The internal genitalia of species of 
<taxonomicName id="4C6E4D56FFDFFFF2C6ECFD3EFAFBFD31" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1232,1328,655,680]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C6ECFD3EFAFBFD31" box="[1232,1328,655,680]" italics="true" pageId="2" pageNumber="503">Hypoxys</emphasis>
</taxonomicName>
have not been describe yet. This is the first attempt to compare and use internal genitalia to characterize the groups of species. Studying the internal genitalia of species belonging to the 
<taxonomicName id="4C6E4D56FFDFFFF2C195FD66FBC2FD69" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[937,1033,727,752]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C195FD66FBC2FD69" box="[937,1033,727,752]" italics="true" pageId="2" pageNumber="503">Hypoxys</emphasis>
</taxonomicName>
, we observed some characteristics that support 
<taxonomicName id="4C6E4D56FFDFFFF2C31AFD4AFE4DFC8D" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[294,390,763,788]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C31AFD4AFE4DFC8D" box="[294,390,763,788]" italics="true" pageId="2" pageNumber="503">Hypoxys</emphasis>
</taxonomicName>
and its groups of species. The female internal genitalia present the following characteristics for 
<taxonomicName id="4C6E4D56FFDFFFF2C2FCFCAEFEEBFCA1" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[192,288,799,824]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C2FCFCAEFEEBFCA1" box="[192,288,799,824]" italics="true" pageId="2" pageNumber="503">Hypoxys</emphasis>
</taxonomicName>
: distal part of vesicular area of the ductus receptaculi sclerotized (
<figureCitation id="13552A50FFDFFFF2C62CFCAEFBADFCA0" box="[1040,1126,799,825]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6E</figureCitation>
, yellow arrow); base of the capsula seminalis spherical and sclerotized (
<figureCitation id="13552A50FFDFFFF2C0BBFCF2FD14FCC4" box="[647,735,835,861]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6D</figureCitation>
, green arrow); capsula seminalis dilated with thick projections well-developed (
<figureCitation id="13552A50FFDFFFF2C36FFCD6FE61FC18" box="[339,426,871,897]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6B</figureCitation>
); posterior annular crest more developed than anterior annular crest (
<figureCitation id="13552A50FFDFFFF2C697FCD6FACEFC18" box="[1195,1285,871,897]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6A</figureCitation>
). The 
<emphasis id="B91AEAC7FFDFFFF2C776FCD6FA57FC18" box="[1354,1436,871,897]" italics="true" pageId="2" pageNumber="503">phallus</emphasis>
of 
<taxonomicName id="4C6E4D56FFDFFFF2C28BFC3AFEBAFC3C" box="[183,369,907,933]" pageId="2" pageNumber="503">
<emphasis id="B91AEAC7FFDFFFF2C28BFC3AFEDCFC3D" box="[183,279,907,932]" italics="true" pageId="2" pageNumber="503">Hypoxys</emphasis>
species
</taxonomicName>
have the phallotheca cylindrical and curved dorsoventrally (
<figureCitation id="13552A50FFDFFFF2C61FFC3AFBA2FC3C" box="[1059,1129,907,933]" captionStart="FIGURE 7" captionStartId="12.[151,250,1114,1138]" captionTargetBox="[179,1370,196,1079]" captionTargetId="figure-25@12.[160,1427,181,1090]" captionTargetPageId="12" captionText="FIGURE 7. Phallus A—H. triangularis; B—H. subrastratus; C—Hypoxys myrtacivorus sp. n. Acronyms: ph—phallotheca; vs—vesica. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911522" httpUri="https://zenodo.org/record/6911522/files/figure.png" pageId="2" pageNumber="503">Fig. 7</figureCitation>
). The differences observed between the species belonging to different groups of 
<taxonomicName id="4C6E4D56FFDFFFF2C0E4FC1EFCF3FC51" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[728,824,943,968]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAC7FFDFFFF2C0E4FC1EFCF3FC51" box="[728,824,943,968]" italics="true" pageId="2" pageNumber="503">Hypoxys</emphasis>
</taxonomicName>
were: the 
<taxonomicName id="4C6E4D56FFDFFFF2C194FC1EFB80FC50" baseAuthorityName="Fabricius" baseAuthorityYear="1803" box="[936,1099,942,969]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B91AEAC7FFDFFFF2C194FC1EFB80FC50" bold="true" box="[936,1099,942,969]" italics="true" pageId="2" pageNumber="503">H. quadridens</emphasis>
</taxonomicName>
(
<figureCitation id="13552A50FFDFFFF2C664FC1EFB65FC50" box="[1112,1198,943,969]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6A</figureCitation>
) and 
<taxonomicName id="4C6E4D56FFDFFFF2C6D4FC1EFA57FC50" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[1256,1436,942,969]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="B91AEAC7FFDFFFF2C6D4FC1EFA57FC50" bold="true" box="[1256,1436,942,969]" italics="true" pageId="2" pageNumber="503">H. oxyacanthus</emphasis>
</taxonomicName>
(
<figureCitation id="13552A50FFDFFFF2C2A3FC62FF3DFC74" box="[159,246,979,1005]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6B</figureCitation>
) groups share the projections of capsula seminalis wider and more dilated than the pars intermedialis. In 
<taxonomicName id="4C6E4D56FFDFFFF2C2ABFC46FE8FFB88" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[151,324,1014,1041]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C2ABFC46FE8FFB88" bold="true" box="[151,324,1014,1041]" italics="true" pageId="2" pageNumber="503">H. triangularis</emphasis>
</taxonomicName>
(
<figureCitation id="13552A50FFDFFFF2C36FFC46FE2DFB88" box="[339,486,1015,1041]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6D, E, F</figureCitation>
) and 
<taxonomicName id="4C6E4D56FFDFFFF2C018FC46FD64FB88" authorityName="Walker" authorityYear="1868" box="[548,687,1014,1041]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="balteatus">
<emphasis id="B91AEAC7FFDFFFF2C018FC46FD64FB88" bold="true" box="[548,687,1014,1041]" italics="true" pageId="2" pageNumber="503">H. balteatus</emphasis>
</taxonomicName>
(
<figureCitation id="13552A50FFDFFFF2C082FC46FCDFFB88" box="[702,788,1015,1041]" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6C</figureCitation>
) groups the projections of capsula seminalis are almost as wide as pars intermedialis. Groups 
<taxonomicName id="4C6E4D56FFDFFFF2C019FBAAFD01FBAC" baseAuthorityName="Fabricius" baseAuthorityYear="1803" box="[549,714,1050,1077]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B91AEAC7FFDFFFF2C019FBAAFD01FBAC" bold="true" box="[549,714,1050,1077]" italics="true" pageId="2" pageNumber="503">H. quadridens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6E4D56FFDFFFF2C13EFBAAFC73FBAC" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[770,952,1050,1077]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="B91AEAC7FFDFFFF2C13EFBAAFC73FBAC" bold="true" box="[770,952,1050,1077]" italics="true" pageId="2" pageNumber="503">H. oxyacanthus</emphasis>
</taxonomicName>
share median portion of pars intermedialis thickened and spherical, while in 
<taxonomicName id="4C6E4D56FFDFFFF2C031FB8EFD72FBC0" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[525,697,1086,1113]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C031FB8EFD72FBC0" bold="true" box="[525,697,1086,1113]" italics="true" pageId="2" pageNumber="503">H. triangularis</emphasis>
</taxonomicName>
group this structure is spherical or cylindrical, and in 
<taxonomicName id="4C6E4D56FFDFFFF2C72DFB8EFA57FBC0" authorityName="Walker" authorityYear="1868" box="[1297,1436,1086,1113]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="balteatus">
<emphasis id="B91AEAC7FFDFFFF2C72DFB8EFA57FBC0" bold="true" box="[1297,1436,1086,1113]" italics="true" pageId="2" pageNumber="503">H. balteatus</emphasis>
</taxonomicName>
group the same structure is cylindrical. The 
<taxonomicName id="4C6E4D56FFDFFFF2C0BFFBD2FCECFBE4" baseAuthorityName="Fabricius" baseAuthorityYear="1803" box="[643,807,1122,1149]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B91AEAC7FFDFFFF2C0BFFBD2FCECFBE4" bold="true" box="[643,807,1122,1149]" italics="true" pageId="2" pageNumber="503">H. quadridens</emphasis>
</taxonomicName>
group shows the base of capsula seminalis conical (
<figureCitation id="13552A50FFDFFFF2C753FBD2FF77FB38" captionStart="FIGURE 6" captionStartId="11.[151,250,1824,1848]" captionTargetBox="[229,1359,182,1800]" captionTargetId="figure-17@11.[228,1359,181,1800]" captionTargetPageId="11" captionText="FIGURE 6. Internal female genitalia: A—Hypoxys trabeculus (Breddin, 1907) (H. quadridens group); B—H. oxyacanthus (Breddin, 1904) (H. oxyacanthus group); C—H. aspilogaster Mendonça et al., 2021 (H. balteatus group); D—H. triangularis (H. triangularis group); E—H. subrastratus (H. triangularis group); F—H. myrtacivorus sp. n. (H. triangularis group). Acronyms: aac—anterior annular crest; ch—chitinellipsen; cs—capsula seminalis; de—ductus receptaculi; vf9—valvifers 9; la9—laterotergite 9; pac—posterior annular crest; pi—pars intermedialis; tvi—thickening of vaginal intima; x—abdominal segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911520" httpUri="https://zenodo.org/record/6911520/files/figure.png" pageId="2" pageNumber="503">Fig. 6A</figureCitation>
, red arrow). 
<taxonomicName id="4C6E4D56FFDFFFF2C371FB36FD81FB38" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[333,586,1158,1185]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="B91AEAC7FFDFFFF2C371FB36FD81FB38" bold="true" box="[333,586,1158,1185]" italics="true" pageId="2" pageNumber="503">Hypoxys oxyacanthus</emphasis>
</taxonomicName>
group has the globose base of capsula seminalis as wide as the segment X. A comparative study of the internal genitalia in all taxa of Edessinae can help to understand the evolution of the genitalia in the subfamily and add diagnostic characteristics to the genera.
</paragraph>
</subSubSection>
<subSubSection id="C374655EFFDFFFF2C2FBFB43FD27FAA8" pageId="2" pageNumber="503" type="distribution">
<paragraph id="8BD136D5FFDFFFF2C2FBFB43FD27FAA8" blockId="2.[151,1437,150,1329]" pageId="2" pageNumber="503">
<emphasis id="B91AEAC7FFDFFFF2C2FBFB43FE92FA95" bold="true" box="[199,345,1266,1292]" pageId="2" pageNumber="503">Distribution</emphasis>
(
<figureCitation id="13552A50FFDFFFF2C354FB42FE71FA94" box="[360,442,1267,1293]" captionStart="FIGURE 12" captionStartId="17.[151,250,1988,2012]" captionTargetBox="[162,1430,319,1961]" captionTargetId="figure-81@17.[151,1435,307,1971]" captionTargetPageId="17" captionText="FIGURE 12. Distributional map of the species H. triangularis group." figureDoi="http://doi.org/10.5281/zenodo.6911540" httpUri="https://zenodo.org/record/6911540/files/figure.png" pageId="2" pageNumber="503">Fig. 12</figureCitation>
). 
<collectingCountry id="F3797645FFDFFFF2C3F1FB42FDFDFA94" box="[461,566,1267,1293]" name="Brazil" pageId="2" pageNumber="503">BRAZIL</collectingCountry>
; FRENCH GUIANA; 
<collectingCountry id="F3797645FFDFFFF2C10CFB42FC11FA94" box="[816,986,1267,1293]" name="Venezuela" pageId="2" pageNumber="503">VENEZUELA</collectingCountry>
; 
<collectingCountry id="F3797645FFDFFFF2C1D8FB42FBB7FA94" box="[996,1148,1267,1293]" name="Paraguay" pageId="2" pageNumber="503">PARAGUAY</collectingCountry>
; 
<collectingCountry id="F3797645FFDFFFF2C6BAFB42FAD7FA94" box="[1158,1308,1267,1293]" name="Suriname" pageId="2" pageNumber="503">SURINAME</collectingCountry>
; 
<collectingCountry id="F3797645FFDFFFF2C71AFB42FA53FA94" box="[1318,1432,1267,1293]" name="Guyana" pageId="2" pageNumber="503">GUIANA</collectingCountry>
; 
<collectingCountry id="F3797645FFDFFFF2C2ABFAA6FEC5FAA8" box="[151,270,1303,1329]" name="Bolivia" pageId="2" pageNumber="503">BOLIVIA</collectingCountry>
; 
<collectingCountry id="F3797645FFDFFFF2C325FAA6FEAFFAA8" box="[281,356,1303,1329]" name="Peru" pageId="2" pageNumber="503">PERU</collectingCountry>
; 
<collectingCountry id="F3797645FFDFFFF2C353FAA6FDCCFAA8" box="[367,519,1303,1329]" name="Colombia" pageId="2" pageNumber="503">COLOMBIA</collectingCountry>
, and 
<collectingCountry id="F3797645FFDFFFF2C07DFAA6FD23FAA8" box="[577,744,1303,1329]" name="Argentina" pageId="2" pageNumber="503">ARGENTINA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C374655EFFDFFFF2C2ABFAC9FD26FA0B" box="[151,749,1399,1426]" pageId="2" pageNumber="503" type="multiple">
<paragraph id="8BD136D5FFDFFFF2C2ABFAC9FD26FA0B" blockId="2.[151,749,1399,1426]" box="[151,749,1399,1426]" pageId="2" pageNumber="503">
<heading id="D09981B9FFDFFFF2C2ABFAC9FD26FA0B" bold="true" box="[151,749,1399,1426]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<emphasis id="B91AEAC7FFDFFFF2C2ABFAC9FD26FA0B" bold="true" box="[151,749,1399,1426]" pageId="2" pageNumber="503">
Key to species of the 
<taxonomicName id="4C6E4D56FFDFFFF2C3A1FAC9FD50FA0B" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[413,667,1399,1426]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C3A1FAC9FD50FA0B" bold="true" box="[413,667,1399,1426]" italics="true" pageId="2" pageNumber="503">Hypoxys triangularis</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374655EFFDFFFF3C2ABFA09FA57FDB3" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="key">
<keyStep id="309A2E70FFDFFFF2C2ABFA09FA57F99E" pageId="2" pageNumber="503">
<paragraph id="8BD136D5FFDFFFF2C2ABFA09FA57FA73" blockId="2.[151,1437,1464,2014]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABFA09FA57FA73" pageId="2" pageNumber="503">
1. Pronotum with posterior smoky brown stripe; ventral surface of the body with small yellow calli (See 
<bibRefCitation id="EFFF4B24FFDFFFF2C682FA09FAA6FA54" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[1214,1389,1464,1485]" pageId="2" pageNumber="503" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDFFFF2C73EFA09FAF8FA54" box="[1282,1331,1464,1485]" italics="true" pageId="2" pageNumber="503">et al.</emphasis>
2020
</bibRefCitation>
: Fig. 19E–F).................................................................................. 
<taxonomicName id="4C6E4D56FFDFFFF2C734FA64FA57FA73" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Dallas" baseAuthorityYear="1851" box="[1288,1436,1492,1514]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="B91AEAC7FFDFFFF2C734FA64FA57FA73" bold="true" box="[1288,1436,1492,1514]" italics="true" pageId="2" pageNumber="503">H. triangularis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDFFFF2C2ABFA43FA57F99E" blockId="2.[151,1437,1464,2014]" box="[151,1436,1522,1544]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABFA43FA57F99E" box="[151,1436,1522,1544]" pageId="2" pageNumber="503">
1’. Pronotum without dark stripe; ventral surface of the body without yellow calli (
<figureCitation id="13552A50FFDFFFF2C1E1FA43FBF5F99E" box="[989,1086,1522,1544]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="2" pageNumber="503">Figs. 10A</figureCitation>
, 
<figureCitation id="13552A50FFDFFFF2C674FA43FBBAF99E" box="[1096,1137,1522,1543]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="2" pageNumber="503">11A</figureCitation>
)...................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDFFFF2C2ABF9A1FA57F9DB" pageId="2" pageNumber="503">
<paragraph id="8BD136D5FFDFFFF2C2ABF9A1FA57F9BC" blockId="2.[151,1437,1464,2014]" box="[151,1436,1552,1574]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF9A1FA57F9BC" box="[151,1436,1552,1574]" pageId="2" pageNumber="503">2. Male............................................................................................... 3</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDFFFF2C2ABF99CFA57F9DB" blockId="2.[151,1437,1464,2014]" box="[151,1436,1581,1603]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF99CFA57F9DB" box="[151,1436,1581,1603]" pageId="2" pageNumber="503">2’. Female..............................................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDFFFF2C2ABF9FBFA57F9E4" pageId="2" pageNumber="503">
<paragraph id="8BD136D5FFDFFFF2C2ABF9FBFA57F9C6" blockId="2.[151,1437,1464,2014]" box="[151,1436,1610,1632]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF9FBFA57F9C6" box="[151,1436,1610,1632]" pageId="2" pageNumber="503">
3. Proctiger strongly (
<figureCitation id="13552A50FFDFFFF2C3A0F9FBFE3BF9C6" box="[412,496,1610,1632]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="2" pageNumber="503">Figs. 1B</figureCitation>
, 
<figureCitation id="13552A50FFDFFFF2C3C6F9FBFDD3F9C6" box="[506,536,1610,1631]" captionStart="FIGURE 3" captionStartId="7.[151,250,1427,1451]" captionTargetBox="[164,1409,212,1354]" captionTargetId="figure-17@7.[151,1435,181,1403]" captionTargetPageId="7" captionText="FIGURE 3. Hypoxys saltensis sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911509" httpUri="https://zenodo.org/record/6911509/files/figure.png" pageId="2" pageNumber="503">3B</figureCitation>
, See 
<bibRefCitation id="EFFF4B24FFDFFFF2C077F9FAFD33F9C6" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[587,760,1610,1632]" pageId="2" pageNumber="503" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDFFFF2C0B2F9FAFD73F9C6" box="[654,696,1610,1632]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
. 2020
</bibRefCitation>
: 
<figureCitation id="13552A50FFDFFFF2C13CF9FBFCB9F9C6" box="[768,882,1610,1632]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="2" pageNumber="503">Fig. 11A–B</figureCitation>
) or moderately excavated laterally (
<figureCitation id="13552A50FFDFFFF2C6F1F9FBFAD3F9C6" box="[1229,1304,1610,1632]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="2" pageNumber="503">Fig. 4D</figureCitation>
).........4
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDFFFF2C2ABF9D9FA57F9E4" blockId="2.[151,1437,1464,2014]" box="[151,1436,1640,1662]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF9D9FA57F9E4" box="[151,1436,1640,1662]" pageId="2" pageNumber="503">
3’. Proctiger shallowly excavated laterally (
<figureCitation id="13552A50FFDFFFF2C059F9D9FD70F9E4" box="[613,699,1640,1662]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="2" pageNumber="503">Figs. 2D</figureCitation>
, 
<figureCitation id="13552A50FFDFFFF2C0F8F9D9FD2AF9E4" box="[708,737,1640,1661]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="2" pageNumber="503">8B</figureCitation>
)........................................................ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDFFFF2C2ABF934FA57F96A" pageId="2" pageNumber="503">
<paragraph id="8BD136D5FFDFFFF2C2ABF934FA57F921" blockId="2.[151,1437,1464,2014]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF934FA57F921" pageId="2" pageNumber="503">
4. Dorsal rim of pygophore medially notched (See 
<bibRefCitation id="EFFF4B24FFDFFFF2C080F934FCA7F902" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[700,876,1669,1691]" pageId="2" pageNumber="503" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDFFFF2C13DF937FCE6F903" box="[769,813,1669,1691]" italics="true" pageId="2" pageNumber="503">et al</emphasis>
. 2020
</bibRefCitation>
– 
<figureCitation id="13552A50FFDFFFF2C144F934FC05F903" box="[888,974,1669,1691]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="2" pageNumber="503">Fig. 11A</figureCitation>
). Superior process of genital cup subtriangular, almost the same size on swollen area on inner wall of pygophore (
<figureCitation id="13552A50FFDFFFF2C161F913FC6DF92E" box="[861,934,1698,1720]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="2" pageNumber="503">Fig. 8C</figureCitation>
)............................ 
<taxonomicName id="4C6E4D56FFDFFFF2C73FF912FA57F921" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Bergroth" baseAuthorityYear="1891" box="[1283,1436,1698,1720]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="subrastratus">
<emphasis id="B91AEAC7FFDFFFF2C73FF912FA57F921" bold="true" box="[1283,1436,1698,1720]" italics="true" pageId="2" pageNumber="503">H. subrastratus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDFFFF2C2ABF971FA57F96A" blockId="2.[151,1437,1464,2014]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF971FA57F96A" pageId="2" pageNumber="503">
4’. Dorsal rim entire. Superior process of genital cup elliptic, bigger than swollen area on inner wall of pygophore (
<figureCitation id="13552A50FFDFFFF2C727F971FAA4F94C" box="[1307,1391,1728,1750]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="2" pageNumber="503">Figs. 1B</figureCitation>
, 
<figureCitation id="13552A50FFDFFFF2C746F971FA53F94C" box="[1402,1432,1728,1749]" captionStart="FIGURE 3" captionStartId="7.[151,250,1427,1451]" captionTargetBox="[164,1409,212,1354]" captionTargetId="figure-17@7.[151,1435,181,1403]" captionTargetPageId="7" captionText="FIGURE 3. Hypoxys saltensis sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911509" httpUri="https://zenodo.org/record/6911509/files/figure.png" pageId="2" pageNumber="503">3B</figureCitation>
, 
<figureCitation id="13552A50FFDFFFF2C2DEF96CFF34F96B" box="[226,255,1757,1778]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="2" pageNumber="503">4B</figureCitation>
)................................................................................................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDFFFF2C2ABF94BFA57F8B7" pageId="2" pageNumber="503">
<paragraph id="8BD136D5FFDFFFF2C2ABF94BFA57F896" blockId="2.[151,1437,1464,2014]" box="[151,1436,1786,1808]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF94BFA57F896" box="[151,1436,1786,1808]" pageId="2" pageNumber="503">
5. Superior process of genital cup coarse in posterior view (
<figureCitation id="13552A50FFDFFFF2C13AF94BFC9AF896" box="[774,849,1786,1808]" captionStart="FIGURE 3" captionStartId="7.[151,250,1427,1451]" captionTargetBox="[164,1409,212,1354]" captionTargetId="figure-17@7.[151,1435,181,1403]" captionTargetPageId="7" captionText="FIGURE 3. Hypoxys saltensis sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911509" httpUri="https://zenodo.org/record/6911509/files/figure.png" pageId="2" pageNumber="503">Fig. 3B</figureCitation>
). Paramere spearhead-shaped (
<figureCitation id="13552A50FFDFFFF2C647F94BFB0DF896" box="[1147,1222,1786,1808]" captionStart="FIGURE 3" captionStartId="7.[151,250,1427,1451]" captionTargetBox="[164,1409,212,1354]" captionTargetId="figure-17@7.[151,1435,181,1403]" captionTargetPageId="7" captionText="FIGURE 3. Hypoxys saltensis sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911509" httpUri="https://zenodo.org/record/6911509/files/figure.png" pageId="2" pageNumber="503">Fig. 3D</figureCitation>
).. 
<emphasis id="B91AEAC7FFDFFFF2C6D1F94AFA57F896" bold="true" box="[1261,1436,1786,1808]" pageId="2" pageNumber="503">
<taxonomicName id="4C6E4D56FFDFFFF2C6D1F94AFAABF889" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1261,1376,1786,1808]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="saltensis" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C6D1F94AFAABF889" bold="true" box="[1261,1376,1786,1808]" italics="true" pageId="2" pageNumber="503">H. saltensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957BCFFDFFFF2C75AF94BFA57F896" box="[1382,1436,1786,1807]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDFFFF2C2ABF8A6FA57F8B7" blockId="2.[151,1437,1464,2014]" box="[151,1436,1815,1838]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF8A6FA57F8B7" box="[151,1436,1815,1838]" pageId="2" pageNumber="503">
5’. Superior process of genital cup smooth in posterior view (
<figureCitation id="13552A50FFDFFFF2C133F8A9FCA8F8B4" box="[783,867,1816,1838]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="2" pageNumber="503">Figs. 1B</figureCitation>
, 
<figureCitation id="13552A50FFDFFFF2C151F8A9FC47F8B4" box="[877,908,1816,1837]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="2" pageNumber="503">4B</figureCitation>
). Paramere ax-shaped (
<figureCitation id="13552A50FFDFFFF2C650F8A9FB73F8B4" box="[1132,1208,1816,1838]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="2" pageNumber="503">Fig. 1D</figureCitation>
, 
<figureCitation id="13552A50FFDFFFF2C6FEF8A9FB2AF8B4" box="[1218,1249,1816,1837]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="2" pageNumber="503">4D</figureCitation>
)............. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDFFFF2C2ABF884FA57F83B" pageId="2" pageNumber="503">
<paragraph id="8BD136D5FFDFFFF2C2ABF884FA57F8FE" blockId="2.[151,1437,1464,2014]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF884FA57F8FE" pageId="2" pageNumber="503">
6. Swollen area on inner wall globose, developed, close to posterolateral angles (
<figureCitation id="13552A50FFDFFFF2C1ECF884FBD6F8D3" box="[976,1053,1845,1867]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="2" pageNumber="503">Fig. 8D</figureCitation>
). Ventral rim with bottom of excavation convex (
<figureCitation id="13552A50FFDFFFF2C308F8E3FEB6F8FE" box="[308,381,1874,1896]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="2" pageNumber="503">Fig. 1C</figureCitation>
)..................................................................... 
<emphasis id="B91AEAC7FFDFFFF2C6FCF8E2FA57F8FE" bold="true" box="[1216,1436,1874,1896]" pageId="2" pageNumber="503">
<taxonomicName id="4C6E4D56FFDFFFF2C6FCF8E2FAABF8FE" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1216,1376,1874,1896]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="myrtacivorus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C6FCF8E2FAABF8FE" bold="true" box="[1216,1376,1874,1896]" italics="true" pageId="2" pageNumber="503">H. myrtacivorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957BCFFDFFFF2C75AF8E3FA57F8FE" box="[1382,1436,1874,1895]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDFFFF2C2ABF8C1FA57F83B" blockId="2.[151,1437,1464,2014]" pageId="2" pageNumber="503">
<keyLead id="309F95E0FFDFFFF2C2ABF8C1FA57F83B" pageId="2" pageNumber="503">
6”. Swollen area on inner wall narrow, slightly developed, contiguous to posterolateral angles (
<figureCitation id="13552A50FFDFFFF2C66BF8C1FB69F81C" box="[1111,1186,1904,1926]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="2" pageNumber="503">Fig. 8E</figureCitation>
). Ventral rim with bottom of excavation flat (
<figureCitation id="13552A50FFDFFFF2C3ABF83CFE2AF83B" box="[407,481,1933,1955]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="2" pageNumber="503">Fig. 4C</figureCitation>
)............................................................... 
<emphasis id="B91AEAC7FFDFFFF2C6D0F83CFA57F83B" bold="true" box="[1260,1436,1933,1954]" pageId="2" pageNumber="503">
<taxonomicName id="4C6E4D56FFDFFFF2C6D0F83CFAABF83B" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1260,1376,1933,1954]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="curvatus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C6D0F83CFAABF83B" bold="true" box="[1260,1376,1933,1954]" italics="true" pageId="2" pageNumber="503">H. curvatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957BCFFDFFFF2C75AF83CFA57F83B" box="[1382,1436,1933,1954]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</keyStep>
<keyStep id="309A2E70FFDFFFF3C2ABF81AFA57FF53" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
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<keyLead id="309F95E0FFDFFFF2C2ABF81AFA57F844" pageId="2" pageNumber="503">
7. Superior process of genital cup subtriangular, small, swollen in posterior view (
<figureCitation id="13552A50FFDFFFF2C1CAF81BFB88F826" box="[1014,1091,1962,1984]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="2" pageNumber="503">Fig. 2B</figureCitation>
). Expansions of ventral rim hardly visible (
<figureCitation id="13552A50FFDFFFF2C313F879FEB2F844" box="[303,377,1992,2014]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="2" pageNumber="503">Fig. 2C</figureCitation>
).......................................................................... 
<emphasis id="B91AEAC7FFDFFFF2C6C4F879FA57F844" bold="true" box="[1272,1436,1991,2013]" pageId="2" pageNumber="503">
<taxonomicName id="4C6E4D56FFDFFFF2C6C4F879FAABF845" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1272,1376,1991,2013]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="B91AEAC7FFDFFFF2C6C4F879FAABF845" bold="true" box="[1272,1376,1991,2013]" italics="true" pageId="2" pageNumber="503">H. striatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957BCFFDFFFF2C75AF879FA57F844" box="[1382,1436,1992,2013]" pageId="2" pageNumber="503" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDEFFF3C2ABFF29FA57FF53" blockId="3.[151,1436,151,555]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFF29FA57FF53" pageId="3" pageNumber="504">
7’. Superior process of genital cup elliptic, narrow, smooth in posterior view (
<figureCitation id="13552A50FFDEFFF3C1FEFF26FBC4FF34" box="[962,1039,151,173]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="3" pageNumber="504">Fig. 8B</figureCitation>
). Expansions of ventral rim visible (See 
<bibRefCitation id="EFFF4B24FFDEFFF3C2DEFF04FE44FF53" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[226,399,181,202]" pageId="3" pageNumber="504" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDEFFF3C319FF04FE9BFF53" box="[293,336,181,202]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
. 2020
</bibRefCitation>
: 
<figureCitation id="13552A50FFDEFFF3C3A4FF04FE26FF53" box="[408,493,181,203]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="3" pageNumber="504">Fig. 10C</figureCitation>
)...................................................................... 
<taxonomicName id="4C6E4D56FFDEFFF3C77EFF04FA57FF53" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[1346,1436,181,202]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="capito">
<emphasis id="B91AEAC7FFDEFFF3C77EFF04FA57FF53" bold="true" box="[1346,1436,181,202]" italics="true" pageId="3" pageNumber="504">H. capito</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDEFFF3C2ABFF63FA57FE9C" pageId="3" pageNumber="504">
<paragraph id="8BD136D5FFDEFFF3C2ABFF63FA57FF7E" blockId="3.[151,1436,151,555]" box="[151,1436,210,232]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFF63FA57FF7E" box="[151,1436,210,232]" pageId="3" pageNumber="504">
8. Valvifers VIII with outer lateral angle rounded (
<figureCitation id="13552A50FFDEFFF3C091FF63FCCBFF7E" box="[685,768,210,232]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="3" pageNumber="504">Figs. 2E</figureCitation>
, See 
<bibRefCitation id="EFFF4B24FFDEFFF3C10FFF62FC2BFF7E" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[819,992,210,232]" pageId="3" pageNumber="504" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDEFFF3C14AFF62FC6BFF7E" box="[886,928,210,232]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
. 2020
</bibRefCitation>
: 
<figureCitation id="13552A50FFDEFFF3C1D5FF63FB82FF7E" box="[1001,1097,210,232]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="3" pageNumber="504">Figs. 10D</figureCitation>
, 
<figureCitation id="13552A50FFDEFFF3C66FFF63FBB7FF7E" box="[1107,1148,210,231]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="3" pageNumber="504">11D</figureCitation>
)..................... 9
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDEFFF3C2ABFF41FA57FE9C" blockId="3.[151,1436,151,555]" box="[151,1436,239,261]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFF41FA57FE9C" box="[151,1436,239,261]" pageId="3" pageNumber="504">
8’. Valvifers VIII with outer lateral angle acute (
<figureCitation id="13552A50FFDEFFF3C0ADFF5EFD28FE9C" box="[657,739,239,261]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="3" pageNumber="504">Figs. 1E</figureCitation>
, 
<figureCitation id="13552A50FFDEFFF3C0D2FF41FCC0FE9C" box="[750,779,240,261]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="3" pageNumber="504">4E</figureCitation>
, 
<figureCitation id="13552A50FFDEFFF3C129FF5EFCF8FE9C" box="[789,819,239,261]" captionStart="FIGURE 5" captionStartId="10.[151,250,890,914]" captionTargetBox="[166,1415,197,851]" captionTargetId="figure-25@10.[151,1435,181,866]" captionTargetPageId="10" captionText="FIGURE 5. Hypoxys ocanensis sp. n. Female genital plates: A—ventral view, B—oblique view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911516" httpUri="https://zenodo.org/record/6911516/files/figure.png" pageId="3" pageNumber="504">5A</figureCitation>
)................................................ 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDEFFF3C2ABFEBCFA57FEA6" pageId="3" pageNumber="504">
<paragraph id="8BD136D5FFDEFFF3C2ABFEBCFA57FEBB" blockId="3.[151,1436,151,555]" box="[151,1436,269,291]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFEBCFA57FEBB" box="[151,1436,269,291]" pageId="3" pageNumber="504">
9. Valvifers VIII with excavation in “V” (See 
<bibRefCitation id="EFFF4B24FFDEFFF3C0BEFEBCFCE4FEBB" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[642,815,269,290]" pageId="3" pageNumber="504" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDEFFF3C0F9FEBCFD3BFEBB" box="[709,752,269,290]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
. 2020
</bibRefCitation>
: 
<figureCitation id="13552A50FFDEFFF3C104FEBCFC45FEBB" box="[824,910,269,291]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1977]" captionTargetBox="[365,1220,179,1908]" captionTargetId="figure-44@15.[179,1412,151,1930]" captionTargetPageId="15" captionText="FIGURE 10. Dorsal and ventral view. A–B Hypoxys myrtacivorus sp. n.; C–D H. striatus sp. n.; E–F H. saltensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911532" httpUri="https://zenodo.org/record/6911532/files/figure.png" pageId="3" pageNumber="504">Fig. 10D</figureCitation>
).................................. ... 
<taxonomicName id="4C6E4D56FFDEFFF3C77EFEBCFA57FEBB" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[1346,1436,269,290]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="capito">
<emphasis id="B91AEAC7FFDEFFF3C77EFEBCFA57FEBB" bold="true" box="[1346,1436,269,290]" italics="true" pageId="3" pageNumber="504">H. capito</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDEFFF3C2ABFE9BFA57FEA6" blockId="3.[151,1436,151,555]" box="[151,1436,298,320]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFE9BFA57FEA6" box="[151,1436,298,320]" pageId="3" pageNumber="504">
9’. Valvifers VIII with excavation in “U” (
<figureCitation id="13552A50FFDEFFF3C064FE9BFD60FEA6" box="[600,683,298,320]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="3" pageNumber="504">Figs. 2E</figureCitation>
, See 
<bibRefCitation id="EFFF4B24FFDEFFF3C0E2FE9AFC40FEA6" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[734,907,298,320]" pageId="3" pageNumber="504" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDEFFF3C11DFE9AFC87FEA6" box="[801,844,298,320]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
. 2020
</bibRefCitation>
: 
<figureCitation id="13552A50FFDEFFF3C1A8FE9BFC22FEA6" box="[916,1001,298,320]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="3" pageNumber="504">Fig. 11D</figureCitation>
)................................. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E70FFDEFFF3C2ABFEF9FA57FE2C" pageId="3" pageNumber="504">
<paragraph id="8BD136D5FFDEFFF3C2ABFEF9FA57FEE3" blockId="3.[151,1436,151,555]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFEF9FA57FEE3" pageId="3" pageNumber="504">
10. Valvifers VIII with mediolateral crest slightly swollen (
<figureCitation id="13552A50FFDEFFF3C134FEF6FC9FFEC4" box="[776,852,327,349]" captionStart="FIGURE 9" captionStartId="14.[151,250,1838,1862]" captionTargetBox="[247,1354,212,1787]" captionTargetId="figure-25@14.[199,1389,181,1814]" captionTargetPageId="14" captionText="FIGURE 9. Female genital plates in oblique view, mediolateral crest swollen on valvifers VIII indicated by the arrow: A— Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911528" httpUri="https://zenodo.org/record/6911528/files/figure.png" pageId="3" pageNumber="504">Fig. 9F</figureCitation>
); median excavation narrow, clearly narrower than base of valvifers IX (
<figureCitation id="13552A50FFDEFFF3C35EFED4FE60FEE3" box="[354,427,357,379]" captionStart="FIGURE 2" captionStartId="6.[151,250,1949,1973]" captionTargetBox="[171,1432,202,1893]" captionTargetId="figure-25@6.[151,1435,181,1926]" captionTargetPageId="6" captionText="FIGURE 2. Hypoxys striatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911507" httpUri="https://zenodo.org/record/6911507/files/figure.png" pageId="3" pageNumber="504">Fig. 2E</figureCitation>
).................................................................... 
<emphasis id="B91AEAC7FFDEFFF3C6C4FED4FA57FEE3" bold="true" box="[1272,1436,357,378]" pageId="3" pageNumber="504">
<taxonomicName id="4C6E4D56FFDEFFF3C6C4FED4FAABFEE3" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1272,1376,357,378]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="B91AEAC7FFDEFFF3C6C4FED4FAABFEE3" bold="true" box="[1272,1376,357,378]" italics="true" pageId="3" pageNumber="504">H. striatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957BCFFDEFFF3C75AFED4FA57FEE3" box="[1382,1436,357,378]" pageId="3" pageNumber="504" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDEFFF3C2ABFE33FA57FE2C" blockId="3.[151,1436,151,555]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFE33FA57FE2C" pageId="3" pageNumber="504">
10’. Valvifers VIII with mediolateral crest swollen (
<figureCitation id="13552A50FFDEFFF3C095FE33FD3EFE0E" box="[681,757,386,408]" captionStart="FIGURE 9" captionStartId="14.[151,250,1838,1862]" captionTargetBox="[247,1354,212,1787]" captionTargetId="figure-25@14.[199,1389,181,1814]" captionTargetPageId="14" captionText="FIGURE 9. Female genital plates in oblique view, mediolateral crest swollen on valvifers VIII indicated by the arrow: A— Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911528" httpUri="https://zenodo.org/record/6911528/files/figure.png" pageId="3" pageNumber="504">Fig. 9C</figureCitation>
); median excavation wide, almost as wide as base of valvifers IX (See 
<bibRefCitation id="EFFF4B24FFDEFFF3C2DEFE11FE44FE2C" author="Nunes, B. M. &amp; Campos, L. D. &amp; Mendonca, M. T. S. &amp; Cunha, E. V. P. &amp; Fernandes, J. A. M." box="[226,399,415,437]" pageId="3" pageNumber="504" pagination="445 - 470" refId="ref10667" refString="Nunes, B. M., Campos, L. D., Mendonca, M. T. S., Cunha, E. V. P. &amp; Fernandes, J. A. M. (2020) Revision of Hypoxys Amyot &amp; Serville, 1843 stat. res. (Heteroptera: Pentatomidae). Zootaxa, 4742 (3), 445 - 470. https: // doi. org / 10.11646 / zootaxa. 4742.3.1" type="journal article" year="2020">
Nunes 
<emphasis id="B91AEAC7FFDEFFF3C319FE11FE9BFE2D" box="[293,336,415,437]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
. 2020
</bibRefCitation>
: 
<figureCitation id="13552A50FFDEFFF3C3A4FE2EFE26FE2C" box="[408,493,415,437]" captionStart="FIGURE 11" captionStartId="16.[151,250,1899,1923]" captionTargetBox="[321,1260,659,1832]" captionTargetId="figure-155@16.[151,1435,622,1874]" captionTargetPageId="16" captionText="FIGURE 11. Dorsal and ventral view. A–B H. curvatus sp. n.; C–D Hypoxys ocanensis sp. n. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.6911536" httpUri="https://zenodo.org/record/6911536/files/figure.png" pageId="3" pageNumber="504">Fig. 11D</figureCitation>
)................................................................ 
<taxonomicName id="4C6E4D56FFDEFFF3C73FFE11FA57FE2C" authorityName="Nunes et al." authorityYear="2020" baseAuthorityName="Bergroth" baseAuthorityYear="1891" box="[1283,1436,415,437]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="subrastratus">
<emphasis id="B91AEAC7FFDEFFF3C73FFE11FA57FE2C" bold="true" box="[1283,1436,415,437]" italics="true" pageId="3" pageNumber="504">H. subrastratus</emphasis>
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<keyStep id="309A2E70FFDEFFF3C2ABFE0CFA57FE76" pageId="3" pageNumber="504">
<paragraph id="8BD136D5FFDEFFF3C2ABFE0CFA57FE4B" blockId="3.[151,1436,151,555]" box="[151,1436,445,467]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFE0CFA57FE4B" box="[151,1436,445,467]" pageId="3" pageNumber="504">
11. Mediolateral crest of valvifers VIII carinated (
<figureCitation id="13552A50FFDEFFF3C0A2FE0CFD2CFE4A" box="[670,743,445,467]" captionStart="FIGURE 5" captionStartId="10.[151,250,890,914]" captionTargetBox="[166,1415,197,851]" captionTargetId="figure-25@10.[151,1435,181,866]" captionTargetPageId="10" captionText="FIGURE 5. Hypoxys ocanensis sp. n. Female genital plates: A—ventral view, B—oblique view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911516" httpUri="https://zenodo.org/record/6911516/files/figure.png" pageId="3" pageNumber="504">Fig. 5B</figureCitation>
)......................................... 
<emphasis id="B91AEAC7FFDEFFF3C6DCFE0CFA57FE4B" bold="true" box="[1248,1436,445,466]" pageId="3" pageNumber="504">
<taxonomicName id="4C6E4D56FFDEFFF3C6DCFE0CFAABFE4B" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1248,1376,445,466]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="ocanensis" status="sp. nov.">
<emphasis id="B91AEAC7FFDEFFF3C6DCFE0CFAABFE4B" bold="true" box="[1248,1376,445,466]" italics="true" pageId="3" pageNumber="504">H. ocanensis</emphasis>
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<taxonomicNameLabel id="A22957BCFFDEFFF3C75AFE0CFA57FE4B" box="[1382,1436,445,466]" pageId="3" pageNumber="504" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BD136D5FFDEFFF3C2ABFE6BFA57FE76" blockId="3.[151,1436,151,555]" box="[151,1436,474,496]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFE6BFA57FE76" box="[151,1436,474,496]" pageId="3" pageNumber="504">
11’. Mediolateral crest of valvifers VIII not carinated (
<figureCitation id="13552A50FFDEFFF3C0FEFE6BFCFFFE76" box="[706,820,474,496]" captionStart="FIGURE 8" captionStartId="13.[151,250,1859,1883]" captionTargetBox="[250,1339,204,1800]" captionTargetId="figure-17@13.[199,1389,181,1835]" captionTargetPageId="13" captionText="FIGURE 8. Male genital capsule in posterolateral view, swollen area on inner wall of the pygophore indicated by the arrow: A—Hypoxys triangularis; B—H. capito; C—H. subrastratus; D—H. myrtacivorus sp. n.; E—H. curvatus sp. n.; F—H. striatus sp. n. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.6911526" httpUri="https://zenodo.org/record/6911526/files/figure.png" pageId="3" pageNumber="504">Fig. 8D, 8E</figureCitation>
)................................................ 12
</keyLead>
</paragraph>
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<keyStep id="309A2E70FFDEFFF3C2ABFE49FA57FDB3" pageId="3" pageNumber="504">
<paragraph id="8BD136D5FFDEFFF3C2ABFE49FA57FD94" blockId="3.[151,1436,151,555]" box="[151,1436,503,525]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFE49FA57FD94" box="[151,1436,503,525]" pageId="3" pageNumber="504">
12. Valvifers VIII with mesial borders forming a spiniform median projection (
<figureCitation id="13552A50FFDEFFF3C18FFE46FC30FD94" box="[947,1019,503,525]" captionStart="FIGURE 1" captionStartId="4.[151,250,1905,1929]" captionTargetBox="[182,1399,216,1846]" captionTargetId="figure-25@4.[151,1435,181,1881]" captionTargetPageId="4" captionText="FIGURE 1. Hypoxys myrtacivorus sp. n. Male (pygophore): A—dorsal view, B —posterior view, C—ventral view, D— posterolateral view. Female: E—genital plates. Acronyms: Dr—dorsal rim; Pc—superior process of the genital cup; Pa— parameres; Proc—proctiger; Vr—ventral rim; Vf8—valvifers VIII; Vf9—valvifers IX; Lat 8—laterotergites VIII; Lat 9— laterotergites IX; Va9—valvulae IX; X—tenth segment. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911505" httpUri="https://zenodo.org/record/6911505/files/figure.png" pageId="3" pageNumber="504">Fig. 1E</figureCitation>
)............... 
<emphasis id="B91AEAC7FFDEFFF3C6FCFE49FA57FD94" bold="true" box="[1216,1436,503,525]" pageId="3" pageNumber="504">
<taxonomicName id="4C6E4D56FFDEFFF3C6FCFE49FAABFD95" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1216,1376,503,525]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="myrtacivorus" status="sp. nov.">
<emphasis id="B91AEAC7FFDEFFF3C6FCFE49FAABFD95" bold="true" box="[1216,1376,503,525]" italics="true" pageId="3" pageNumber="504">H. myrtacivorus</emphasis>
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<taxonomicNameLabel id="A22957BCFFDEFFF3C75AFE49FA57FD94" box="[1382,1436,504,525]" pageId="3" pageNumber="504" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BD136D5FFDEFFF3C2ABFDA4FA57FDB3" blockId="3.[151,1436,151,555]" box="[151,1436,533,555]" pageId="3" pageNumber="504">
<keyLead id="309F95E0FFDEFFF3C2ABFDA4FA57FDB3" box="[151,1436,533,555]" pageId="3" pageNumber="504">
12’. Valvifers VIII with mesial borders not forming a spiniform projection. (
<figureCitation id="13552A50FFDEFFF3C1ADFDA4FC11FDB3" box="[913,986,533,555]" captionStart="FIGURE 4" captionStartId="9.[151,250,1895,1919]" captionTargetBox="[201,1400,209,1833]" captionTargetId="figure-17@9.[151,1435,181,1871]" captionTargetPageId="9" captionText="FIGURE 4. Hypoxys curvatus sp. n. Male (pygophore):A—dorsal view, B —posterior view, C—ventral view, D—posterolateral view. Female: E—genital plates. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6911514" httpUri="https://zenodo.org/record/6911514/files/figure.png" pageId="3" pageNumber="504">Fig. 4E</figureCitation>
)..................... 
<emphasis id="B91AEAC7FFDEFFF3C6D0FDA4FA57FDB3" bold="true" box="[1260,1436,533,554]" pageId="3" pageNumber="504">
<taxonomicName id="4C6E4D56FFDEFFF3C6D0FDA4FAABFDB3" authority="Cunha &amp; Fernandes, 2022" authorityName="Cunha &amp; Fernandes" authorityYear="2022" box="[1260,1376,533,554]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="curvatus" status="sp. nov.">
<emphasis id="B91AEAC7FFDEFFF3C6D0FDA4FAABFDB3" bold="true" box="[1260,1376,533,554]" italics="true" pageId="3" pageNumber="504">H. curvatus</emphasis>
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<taxonomicNameLabel id="A22957BCFFDEFFF3C75AFDA4FA57FDB3" box="[1382,1436,533,554]" pageId="3" pageNumber="504" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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