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<document id="480AC6A22FBE0CC858E971B134B9297E" ID-CLB-Dataset="21836" ID-DOI="10.5852/ejt.2012.17" ID-GBIF-Dataset="ac80d266-8f8e-4fc5-a172-599a331449a4" ID-ISSN="2118-9773" ID-Zenodo-Dep="3857844" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590517236165" checkinUser="carolina" docAuthor="Karner, Michael" docDate="2012" docId="03BB8D13E10EFF80FD987B24851FFCAD" docLanguage="en" docName="ejt-2012-17.pdf.imf" docOrigin="European Journal of Taxonomy 17" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Psammoecus spinosus Grouvelle 1882" docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="FF82F56BE117FF9BFFA87F6B8405FFDA" masterDocTitle="A revision of African Psammoecus (Coleoptera, Silvanidae) and descriptions of two new species from the collection of the Musée royal de l’Afrique centrale" masterLastPageNumber="31" masterPageNumber="1" pageNumber="26" updateTime="1698841482259" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="5BD7963C405146D8E8CFC618ED9D1C13" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F2893FFE5F44C4A0A11751D0F2930465">
<mods:title id="8519F081CB8CA9D049192C61CF2DCA11">A revision of African Psammoecus (Coleoptera, Silvanidae) and descriptions of two new species from the collection of the Musée royal de l’Afrique centrale</mods:title>
</mods:titleInfo>
<mods:name id="D1ADD6C364DC2E9F91F11CE0B5C47245" type="personal">
<mods:role id="A9B58985D4F0AA633D6209FC8F5B92DB">
<mods:roleTerm id="5D2368E0C935815C73AB05694A72A2EB">Author</mods:roleTerm>
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<mods:namePart id="3D11157DC9AB75688E5956D54E731253">Karner, Michael</mods:namePart>
</mods:name>
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<mods:title id="C3DC24A2C5347646AFCA48E9D3CEBC79">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="4DE3CEC440703D02A13935A312112DB3">
<mods:date id="D76A53D1051E3FBEBC3A9A57DBAC0B63">2012</mods:date>
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<mods:number id="2FD8EE5CE1F92D396C9081F055834BB6">2012-06-04</mods:number>
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<mods:number id="E583AF3B9FD86E72EBF1A18E8C342E92">17</mods:number>
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<mods:classification id="0E37FEA7D4BF309711C46E88133BA51A">journal article</mods:classification>
<mods:identifier id="DF3CEAF5375BB633B6B0A9DF1F8F81C6" type="CLB-Dataset">21836</mods:identifier>
<mods:identifier id="CE5EB62D071A99C7297AE32BC556B1C6" type="DOI">10.5852/ejt.2012.17</mods:identifier>
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<mods:identifier id="5EE1C0C39EC6DE392D99E5D052796C68" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="605F005FFE2804801B7BBA5B8C10F12F" type="Zenodo-Dep">3857844</mods:identifier>
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<treatment id="03BB8D13E10EFF80FD987B24851FFCAD" ID-DOI="http://doi.org/10.5281/zenodo.3858702" ID-GBIF-Taxon="164302231" ID-Zenodo-Dep="3858702" LSID="urn:lsid:plazi:treatment:03BB8D13E10EFF80FD987B24851FFCAD" httpUri="http://treatment.plazi.org/id/03BB8D13E10EFF80FD987B24851FFCAD" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="C3086F8EE10EFF82FD987B248001FBB2" box="[560,1028,1102,1129]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BAD3C05E10EFF82FD987B248001FBB2" blockId="25.[560,1028,1102,1164]" box="[560,1028,1102,1129]" pageId="25" pageNumber="26">
<heading id="D0E58B69E10EFF82FD987B248001FBB2" box="[560,1028,1102,1129]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="4C124786E10EFF82FD987B248001FBB2" authority="Grouvelle, 1882" authorityName="Grouvelle" authorityYear="1882" box="[560,1028,1102,1129]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B966E017E10EFF82FD987B248738FBB2" bold="true" box="[560,829,1102,1129]" italics="true" pageId="25" pageNumber="26">Psammoecus spinosus</emphasis>
<bibRefCitation id="EF8341F4E10EFF82FCED7B258001FBB2" author="Grouvelle A." box="[837,1028,1102,1128]" pageId="25" pageNumber="26" pagination="275 - 296" refId="ref14966" refString="Grouvelle A. 1882. Cucujides nouveaux du Musee Civique de Genes. Annali del Museo civico di storia naturale Giacomo Doria 18: 275 - 296, pl. 7." type="journal article" year="1882">Grouvelle, 1882</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE10EFF82FD4E7B198748FB56" box="[742,845,1138,1164]" pageId="25" pageNumber="26" type="description">
<paragraph id="8BAD3C05E10EFF82FD4E7B198748FB56" blockId="25.[560,1028,1102,1164]" box="[742,845,1138,1164]" pageId="25" pageNumber="26">
(
<figureCitation id="13292080E10EFF82FD477B198741FB56" box="[751,836,1138,1164]" captionStart="Fig" captionStartId="26.[189,232,1965,1991]" captionTargetBox="[275,1306,729,1878]" captionTargetId="figure@26.[189,1397,667,1925]" captionTargetPageId="26" captionText="Fig. 12. Psammoecus spinosus Grouvelle,1882, specimen from Bingerville. A. Habitus. B. Right antenna. C. Parameres. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857870" httpUri="https://zenodo.org/record/3857870/files/figure.png" pageId="25" pageNumber="26">Fig. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE10EFF82FF157BD8852AFA2E" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BAD3C05E10EFF82FF157BD885AFFB17" blockId="25.[189,426,1203,1229]" box="[189,426,1203,1229]" pageId="25" pageNumber="26">
<heading id="D0E58B69E10EFF82FF157BD885AFFB17" bold="true" box="[189,426,1203,1229]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B966E017E10EFF82FF157BD885AFFB17" bold="true" box="[189,426,1203,1229]" pageId="25" pageNumber="26">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E10EFF82FF157B89852AFA2E" blockId="25.[189,1399,1250,1524]" pageId="25" pageNumber="26">
1 spm ‘I.R.S.A.C.-MUS.
<collectingCountry id="F3057C95E10EFF82FE757B898651FB26" box="[477,596,1250,1276]" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">CONGO</collectingCountry>
|
<collectingRegion id="49D6F2E7E10EFF82FDDD7B8986B1FB26" box="[629,692,1250,1276]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="25" pageNumber="26">Kivu</collectingRegion>
: Kavimvira (Uvira) | (à la lumière)
<date id="FFAC1AC5E10EFF82FBD57B8980C9FB27" box="[1149,1228,1250,1277]" pageId="25" pageNumber="26" value="1956-01">I-1956</date>
| G. Marlier’ (
<collectionCode id="ED03A4C0E10EFF82FF6D7A6D8527FAFA" box="[197,290,1286,1312]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="25" pageNumber="26" type="Museum">MRAC</collectionCode>
). 1 spm ‘I.R.S.A.C.-MUS.
<collectingCountry id="F3057C95E10EFF82FDE67A6D86C0FAFA" box="[590,709,1286,1312]" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">CONGO</collectingCountry>
|
<collectingRegion id="49D6F2E7E10EFF82FD757A6E8719FAC5" box="[733,796,1285,1311]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="25" pageNumber="26">Kivu</collectingRegion>
: Kavimvira (Uvira) | (à la lumière) II/
<date id="FFAC1AC5E10EFF82FB4A7A6E8146FAFA" box="[1250,1347,1285,1312]" pageId="25" pageNumber="26" value="1956-03">III-1956</date>
| G. Marlier’ (
<collectionCode id="ED03A4C0E10EFF82FE857A42858FFA99" box="[301,394,1321,1347]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="25" pageNumber="26" type="Museum">MRAC</collectionCode>
). 1 spm ‘COLL. MUS.
<collectingCountry id="F3057C95E10EFF82FD087A42870DFA99" box="[672,776,1321,1347]" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">CONGO</collectingCountry>
| Tanganika: Moba,
<specimenCount id="9D14F78CE10EFF82FC517A42804EFA99" box="[1017,1099,1321,1347]" pageId="25" pageNumber="26" type="generic">
<quantity id="4CEA91E0E10EFF82FC517A428042FA99" box="[1017,1095,1321,1347]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="25" pageNumber="26" unit="m" value="780.0">780 m</quantity>
.
</specimenCount>
| (à la lumière)
<date id="FFAC1AC5E10EFF82FAAE7A42816CFA9E" box="[1286,1385,1321,1348]" pageId="25" pageNumber="26" value="1953-11">XI-1953</date>
| H. Bomans’ (
<collectionCode id="ED03A4C0E10EFF82FEFE7A2785B6FABC" box="[342,435,1356,1382]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="25" pageNumber="26" type="Museum">MRAC</collectionCode>
). 1 spm ‘
<collectingCountry id="F3057C95E10EFF82FDBD7A278670FABC" box="[533,629,1356,1382]" name="Angola" pageId="25" pageNumber="26">Angola</collectingCountry>
: Cazombo, | Alto Zambeze
<date id="FFAC1AC5E10EFF82FC1B7A27800EFABD" box="[947,1035,1356,1383]" pageId="25" pageNumber="26" value="1955-02">II-1955</date>
| 4987.1 | E. Luna de Carvalho’ (
<collectionCode id="ED03A4C0E10EFF82FF6D7A1B8527FA50" box="[197,290,1392,1418]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="25" pageNumber="26" type="Museum">MRAC</collectionCode>
).
<specimenCount id="9D14F78CE10EFF82FE9F7A1B855EFA50" box="[311,347,1392,1418]" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
‘
<collectingCountry id="F3057C95E10EFF82FEC07A1B85C0FA50" box="[360,453,1392,1418]" name="Nigeria" pageId="25" pageNumber="26">Nigeria</collectingCountry>
VdoFR | m State |
<date id="FFAC1AC5E10EFF82FD037A1B8708FA51" box="[683,781,1392,1419]" pageId="25" pageNumber="26" value="1975-04">IV 1975</date>
| J. T. Nedler Coll.’ (
<collectionCode id="ED03A4C0E10EFF82FBA07A1B8060FA50" box="[1032,1125,1392,1418]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="25" pageNumber="26" type="Museum">MRAC</collectionCode>
). 1 spm ‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E10EFF82FEC67AF8860BFA77" box="[366,526,1427,1453]" name="Ivory Coast" pageId="25" pageNumber="26">Côte d’ivoire</collectingCountry>
: Bingerville |
<date id="FFAC1AC5E10EFF82FD1D7AF8870EFA77" box="[693,779,1427,1453]" pageId="25" pageNumber="26" value="1962-02">II.1962</date>
| J. Decelle’, ‘
<taxonomicName id="4C124786E10EFF82FC1C7AF88056FA77" authorityName="Latreille" authorityYear="1829" box="[948,1107,1427,1453]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="30" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
sp. | D. G. H. Halstead | det. 1978’ (
<collectionCode id="ED03A4C0E10EFF82FE967ADC8581FA0B" box="[318,388,1463,1489]" pageId="25" pageNumber="26">MKF</collectionCode>
).
<specimenCount id="9D14F78CE10EFF82FE3D7ADC85BFFA0B" box="[405,442,1463,1489]" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E10EFF82FCB27ADC87BDFA0A" box="[794,952,1462,1489]" name="Ivory Coast" pageId="25" pageNumber="26">Côte d’ivoire</collectingCountry>
: Bingerville | I/
<date id="FFAC1AC5E10EFF82FBC27ADD80E9FA0A" box="[1130,1260,1462,1488]" pageId="25" pageNumber="26" value="1962-03-12">12.III.1962</date>
| J. Decelle’ (
<collectionCode id="ED03A4C0E10EFF82FF6D7AB18527FA2E" box="[197,290,1498,1524]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="25" pageNumber="26" type="Museum">MRAC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE10EFF82FF15797086CFF952" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8BAD3C05E10EFF82FF15797085CCF9EF" blockId="25.[189,457,1563,1589]" box="[189,457,1563,1589]" pageId="25" pageNumber="26">
<heading id="D0E58B69E10EFF82FF15797085CCF9EF" bold="true" box="[189,457,1563,1589]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B966E017E10EFF82FF15797085CCF9EF" bold="true" box="[189,457,1563,1589]" pageId="25" pageNumber="26">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E10EFF82FF15792086CFF952" blockId="25.[189,1399,1610,1672]" pageId="25" pageNumber="26">
<taxonomicName id="4C124786E10EFF82FF157920854AF9BE" authorityName="Grouvelle" authorityYear="1882" box="[189,335,1611,1636]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B966E017E10EFF82FF157920854AF9BE" box="[189,335,1611,1636]" italics="true" pageId="25" pageNumber="26">Ps. spinosus</emphasis>
</taxonomicName>
differs by the very characteristic shape of the pronotum and the distinctly toothed elytral margins from all other African
<taxonomicName id="4C124786E10EFF82FD82790486C6F952" authorityName="Latreille" authorityYear="1829" box="[554,707,1647,1672]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B966E017E10EFF82FD82790486C6F952" box="[554,707,1647,1672]" italics="true" pageId="25" pageNumber="26">Psammoecus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE10EFF80FF1579C487ADFD6C" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="8BAD3C05E10EFF82FF1579C4856EF913" blockId="25.[189,363,1711,1737]" box="[189,363,1711,1737]" pageId="25" pageNumber="26">
<heading id="D0E58B69E10EFF82FF1579C4856EF913" bold="true" box="[189,363,1711,1737]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B966E017E10EFF82FF1579C4856EF913" bold="true" box="[189,363,1711,1737]" pageId="25" pageNumber="26">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E10EFF82FF1579B486CCF85C" blockId="25.[189,1399,1758,1926]" pageId="25" pageNumber="26">
BODY. Oval, total length
<quantity id="4CEA91E0E10EFF82FE5C79B5869BF923" box="[500,670,1758,1785]" metricMagnitude="-3" metricUnit="m" metricValue="2.775" metricValueMax="3.15" metricValueMin="2.4" pageId="25" pageNumber="26" unit="mm" value="2.775" valueMax="3.15" valueMin="2.4">2.40-3.15 mm</quantity>
(
<figureCitation id="13292080E10EFF82FD1B79B58720F922" box="[691,805,1758,1784]" captionStart="Fig" captionStartId="26.[189,232,1965,1991]" captionTargetBox="[275,1306,729,1878]" captionTargetId="figure@26.[189,1397,667,1925]" captionTargetPageId="26" captionText="Fig. 12. Psammoecus spinosus Grouvelle,1882, specimen from Bingerville. A. Habitus. B. Right antenna. C. Parameres. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857870" httpUri="https://zenodo.org/record/3857870/files/figure.png" pageId="25" pageNumber="26">Fig. 12A</figureCitation>
), reddish-brown, elytra with darkened basis and with dark lateral macula near their middle, connected to the darkened suture in posterior half which is connected to another lateral macula close to the apex; the maculae forming a roughly x-shaped pattern on posterior two thirds of elytra. Antennae reddish-brown, 7
<superScript id="7C67914DE10EFF82FC22782C8792F88F" attach="left" box="[906,919,1863,1877]" fontSize="6" pageId="25" pageNumber="26">th</superScript>
to 10
<superScript id="7C67914DE10EFF82FC75782C87EFF88F" attach="left" box="[989,1002,1863,1877]" fontSize="6" pageId="25" pageNumber="26">th</superScript>
antennomeres darker brown, 11
<superScript id="7C67914DE10EFF82FAC1782C8173F88F" attach="none" box="[1385,1398,1863,1877]" fontSize="6" pageId="25" pageNumber="26">th</superScript>
antennomere bright yellowish, almost white.
</paragraph>
<paragraph id="8BAD3C05E10EFF81FF1578C587C6FE68" blockId="25.[189,1399,1965,2027]" lastBlockId="26.[189,1399,267,435]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
HEAD. Wide, temples strongly narrowed immediately behind eyes; width
<quantity id="4CEA91E0E10EFF82FB8578C680DDF812" box="[1069,1240,1965,1992]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="25" pageNumber="26" unit="mm" value="0.675" valueMax="0.75" valueMin="0.6">0.60-0.75 mm</quantity>
, length
<quantity id="4CEA91E0E10EFF82FA9178C68528F831" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.5" metricValueMin="3.8" pageId="25" pageNumber="26" unit="mm" value="0.415" valueMax="0.45" valueMin="0.38">0.38- 0.45 mm</quantity>
, 1.58-1.71 times as wide as long. Eyes very large and protuberant, length
<quantity id="4CEA91E0E10EFF82FB6E78BA8176F831" box="[1222,1395,2001,2027]" metricMagnitude="-4" metricUnit="m" metricValue="2.0500000000000003" metricValueMax="2.3000000000000003" metricValueMin="1.7999999999999998" pageId="25" pageNumber="26" unit="mm" value="0.205" valueMax="0.23" valueMin="0.18">0.18-0.23 mm</quantity>
, distance of inner margins
<quantity id="4CEA91E0E10DFF81FE567E6086AEFEFC" box="[510,683,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.8" metricValueMin="3.5999999999999996" pageId="26" pageNumber="27" unit="mm" value="0.42" valueMax="0.48" valueMin="0.36">0.36-0.48 mm</quantity>
. Puncturation on vertex irregular, punctures comparatively small and sparse, pubescence composed of moderately long, semierect setae, microsculpture absent. Longitudinal impressions on vertex very distinct, deep, slightly curved, attaining slightly beyond the middle of eyes. Antennae as in
<figureCitation id="13292080E10DFF81FD8E7E1E868BFE55" box="[550,654,373,399]" captionStart="Fig" captionStartId="26.[189,232,1965,1991]" captionTargetBox="[275,1306,729,1878]" captionTargetId="figure@26.[189,1397,667,1925]" captionTargetPageId="26" captionText="Fig. 12. Psammoecus spinosus Grouvelle,1882, specimen from Bingerville. A. Habitus. B. Right antenna. C. Parameres. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857870" httpUri="https://zenodo.org/record/3857870/files/figure.png" pageId="26" pageNumber="27">Fig. 12B</figureCitation>
, long and slender,
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long; antennomere proportions as follows: 2.8: 1.0: 1.3: 1.2: 1.3: 1.4: 1.3: 1.0: 1.0: 1.1: 2.2.
</paragraph>
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PRONOTUM. Transverse, posterior and lateral margins with distinct impressions, close to the middle of the anterior margin with shallow impression; width
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, length
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, 1.38-1.44 times as wide as long. Anterior angle with very distinct group or large teeth; lateral margin characteristic pattern of four teeth; tooth I large with wide basis, teeth II and III almost fused, forming a large forked tooth, tooth IV as large as tooth I; posterior angle with small group of teeth. Anterior margin with a small short tooth right between anterior group of teeth and middle of anterior margin. Puncturation on disc somewhat coarser as on vertex, pubescence as on vertex, microsculpture absent.
</paragraph>
<caption id="DF6D6C8DE10DFF81FF1578C6876EF831" ID-DOI="http://doi.org/10.5281/zenodo.3857870" ID-Zenodo-Dep="3857870" httpUri="https://zenodo.org/record/3857870/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1965,1991]" targetBox="[275,1306,729,1878]" targetPageId="26">
<paragraph id="8BAD3C05E10DFF81FF1578C6876EF831" blockId="26.[189,1398,1965,2027]" pageId="26" pageNumber="27">
<emphasis id="B966E017E10DFF81FF1578C68525F81D" bold="true" box="[189,288,1965,1991]" pageId="26" pageNumber="27">Fig. 12.</emphasis>
<taxonomicName id="4C124786E10DFF81FE8678C58707F81D" authority="Grouvelle, 1882" authorityName="Grouvelle" authorityYear="1882" box="[302,770,1965,1991]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B966E017E10DFF81FE8678C5863FF81D" box="[302,570,1966,1991]" italics="true" pageId="26" pageNumber="27">Psammoecus spinosus</emphasis>
<bibRefCitation id="EF8341F4E10DFF81FDE078C68707F81D" author="Grouvelle A." box="[584,770,1965,1991]" pageId="26" pageNumber="27" pagination="275 - 296" refId="ref14966" refString="Grouvelle A. 1882. Cucujides nouveaux du Musee Civique de Genes. Annali del Museo civico di storia naturale Giacomo Doria 18: 275 - 296, pl. 7." type="journal article" year="1882">Grouvelle,1882</bibRefCitation>
</taxonomicName>
, specimen from Bingerville.
<emphasis id="B966E017E10DFF81FBD978C68083F81D" bold="true" box="[1137,1158,1965,1991]" pageId="26" pageNumber="27">A</emphasis>
. Habitus.
<emphasis id="B966E017E10DFF81FAA478C58125F812" bold="true" box="[1292,1312,1966,1992]" pageId="26" pageNumber="27">B</emphasis>
. Right antenna.
<emphasis id="B966E017E10DFF81FE8F78BB8539F830" bold="true" box="[295,316,2000,2026]" pageId="26" pageNumber="27">C</emphasis>
. Parameres. Scale line A, B: 1 mm; C: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BAD3C05E10CFF80FF157EF085CFFD96" blockId="27.[189,1399,411,588]" pageId="27" pageNumber="28">
ELYTRA. Oval, length
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, combined width
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, 1.46-1.59 times as long as combined width. Rows of punctures on the disc about as wide as interstices; pubescence composed of long semierect setae, directed posteriorly; interstices of lateral rows of punctures with tubercles that are enlarged to form distinct teeth; towards the elytral disc, these teeth become successively smaller. Microsculpture absent.
</paragraph>
<paragraph id="8BAD3C05E10CFF80FF157D1C87ADFD6C" blockId="27.[189,1399,630,694]" pageId="27" pageNumber="28">
PARAMERES. Simple, narrowed towards apex, without well-defined basal part. Inner portion of parameres with numerous small setae; apex with one large seta (
<figureCitation id="13292080E10CFF80FC877DF7879EFD6C" box="[815,923,668,694]" captionStart="Fig" captionStartId="26.[189,232,1965,1991]" captionTargetBox="[275,1306,729,1878]" captionTargetId="figure@26.[189,1397,667,1925]" captionTargetPageId="26" captionText="Fig. 12. Psammoecus spinosus Grouvelle,1882, specimen from Bingerville. A. Habitus. B. Right antenna. C. Parameres. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857870" httpUri="https://zenodo.org/record/3857870/files/figure.png" pageId="27" pageNumber="28">Fig. 12C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE10CFF80FF157D8B851FFCAD" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BAD3C05E10CFF80FF157D8B852AFD20" blockId="27.[189,303,736,762]" box="[189,303,736,762]" pageId="27" pageNumber="28">
<heading id="D0E58B69E10CFF80FF157D8B852AFD20" bold="true" box="[189,303,736,762]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B966E017E10CFF80FF157D8B852AFD20" bold="true" box="[189,303,736,762]" pageId="27" pageNumber="28">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E10CFF80FF157C79851FFCAD" blockId="27.[189,1398,785,887]" pageId="27" pageNumber="28">
It has not been possible to locate and study
<typeStatus id="54A982A7E10CFF80FD607C7986FEFCF6" box="[712,763,786,812]" pageId="27" pageNumber="28">type</typeStatus>
material of
<taxonomicName id="4C124786E10CFF80FC257C798025FCF1" authorityName="Grouvelle" authorityYear="1882" box="[909,1056,786,811]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B966E017E10CFF80FC257C798025FCF1" box="[909,1056,786,811]" italics="true" pageId="27" pageNumber="28">Ps. spinosus</emphasis>
</taxonomicName>
. However,
<bibRefCitation id="EF8341F4E10CFF80FB027C7A8173FCF1" author="Grouvelle A." box="[1194,1398,785,811]" pageId="27" pageNumber="28" pagination="275 - 296" refId="ref14966" refString="Grouvelle A. 1882. Cucujides nouveaux du Musee Civique de Genes. Annali del Museo civico di storia naturale Giacomo Doria 18: 275 - 296, pl. 7." type="journal article" year="1882">Grouvelle (1882)</bibRefCitation>
provides an illustration that leaves little doubt regarding the identity of this species, given its peculiar habitus.
</paragraph>
</subSubSection>
</treatment>
</document>
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