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<document id="395C4BCAE943070FE403FB46C75108C9" ID-CLB-Dataset="21836" ID-DOI="10.5852/ejt.2012.17" ID-GBIF-Dataset="ac80d266-8f8e-4fc5-a172-599a331449a4" ID-ISSN="2118-9773" ID-Zenodo-Dep="3857844" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590517236165" checkinUser="carolina" docAuthor="Karner, Michael" docDate="2012" docId="03BB8D13E100FF8CFDAF7E67865BFA99" docLanguage="en" docName="ejt-2012-17.pdf.imf" docOrigin="European Journal of Taxonomy 17" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Psammoecus trimaculatus Motschulsky 1858" docType="treatment" docVersion="7" lastPageNumber="24" masterDocId="FF82F56BE117FF9BFFA87F6B8405FFDA" masterDocTitle="A revision of African Psammoecus (Coleoptera, Silvanidae) and descriptions of two new species from the collection of the Musée royal de lAfrique centrale" masterLastPageNumber="31" masterPageNumber="1" pageNumber="24" updateTime="1698841482259" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8E578403175F3C1431E7D9FECDD04A3E">A revision of African Psammoecus (Coleoptera, Silvanidae) and descriptions of two new species from the collection of the Musée royal de lAfrique centrale</mods:title>
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<mods:namePart id="1EB2E2269258B824565534C7FAD47A08">Karner, Michael</mods:namePart>
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<mods:title id="586ADBC875C6BCB35B2B56431358EE9C">European Journal of Taxonomy</mods:title>
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<mods:date id="AC7182220A9128EE9E99B7B508AEB109">2012</mods:date>
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<treatment id="03BB8D13E100FF8CFDAF7E67865BFA99" ID-DOI="http://doi.org/10.5281/zenodo.3858700" ID-GBIF-Taxon="164302236" ID-Zenodo-Dep="3858700" LSID="urn:lsid:plazi:treatment:03BB8D13E100FF8CFDAF7E67865BFA99" httpUri="http://treatment.plazi.org/id/03BB8D13E100FF8CFDAF7E67865BFA99" lastPageNumber="24" pageId="23" pageNumber="24">
<subSubSection id="C3086F8EE100FF8CFDAF7E678028FEFC" box="[519,1069,267,294]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BAD3C05E100FF8CFDAF7E678028FEFC" blockId="23.[519,1069,267,294]" box="[519,1069,267,294]" pageId="23" pageNumber="24">
<heading id="D0E58B69E100FF8CFDAF7E678028FEFC" box="[519,1069,267,294]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="4C124786E100FF8CFDAF7E678028FEFC" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[519,1069,267,294]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="B966E017E100FF8CFDAF7E678743FEFC" bold="true" box="[519,838,267,294]" italics="true" pageId="23" pageNumber="24">Psammoecus trimaculatus</emphasis>
Motschulsky, 1858
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE100FF8CFF157E248739FE54" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="8BAD3C05E100FF8CFF157E248754FEB3" blockId="23.[189,849,334,398]" box="[189,849,334,361]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB021B2BE100FF8CFF157E248754FEB3" box="[189,849,334,361]" pageId="23" pageNumber="24">
<treatmentCitation id="0AB31A14E100FF8CFF157E2486DCFEB3" author="Grouvelle A." box="[189,729,334,361]" page="115" pageId="23" pageNumber="24" year="1908">
<taxonomicName id="4C124786E100FF8CFF157E2486DCFEB3" authority="Grouvelle, 1908 a: 115" authorityName="Grouvelle" authorityPageNumber="115" authorityYear="1908" box="[189,729,334,361]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="B966E017E100FF8CFF157E2485CCFEB2" box="[189,457,334,360]" italics="true" pageId="23" pageNumber="24">Psammoecus excellens</emphasis>
<bibRefCitation id="EF8341F4E100FF8CFE787E2586DCFEB3" author="Grouvelle A." box="[464,729,334,361]" pageId="23" pageNumber="24" pagination="97 - 126" refId="ref15074" refString="Grouvelle A. 1908 a. Clavicornes recoltes dans l'Afrique orientale Allemande par le Dr. F. Eichelbaum (2 e liste). Revue d'Entomologie 27: 97 - 126." type="journal article" year="1908">Grouvelle, 1908a: 115</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B966E017E100FF8CFD4D7E248754FEB3" bold="true" box="[741,849,335,361]" pageId="23" pageNumber="24">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAD3C05E100FF8CFF157E1F8739FE54" blockId="23.[189,849,334,398]" box="[189,828,371,398]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB021B2BE100FF8CFF157E1F8739FE54" box="[189,828,371,398]" pageId="23" pageNumber="24">
<treatmentCitation id="0AB31A14E100FF8CFF157E1F86C0FE54" author="Grouvelle A." box="[189,709,371,398]" page="409" pageId="23" pageNumber="24" year="1912">
<taxonomicName id="4C124786E100FF8CFF157E1F86C0FE54" authority="Grouvelle, 1912: 409" authorityName="Grouvelle" authorityPageNumber="409" authorityYear="1912" box="[189,709,371,398]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B966E017E100FF8CFF157E1F85BAFE57" box="[189,447,371,397]" italics="true" pageId="23" pageNumber="24">Psammoecus alluaudi</emphasis>
<bibRefCitation id="EF8341F4E100FF8CFE6F7E1886C0FE54" author="Grouvelle A." box="[455,709,371,398]" pageId="23" pageNumber="24" pagination="409 - 414" refId="ref15165" refString="Grouvelle A. 1912. Coleopteres cucujides nouveaux du genre Psammoecus faisant partie des collections du Museum. Bulletin du Museum d'Histoire Naturelle de Paris 18: 409 - 414." type="journal article" year="1912">Grouvelle, 1912: 409</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B966E017E100FF8CFD677E1F8739FE54" bold="true" box="[719,828,372,398]" pageId="23" pageNumber="24">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE100FF8CFF157EDD8691FD05" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BAD3C05E100FF8CFF157EDD85AFFE0A" blockId="23.[189,426,438,464]" box="[189,426,438,464]" pageId="23" pageNumber="24">
<heading id="D0E58B69E100FF8CFF157EDD85AFFE0A" bold="true" box="[189,426,438,464]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B966E017E100FF8CFF157EDD85AFFE0A" bold="true" box="[189,426,438,464]" pageId="23" pageNumber="24">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E100FF8CFF157E8C8691FD05" blockId="23.[189,1399,486,735]" pageId="23" pageNumber="24">
<specimenCount id="9D14F78CE100FF8CFF157E8C84E7FDDB" box="[189,226,487,513]" pageId="23" pageNumber="24" type="female">1♀</specimenCount>
[round, red label]; [yellow label]; Psammecus [sic!] | trimaculatus | Motch. | Ceylon [Motschulskys hand] (
<collectionCode id="ED03A4C0E100FF8CFEB87D678577FDFC" box="[272,370,524,550]" country="China" name="Zoological Museum, University of Amoy" pageId="23" pageNumber="24">ZMUM</collectionCode>
). 3 spms Af. or. All. | Eichelbaum [Grouvelles hand], Type [red label], MUSEUM PARIS | 1917 | Coll. GROUVELLE [blue label],
<taxonomicName id="4C124786E100FF8CFC8D7D5A87C1FD91" authorityName="Latreille" authorityYear="1829" box="[805,964,561,587]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="30" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
| excellens | G Grouv [Grouvelles hand] (
<collectionCode id="ED03A4C0E100FF8CFEB87D3D8574FDAA" box="[272,369,598,624]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
). One
<specimenCount id="9D14F78CE100FF8CFE167D3D85D1FDAA" box="[446,468,598,624]" pageId="23" pageNumber="24" type="male"></specimenCount>
specimen was designated as lectotpyus of
<taxonomicName id="4C124786E100FF8CFC787D3C80DAFDAA" authority="Grouvelle" authorityName="Grouvelle" authorityYear="1908" box="[976,1247,598,624]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="B966E017E100FF8CFC787D3C8067FDAA" box="[976,1122,598,624]" italics="true" pageId="23" pageNumber="24">Ps excellens</emphasis>
Grouvelle
</taxonomicName>
and labelled accordingly.
<typeStatus id="54A982A7E100FF8CFEFE7D1085C6FD4F" box="[342,451,635,661]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C124786E100FF8CFE447D178671FD4F" authorityName="Grouvelle" authorityYear="1912" box="[492,628,635,661]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B966E017E100FF8CFE447D178671FD4F" box="[492,628,635,661]" italics="true" pageId="23" pageNumber="24">Ps alluaudi</emphasis>
</taxonomicName>
.
<specimenCount id="9D14F78CE100FF8CFD2C7D10869BFD4F" box="[644,670,635,661]" pageId="23" pageNumber="24" type="male"></specimenCount>
:
<collectingCountry id="F3057C95E100FF8CFD077D108747FD4F" box="[687,834,635,661]" name="Madagascar" pageId="23" pageNumber="24">Madagascar</collectingCountry>
| Su[???]ieville [blue label, Grouvelles hand], Type [red label], MUSEUM PARIS | 1917 | Coll. GROUVELLE [blue label],
<taxonomicName id="4C124786E100FF8CFBF67DCB80F8FD60" authorityName="Latreille" authorityYear="1829" box="[1118,1277,672,698]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="30" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
| alluaudi | G. Grouv [Grouvelles hand] (
<collectionCode id="ED03A4C0E100FF8CFD8F7DAE868DFD05" box="[551,648,709,735]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE100FF8CFF157C63865BFA99" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BAD3C05E100FF8CFF157C63852AFCF8" blockId="23.[189,303,776,802]" box="[189,303,776,802]" pageId="23" pageNumber="24">
<heading id="D0E58B69E100FF8CFF157C63852AFCF8" bold="true" box="[189,303,776,802]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B966E017E100FF8CFF157C63852AFCF8" bold="true" box="[189,303,776,802]" pageId="23" pageNumber="24">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E100FF8CFF157C5380EBFC47" blockId="23.[189,1399,824,925]" pageId="23" pageNumber="24">
<bibRefCitation id="EF8341F4E100FF8CFF157C53853EFC89" author="Pal T. K." box="[189,315,824,851]" pageId="23" pageNumber="24" pagination="1 - 54" refId="ref15382" refString="Pal T. K. 1985. A revision of Indian Psammoecus Latreille (Coleoptera, Silvanidae). Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper 71: 1 - 54." type="journal article" year="1985">Pal (1985)</bibRefCitation>
gave a redescription of
<taxonomicName id="4C124786E100FF8CFDC97C528721FC88" box="[609,804,824,850]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="B966E017E100FF8CFDC97C528721FC88" box="[609,804,824,850]" italics="true" pageId="23" pageNumber="24">Ps. trimaculatus</emphasis>
</taxonomicName>
based on material from Motschulskys collection (Museum Moscow) that he accepted as type material. He did not designate a
<typeStatus id="54A982A7E100FF8CFBCB7C3680D7FCAD" box="[1123,1234,861,887]" pageId="23" pageNumber="24" type="lectotype">lectotype</typeStatus>
. The material consisted of five specimens mounted on a single label (Pal 1996, personal communication).
</paragraph>
<paragraph id="8BAD3C05E100FF8CFF157CAD8792FBF3" blockId="23.[189,1399,965,1065]" pageId="23" pageNumber="24">
It was not possible for the present author to see this material, but he had the opportunity to study a single, female specimen from Motschulskys collection (ZMUM). This specimen is not assumed to be a
<typeStatus id="54A982A7E100FF8CFA957C818177FBDE" box="[1341,1394,1002,1028]" pageId="23" pageNumber="24">type</typeStatus>
, while the material mentioned by Pal may be discovered again.
</paragraph>
<paragraph id="8BAD3C05E100FF8CFF157B39866FFB01" blockId="23.[189,1399,1106,1243]" pageId="23" pageNumber="24">
<bibRefCitation id="EF8341F4E100FF8CFF157B398539FBB7" author="Pal T. K." box="[189,316,1106,1133]" pageId="23" pageNumber="24" pagination="1 - 54" refId="ref15382" refString="Pal T. K. 1985. A revision of Indian Psammoecus Latreille (Coleoptera, Silvanidae). Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper 71: 1 - 54." type="journal article" year="1985">Pal (1985)</bibRefCitation>
provided detailed figures of
<taxonomicName id="4C124786E100FF8CFD097B388760FBB6" box="[673,869,1106,1132]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="B966E017E100FF8CFD097B388760FBB6" box="[673,869,1106,1132]" italics="true" pageId="23" pageNumber="24">Ps. trimaculatus</emphasis>
</taxonomicName>
, including the male genitalia, and gives an overview on the taxonomy. He states that it occurs in
<collectingCountry id="F3057C95E100FF8CFCCA7B1C87A4FB4B" box="[866,929,1143,1169]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
,
<collectingCountry id="F3057C95E100FF8CFC1A7B1C87FEFB4B" box="[946,1019,1143,1169]" name="Nepal" pageId="23" pageNumber="24">Nepal</collectingCountry>
,
<collectingCountry id="F3057C95E100FF8CFBA37B1C8066FB4B" box="[1035,1123,1143,1169]" name="Bhutan" pageId="23" pageNumber="24">Bhutan</collectingCountry>
,
<collectingCountry id="F3057C95E100FF8CFBDB7B1C80EBFB4B" box="[1139,1262,1143,1169]" name="Sri Lanka" pageId="23" pageNumber="24">Sri Lanka</collectingCountry>
,
<collectingCountry id="F3057C95E100FF8CFB567B138177FB48" box="[1278,1394,1144,1170]" name="Myanmar" pageId="23" pageNumber="24">Myanmar</collectingCountry>
,
<collectingCountry id="F3057C95E100FF8CFF157BF78529FB6C" box="[189,300,1180,1206]" name="Malaysia" pageId="23" pageNumber="24">Malaysia</collectingCountry>
,
<collectingCountry id="F3057C95E100FF8CFE9E7BF785A0FB6C" box="[310,421,1180,1206]" name="Australia" pageId="23" pageNumber="24">Australia</collectingCountry>
,
<collectingCountry id="F3057C95E100FF8CFE187BF785F3FB6C" box="[432,502,1180,1206]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
, and
<collectingCountry id="F3057C95E100FF8CFD9B7BF786C6FB6C" box="[563,707,1180,1206]" name="Madagascar" pageId="23" pageNumber="24">Madagascar</collectingCountry>
. Recently, it has also been recorded as being established in
<collectingCountry id="F3057C95E100FF8CFF157BAA8506FB01" box="[189,259,1217,1243]" name="Brazil" pageId="23" pageNumber="24">Brasil</collectingCountry>
(
<bibRefCitation id="EF8341F4E100FF8CFEBA7BAA865BFB01" author="Thomas M. C. &amp; Yamamoto P. T." box="[274,606,1217,1243]" pageId="23" pageNumber="24" pagination="612 - 613" refId="ref15420" refString="Thomas M. C. &amp; Yamamoto P. T. 2007. New records of Old World Silvanidae in the New World (Coleoptera: Cucujoidea). Coleopterists Bulletin 61 (4): 612 - 613. http: // dx. doi. org / 10.1649 / 0010 - 065 X (2007) 61 [612: NROOWS] 2.0. CO; 2" type="journal article" year="2007">Thomas &amp; Yamamoto 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8BAD3C05E100FF8CFF157A6F865BFA99" blockId="23.[189,1399,1284,1347]" pageId="23" pageNumber="24">
Both
<taxonomicName id="4C124786E100FF8CFF577A6E8613FAC4" authority="Grouvelle" authorityName="Grouvelle" authorityYear="1908" box="[255,534,1284,1310]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="B966E017E100FF8CFF577A6E859CFAC4" box="[255,409,1284,1310]" italics="true" pageId="23" pageNumber="24">Ps. excellens</emphasis>
Grouvelle
</taxonomicName>
and
<taxonomicName id="4C124786E100FF8CFDFB7A6E8765FAC4" authority="Grouvelle" authorityName="Grouvelle" authorityYear="1912" box="[595,864,1284,1310]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B966E017E100FF8CFDFB7A6E86E6FAC4" box="[595,739,1284,1310]" italics="true" pageId="23" pageNumber="24">Ps. alluaudi</emphasis>
Grouvelle
</taxonomicName>
are conspecific and have to be synonymised with
<taxonomicName id="4C124786E100FF8CFF507A41865BFA99" authority="Motschulsky." box="[248,606,1321,1347]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="B966E017E100FF8CFF507A4185BDFA99" box="[248,440,1321,1347]" italics="true" pageId="23" pageNumber="24">Ps. trimaculatus</emphasis>
Motschulsky.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>