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<document id="E28AAFB2A94828950934BEC3CEEE5D41" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C567E0C2B51BC80E8FE1B497F" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola dunini Zonstein &amp; Kunt &amp; Yağmur 2018, sp. nov." docType="treatment" docUuid="91C857A7-FBD3-4676-8F5C-5C7795232F57" docUuidSource="ZooBank" docVersion="14" lastPageNumber="13" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="10" updateTime="1698518020641" updateUser="ExternalLinkService">
<mods:mods id="0A875A6BEB81CCBB157234B11A8DB721" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5BFD213A8C5D248CF4E6EF9593BFFB76">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="9A536227C6239ACDC23C42442A01FB31">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="CB8802CC5E259816D9E856C5D32B7642">Kunt, Kadir B.</mods:namePart>
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<mods:namePart id="9ADAC1FAABAA05D88A791AE10D6F96CE">Yağmur, Ersen A.</mods:namePart>
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<mods:title id="2B3EAD14DCF74748C0187E856F0D3490">European Journal of Taxonomy</mods:title>
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<mods:date id="E58C859EEC12AB4BBAFEB8821E52D10B">2018</mods:date>
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<mods:number id="90113C721908B18C1DD1B441E9AD5E32">2018-01-30</mods:number>
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<treatment id="03B9B44C567E0C2B51BC80E8FE1B497F" ID-DOI="http://doi.org/10.5281/zenodo.5980279" ID-GBIF-Taxon="140561335" ID-Zenodo-Dep="5980279" LSID="urn:lsid:zoobank.org:act:91C857A7-FBD3-4676-8F5C-5C7795232F57" httpUri="http://treatment.plazi.org/id/03B9B44C567E0C2B51BC80E8FE1B497F" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<subSubSection id="C30A56D1567E0C2E51BC80E8FC6E4E13" box="[640,948,1531,1558]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BAF055A567E0C2E51BC80E8FC6E4E13" blockId="9.[374,1214,1531,1635]" box="[640,948,1531,1558]" pageId="9" pageNumber="10">
<heading id="D0E7B236567E0C2E51BC80E8FC6E4E13" box="[640,948,1531,1558]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="4C107ED9567E0C2E51BC80E8FC884E13" ID-CoL="4RMTQ" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[640,850,1531,1558]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dunini" status="sp. nov.">
<emphasis id="B964D948567E0C2E51BC80E8FC884E13" bold="true" box="[640,850,1531,1558]" italics="true" pageId="9" pageNumber="10">Raveniola dunini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2576433567E0C2E506580EFFC6E4E13" box="[857,948,1532,1558]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567E0C2E524A8331FBA34E66" pageId="9" pageNumber="10" type="description">
<paragraph id="8BAF055A567E0C2E524A8331FB644E38" blockId="9.[374,1214,1531,1635]" box="[374,1214,1570,1597]" pageId="9" pageNumber="10">urn:lsid:zoobank.org:act:91C857A7-FBD3-4676-8F5C-5C7795232F57</paragraph>
<paragraph id="8BAF055A567E0C2E5287835BFBA34E66" blockId="9.[374,1214,1531,1635]" box="[443,1145,1608,1635]" pageId="9" pageNumber="10">
<figureCitation id="132B19DF567E0C2E5287835BFDCF4E66" box="[443,533,1608,1635]" captionStart="Figs43–51" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="9" pageNumber="10">Figs 45</figureCitation>
,
<figureCitation id="132B19DF567E0C2E511D835BFD9B4E67" box="[545,577,1608,1634]" captionStart="Figs 68–73" captionStartId="60.[189,241,1882,1908]" captionTargetBox="[190,1398,265,1816]" captionTargetId="figure@60.[189,1398,265,1816]" captionText="Figs 68–73. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 68. R. adjarica sp. nov. (holotype). 69. R. dunini sp. nov. (holotype). 70. R. hyrcanica Dunin, 1988 (holotype). 71. R. pontica (Spassky, 1937) (Lazarevskoe). 72. R. birecikensis sp. nov. (holotype). 73. Same species as preceding (paratype, Birecik). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163444/files/figure.png" pageId="9" pageNumber="10">69</figureCitation>
,
<figureCitation id="132B19DF567E0C2E5171835BFDB74E66" box="[589,621,1608,1635]" captionStart="Figs 84–89" captionStartId="63.[189,241,1892,1918]" captionTargetBox="[189,1398,266,1873]" captionTargetId="figure@63.[189,1398,265,1878]" captionText="Figs 84–89. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163450/files/figure.png" pageId="9" pageNumber="10">85</figureCitation>
,
<figureCitation id="132B19DF567E0C2E5145835AFD7D4E66" box="[633,679,1609,1635]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="9" pageNumber="10">103</figureCitation>
,
<figureCitation id="132B19DF567E0C2E518F835AFD3A4E66" box="[691,736,1609,1635]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="9" pageNumber="10">118</figureCitation>
,
<figureCitation id="132B19DF567E0C2E51D1835AFCC64E66" box="[749,796,1609,1635]" captionStart="Figs129–132" captionStartId="68.[189,241,1870,1896]" captionTargetBox="[220,1291,268,1789]" captionTargetId="figure@68.[189,1323,265,1791]" captionText="Figs129–132.Raveniolaspp.,♂♂.TibiaandmetatarsusI,retrolateralviews.—129.R.adjaricasp.nov. (holotype).130.R.duninisp.nov.(holotype).131.R.hyrcanicaDunin,1988(holotype).132.R.pontica (Spassky, 1937) (Lazarevskoe). Scalebars: 1.0 mm." httpUri="https://zenodo.org/record/1163460/files/figure.png" pageId="9" pageNumber="10">130</figureCitation>
,
<figureCitation id="132B19DF567E0C2E501B835BFC8C4E66" box="[807,854,1608,1635]" captionStart="Figs 149–152" captionStartId="73.[189,241,1930,1956]" captionTargetBox="[194,1398,266,1866]" captionTargetId="figure@73.[189,1398,265,1875]" captionText="Figs 149–152. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 149. R. adjarica sp. nov. (holotype). 150. R. dunini sp. nov. (holotype). 151. R. hyrcanica Dunin, 1988 (holotype). 152. R. pontica (Spassky, 1937) (Lazarevskoe). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163470/files/figure.png" pageId="9" pageNumber="10">150</figureCitation>
,
<figureCitation id="132B19DF567E0C2E505E835BFC164E66" box="[866,972,1608,1635]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="9" pageNumber="10">174–175</figureCitation>
,
<figureCitation id="132B19DF567E0C2E50EB835AFBDC4E66" box="[983,1030,1609,1635]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="9" pageNumber="10">199</figureCitation>
,
<figureCitation id="132B19DF567E0C2E572E835AFB9B4E66" box="[1042,1089,1609,1635]" captionStart="Figure 220" captionStartId="86.[189,272,1534,1560]" captionTargetBox="[190,1398,265,1471]" captionTargetId="figure@86.[189,1398,265,1471]" captionText="Figure 220. Localities of Raveniola spp. in the eastern part of the Caucasus: R. dunini sp. nov. (), R. hyrcanica Dunin, 1988 () and R. zaitzevi (Charitonov, 1948) ().Abbreviation:TU = Turkey." httpUri="https://zenodo.org/record/1163496/files/figure.png" pageId="9" pageNumber="10">220</figureCitation>
,
<figureCitation id="132B19DF567E0C2E5771835AFBA34E66" box="[1101,1145,1609,1635]" captionStart="Figure 222" captionStartId="87.[189,272,1855,1881]" captionTargetBox="[189,1398,1034,1819]" captionTargetId="figure@87.[189,1398,1033,1819]" captionText="Figure 222. Localities of Raveniola spp. in Iran: R. dunini sp. nov. (), R. marusiki sp. nov. (). R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (), R. niedermeyeri (Brignoli, 1972) () andR.vonwickiZonstein,2000().Abbreviations:AJ=Azerbaijan;AM =Armenia;UZ =Uzbekistan." httpUri="https://zenodo.org/record/1163500/files/figure.png" pageId="9" pageNumber="10">222</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567E0C2E5381839EFAAD4F48" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BAF055A567E0C2E5381839EFEEC4EA2" blockId="9.[189,310,1677,1703]" box="[189,310,1677,1703]" pageId="9" pageNumber="10">
<heading id="D0E7B236567E0C2E5381839EFEEC4EA2" bold="true" box="[189,310,1677,1703]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B964D948567E0C2E5381839EFEEC4EA2" bold="true" box="[189,310,1677,1703]" pageId="9" pageNumber="10">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567E0C2E538183ACFAAD4F48" blockId="9.[189,1399,1727,1870]" pageId="9" pageNumber="10">
Different from other members of the same group by having an embolus with its flagelliform apical part arising laterally from the basal part, carrying a short acute keel forming a paraembolic apophysis (
<figureCitation id="132B19DF567E0C2E53F9821EFEB14F22" box="[197,363,1804,1831]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="9" pageNumber="10">Figs 174–175</figureCitation>
; cf.
<figureCitation id="132B19DF567E0C2E52A1821EFD624F2D" box="[413,696,1805,1832]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="9" pageNumber="10">Figs 171–173, 176–179</figureCitation>
), and by possessing high-arched spermathecae with median and lateral receptacles, subequal in size and located close to each other (
<figureCitation id="132B19DF567E0C2E50C68220FB844F48" box="[1018,1118,1843,1869]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="9" pageNumber="10">Fig. 199</figureCitation>
; cf.
<figureCitation id="132B19DF567E0C2E57B18220FAB04F48" box="[1165,1386,1843,1869]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="9" pageNumber="10">Figs 198, 200–203</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567E0C2E5381826BFB624FEE" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BAF055A567E0C2E5381826BFE9E4F97" blockId="9.[189,324,1912,1938]" box="[189,324,1912,1938]" pageId="9" pageNumber="10">
<heading id="D0E7B236567E0C2E5381826BFE9E4F97" bold="true" box="[189,324,1912,1938]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B964D948567E0C2E5381826BFE9E4F97" bold="true" box="[189,324,1912,1938]" pageId="9" pageNumber="10">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567E0C2E538182B9FB624FEE" blockId="9.[189,1399,1962,2027]" pageId="9" pageNumber="10">The specific epithet is given in honour of Dr. Pyotr Dunin (1952–1998), a prominent Soviet and Russian arachnologist, for his immense contribution to the study of the Caucasian spider fauna.</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567D0C2D53818418FDB54B4D" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BAF055A567D0C2D53818418FE704920" blockId="10.[189,426,267,293]" box="[189,426,267,293]" pageId="10" pageNumber="11">
<heading id="D0E7B236567D0C2D53818418FE704920" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B964D948567D0C2D53818418FE704920" bold="true" box="[189,426,267,293]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B780F07567D0C2D53D68428FAAD49A5" ID-GBIF-Occurrence="1806244362" collectingDate="1987-09-13" collectionCode="TAU" collectorName="S. L. Zonstein" country="Azerbaijan" elevation="650" latitude="39.643333" location="foothills of Karabakh Mts, hills alongside right (south) bank of Kendelanchai River, 1 km WSW of Kadzhar (Qacar" longLatPrecision="116" longitude="47.036667" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8BAF055A567D0C2D53D68428FE864950" blockId="10.[189,1399,315,416]" box="[234,348,315,341]" pageId="10" pageNumber="11">
<typeStatus id="54ABBBF8567D0C2D53D68428FE864950" box="[234,348,315,341]" pageId="10" pageNumber="11" type="holotype">
<emphasis id="B964D948567D0C2D53D68428FE864950" bold="true" box="[234,348,315,341]" pageId="10" pageNumber="11">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BAF055A567D0C2D53818472FAAD49A5" blockId="10.[189,1399,315,416]" pageId="10" pageNumber="11">
<collectingCountry id="F30745CA567D0C2D53818472FEAD497E" box="[189,375,353,379]" name="Azerbaijan" pageId="10" pageNumber="11">AZERBAIJAN</collectingCountry>
:
<specimenCount id="9D16CED3567D0C2D52BD8472FE43497E" box="[385,409,353,379]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
,
<location id="8ECF5381567D0C2D52998473FDF249A5" LSID="urn:lsid:plazi:treatment:03B9B44C567E0C2B51BC80E8FE1B497F:8ECF5381567D0C2D52998473FDF249A5" country="Azerbaijan" latitude="39.643333" longLatPrecision="116" longitude="47.036667" name="foothills of Karabakh Mts, hills alongside right (south) bank of Kendelanchai River, 1 km WSW of Kadzhar (Qacar" pageId="10" pageNumber="11">
foothills of Karabakh Mts, hills alongside right (south) bank of Kendelanchai River,
<quantity id="4CE8A8BF567D0C2D53818495FF2D499A" box="[189,247,389,416]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="km" value="1.0">1 km</quantity>
WSW of Kadzhar (Qacar
</location>
),
<geoCoordinate id="EE24639D567D0C2D51068497FD6649A5" box="[570,700,388,416]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="39.643333">39°38.6′ N</geoCoordinate>
,
<geoCoordinate id="EE24639D567D0C2D51FA8497FC9E49A5" box="[710,836,388,416]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="47.036667">47°02.2′ E</geoCoordinate>
,
<elevation id="003DE269567D0C2D50738496FC4149A5" box="[847,923,389,416]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="11" unit="m" value="650.0">
<quantity id="4CE8A8BF567D0C2D50738496FC4149A5" box="[847,923,389,416]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="11" unit="m" value="650.0">650 m</quantity>
</elevation>
,
<collectingDate id="EFEADA72567D0C2D509A8495FB9A49A5" box="[934,1088,390,416]" pageId="10" pageNumber="11" value="1987-09-13">
<date id="FFAE239A567D0C2D509A8495FB9A49A5" box="[934,1088,390,416]" pageId="10" pageNumber="11" value="1987-09-13">13 Sep. 1987</date>
</collectingDate>
,
<collectorName id="26E5608C567D0C2D57708495FB30499A" box="[1100,1258,389,416]" pageId="10" pageNumber="11">S.L. Zonstein</collectorName>
leg. (
<collectionCode id="ED019D9F567D0C2D56168495FAB149A5" box="[1322,1387,390,416]" collectionName="Israel, Tel Aviv, Tel Aviv University" pageId="10" pageNumber="11">TAU</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BAF055A567D0C2D53D684DAFDDA49E6" blockId="10.[189,1398,457,557]" box="[234,512,457,484]" pageId="10" pageNumber="11">
<emphasis id="B964D948567D0C2D53D684DAFEBD49E6" bold="true" box="[234,359,457,483]" pageId="10" pageNumber="11">Paratypes</emphasis>
(1 ♂, 5 ♀♀)
</paragraph>
<paragraph id="8BAF055A567D0C2D538184FDFC804A28" blockId="10.[189,1398,457,557]" pageId="10" pageNumber="11">
<materialsCitation id="3B780F07567D0C2D538184FDFBDD4A0D" ID-GBIF-Occurrence="1806244430" box="[189,1031,494,520]" pageId="10" pageNumber="11">AZERBAIJAN: 1 ♂, 1 ♀, same collection data as for holotype (TAU)</materialsCitation>
;
<materialsCitation id="3B780F07567D0C2D572984FDFC804A28" ID-GBIF-Occurrence="1806244419" pageId="10" pageNumber="11">4 ♀♀, from same locality but collected 11 Aug. 1986, P.M. Dunin leg. (ZMMU, TAU).</materialsCitation>
</paragraph>
<paragraph id="8BAF055A567D0C2D53D68745FD4E4A74" blockId="10.[189,1399,598,736]" box="[234,660,598,625]" pageId="10" pageNumber="11">
<emphasis id="B964D948567D0C2D53D68745FE3B4A75" bold="true" box="[234,481,598,624]" pageId="10" pageNumber="11">Additional material</emphasis>
(2 ♀♀, 2 juvs)
</paragraph>
<paragraph id="8BAF055A567D0C2D53818768FB8E4AE5" blockId="10.[189,1399,598,736]" pageId="10" pageNumber="11">
<materialsCitation id="3B780F07567D0C2D53818768FAE34A90" ID-GBIF-Occurrence="1806244426" box="[189,1337,633,662]" pageId="10" pageNumber="11">ARMENIA: 1 ♀, Kafan, 39°12′ N, 46°24′ E, 800–900 m, 18 May 1988, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F07567D0C2D567E8768FAF24ABF" ID-GBIF-Occurrence="1806244432" pageId="10" pageNumber="11">1 ♀, Kafan Distr., Shikahoh, 39°06′ N, 46°28′ E, 1000–1200 m, 19 May 1988, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F07567D0C2D561387B3FB8E4AE5" ID-GBIF-Occurrence="1806244431" pageId="10" pageNumber="11">1 juv., Megri, 38°54′ N, 46°14′ E, 700–800 m, 20 May 1988, P.M. Dunin leg. (TAU).</materialsCitation>
</paragraph>
<paragraph id="8BAF055A567D0C2D5381861AFDB54B4D" blockId="10.[189,1399,776,840]" pageId="10" pageNumber="11">IRAN: 1 juv., East Azerbaijan Province,Arasbaran Reserve, Makīdī Valley, 1650–1800 m, 4–5 Jun. 1978, J. Martens and H. Pieper leg. (SMF).</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567D0C2C53818662FCC34F4E" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="8BAF055A567D0C2D53818662FE944B8E" blockId="10.[189,334,881,907]" box="[189,334,881,907]" pageId="10" pageNumber="11">
<heading id="D0E7B236567D0C2D53818662FE944B8E" bold="true" box="[189,334,881,907]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B964D948567D0C2D53818662FE944B8E" bold="true" box="[189,334,881,907]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567D0C2D53D686B1FE714BB9" blockId="10.[189,439,929,993]" box="[234,427,929,956]" pageId="10" pageNumber="11">
<emphasis id="B964D948567D0C2D53D686B1FEF04BB9" bold="true" box="[234,298,930,956]" pageId="10" pageNumber="11">Male</emphasis>
(holotype)
</paragraph>
<paragraph id="8BAF055A567D0C2D538186D4FE6D4BE4" blockId="10.[189,439,929,993]" box="[189,439,966,993]" pageId="10" pageNumber="11">
HABITUS. See
<figureCitation id="132B19DF567D0C2D526186D5FE6D4BE4" box="[349,439,966,993]" captionStart="Figs43–51" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="10" pageNumber="11">Fig. 45.</figureCitation>
</paragraph>
<paragraph id="8BAF055A567D0C2D53818119FB7D4C21" blockId="10.[189,1191,1034,1061]" box="[189,1191,1034,1061]" pageId="10" pageNumber="11">MEASUREMENTS. TBL 13.20, CL 5.38, CW 4.67, LL 0.42, LW 1.05, SL 2.68, SW 2.43.</paragraph>
<paragraph id="8BAF055A567D0C2D5381815EFBC64CF9" blockId="10.[189,1399,1101,1276]" pageId="10" pageNumber="11">COLOUR. Carapace, palps and legs dorsally light reddish brown; eye tubercle with darker spots surrounding AMEs and lateral eyes; chelicerae cherry-red; sternum, labium, maxillae, palps and legs ventrally light brown; abdomen dorsally light brown, with darker brown dorsal pattern represented by weak and narrow median lanceolate spot and four pairs of inclined chevron-like fasciae located posteriorly, ventral abdominal surface and spinnerets pale yellowish grey.</paragraph>
<paragraph id="8BAF055A567D0C2D53818036FAB64DAB" blockId="10.[189,1399,1317,1455]" pageId="10" pageNumber="11">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF567D0C2D503E8036FC864D3A" box="[770,860,1317,1343]" captionStart="Figs 68–73" captionStartId="60.[189,241,1882,1908]" captionTargetBox="[190,1398,265,1816]" captionTargetId="figure@60.[189,1398,265,1816]" captionText="Figs 68–73. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 68. R. adjarica sp. nov. (holotype). 69. R. dunini sp. nov. (holotype). 70. R. hyrcanica Dunin, 1988 (holotype). 71. R. pontica (Spassky, 1937) (Lazarevskoe). 72. R. birecikensis sp. nov. (holotype). 73. Same species as preceding (paratype, Birecik). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163444/files/figure.png" pageId="10" pageNumber="11">Fig. 69</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF567D0C2D57EE8036FAE04D3A" box="[1234,1338,1317,1343]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="10" pageNumber="11">Fig. 103</figureCitation>
. Eye diameters and interdistances: AME 0.14(0.17), ALE 0.18, PLE 0.13, PME: 0.10, AME–AME 0.12(0.09), ALE–AME 0.07(0.06), ALE–PLE 0.06, PLE–PME 0.03, PME–PME 0.38. Each cheliceral furrow with 9–11 promarginal teeth and 6–7 mesobasal denticles. Maxillae with 5–6 relatively long cuspules each.
</paragraph>
<paragraph id="8BAF055A567D0C2D538180CBFE294E65" blockId="10.[189,1398,1495,1633]" pageId="10" pageNumber="11">
LEGS. Tibia and metatarsus I as shown in
<figureCitation id="132B19DF567D0C2D519280C4FCCE4DF4" box="[686,788,1495,1521]" captionStart="Figs129–132" captionStartId="68.[189,241,1870,1896]" captionTargetBox="[220,1291,268,1789]" captionTargetId="figure@68.[189,1323,265,1791]" captionText="Figs129–132.Raveniolaspp.,♂♂.TibiaandmetatarsusI,retrolateralviews.—129.R.adjaricasp.nov. (holotype).130.R.duninisp.nov.(holotype).131.R.hyrcanicaDunin,1988(holotype).132.R.pontica (Spassky, 1937) (Lazarevskoe). Scalebars: 1.0 mm." httpUri="https://zenodo.org/record/1163460/files/figure.png" pageId="10" pageNumber="11">Fig. 130</figureCitation>
. Scopula: distal on metatarsi I–II, entire on tarsi I– II; few scopuliform hairs mixed with bristles on tarsi III–IV. Trichobothria: 2 rows of 9–11 each on tibiae, 11–13 on metatarsi, 11–14 on tarsi, 9 on cymbium. Paired claws: inner margin with 6–8 teeth; 7–8 teeth on outer margin.
</paragraph>
<paragraph id="8BAF055A567D0C2D53818399FB964F58" blockId="10.[189,1399,1674,1886]" pageId="10" pageNumber="11">SPINATION. Palp: femur d1–1–1(0), pd1, rd1; tibia d1–1, p1–1–1(0), pv1–1–1, r1, v1–1; cymbium d4(5). Leg I: femur d1–1–1(0)–1(0), pd1–1; tibia p1–1, v2(3)–2(1)–2(1). Leg II: femur d1–1–1(0)–1(0), pd1– 1; tibia p1–1–1, v1–2–1–3; metatarsus p1, v1–2–2. Leg III: femur d1–1–1(0)–1(0), pd1–1–1, rd1–1; patella p1, r1; tibia d1, p1–1–1, r1–1–1, v2–2–2; metatarsus pd1, rd1–1, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur d1–1–1–1(0)–1(0), pd1, rd1; patella p1(0), r1; tibia d1, p1–1, r1–1–1, v2–2–2; metatarsus pd1–1, rd1, p1–1–1, r1–1, v2–1(2)–3. Metatarsus I and patellae I–II aspinose.</paragraph>
<paragraph id="8BAF055A567D0C2D53818294FF204FEE" blockId="10.[189,1399,1926,2027]" pageId="10" pageNumber="11">
PALP. Tibia, cymbium and palpal organ as shown in
<figureCitation id="132B19DF567D0C2D500D8295FC424FA4" box="[817,920,1926,1953]" captionStart="Figs 149–152" captionStartId="73.[189,241,1930,1956]" captionTargetBox="[194,1398,266,1866]" captionTargetId="figure@73.[189,1398,265,1875]" captionText="Figs 149–152. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 149. R. adjarica sp. nov. (holotype). 150. R. dunini sp. nov. (holotype). 151. R. hyrcanica Dunin, 1988 (holotype). 152. R. pontica (Spassky, 1937) (Lazarevskoe). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163470/files/figure.png" pageId="10" pageNumber="11">Fig. 150</figureCitation>
. Palpal organ provided with moderately short sideward-arising lamellose embolus with keel forming short paraembolic apophysis (
<figureCitation id="132B19DF567D0C2D57C282BFFF374FEE" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="10" pageNumber="11">Figs 174– 175</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A567C0C2C53818418FB794949" blockId="11.[189,1398,267,333]" pageId="11" pageNumber="12">SPINNERETS. PMS: length 0.47; diameter 0.20. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.93, 0.70, 0.82; total length 2.45; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A567C0C2C5381846AFE354997" blockId="11.[189,495,376,403]" box="[189,495,376,403]" pageId="11" pageNumber="12">LEG MEASUREMENTS. ♂(♀).</paragraph>
<paragraph id="8BAF055A567C0C2C525484A5FAB34AA5" pageId="11" pageNumber="12">
<table id="F910F7FA567CF3D853F484A5FAB04AA4" box="[200,1386,438,673]" colsContinueIn="16.[200,1385,731,966]" gridcols="7" gridrows="6" pageId="11" pageNumber="12" rowsContinueIn="13.[200,1385,1322,1557]">
<tr id="35200718567CF3D853F484A5FAB049D5" box="[200,1386,438,464]" gridrow="0" pageId="11" pageNumber="12" rowspan-0="1">
<th id="76F16E64567CF3D8527284A5FE1549D5" box="[334,463,438,464]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">Femur</th>
<th id="76F16E64567CF3D8513F84A5FD5F49D5" box="[515,645,438,464]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">Patella</th>
<th id="76F16E64567CF3D8518584A5FCE149D5" box="[697,827,438,464]" gridcol="3" gridrow="0" pageId="11" pageNumber="12">Tibia</th>
<th id="76F16E64567CF3D8505284A5FC2A49D5" box="[878,1008,438,464]" gridcol="4" gridrow="0" pageId="11" pageNumber="12">Metatarsus</th>
<th id="76F16E64567CF3D8571884A5FB7C49D5" box="[1060,1190,438,464]" gridcol="5" gridrow="0" pageId="11" pageNumber="12">Tarsus</th>
<th id="76F16E64567CF3D857F684A5FAB049D5" box="[1226,1386,438,464]" gridcol="6" gridrow="0" pageId="11" pageNumber="12">Total</th>
</tr>
<tr id="35200718567CF3D853F484CCFAB049FF" box="[200,1386,479,506]" gridrow="1" pageId="11" pageNumber="12">
<th id="76F16E64567CF3D853F484CCFEC649FF" box="[200,284,479,506]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">Palp</th>
<td id="76F16E64567CF3D8527284CCFE1549FF" box="[334,463,479,506]" gridcol="1" gridrow="1" pageId="11" pageNumber="12">2.83 (2.78)</td>
<td id="76F16E64567CF3D8513F84CCFD5F49FF" box="[515,645,479,506]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">1.63 (1.73)</td>
<td id="76F16E64567CF3D8518584CCFCE149FF" box="[697,827,479,506]" gridcol="3" gridrow="1" pageId="11" pageNumber="12">2.17 (1.88)</td>
<td id="76F16E64567CF3D8505284CCFC2A49FF" box="[878,1008,479,506]" gridcol="4" gridrow="1" pageId="11" pageNumber="12">–</td>
<td id="76F16E64567CF3D8571884CCFB7C49FF" box="[1060,1190,479,506]" gridcol="5" gridrow="1" pageId="11" pageNumber="12">1.03 (1.70)</td>
<td id="76F16E64567CF3D857F684CCFAB049FF" box="[1226,1386,479,506]" gridcol="6" gridrow="1" pageId="11" pageNumber="12">7.66 (8.09)</td>
</tr>
<tr id="35200718567CF3D853F4871AFAB04A21" box="[200,1386,521,548]" gridrow="2" pageId="11" pageNumber="12">
<th id="76F16E64567CF3D853F4871AFEC64A21" box="[200,284,521,548]" gridcol="0" gridrow="2" pageId="11" pageNumber="12">Leg I</th>
<td id="76F16E64567CF3D85272871AFE154A21" box="[334,463,521,548]" gridcol="1" gridrow="2" pageId="11" pageNumber="12">4.37 (3.65)</td>
<td id="76F16E64567CF3D8513F871AFD5F4A21" box="[515,645,521,548]" gridcol="2" gridrow="2" pageId="11" pageNumber="12">2.65 (2.40)</td>
<td id="76F16E64567CF3D85185871AFCE14A21" box="[697,827,521,548]" gridcol="3" gridrow="2" pageId="11" pageNumber="12">3.53 (2.77)</td>
<td id="76F16E64567CF3D85052871AFC2A4A21" box="[878,1008,521,548]" gridcol="4" gridrow="2" pageId="11" pageNumber="12">3.55 (2.10)</td>
<td id="76F16E64567CF3D85718871AFB7C4A21" box="[1060,1190,521,548]" gridcol="5" gridrow="2" pageId="11" pageNumber="12">2.03 (1.58)</td>
<td id="76F16E64567CF3D857F6871AFAB04A21" box="[1226,1386,521,548]" gridcol="6" gridrow="2" pageId="11" pageNumber="12">16.13 (12.50)</td>
</tr>
<tr id="35200718567CF3D853F48720FAB04A4B" box="[200,1386,563,590]" gridrow="3" pageId="11" pageNumber="12">
<th id="76F16E64567CF3D853F48720FEC64A4B" box="[200,284,563,590]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">Leg II</th>
<td id="76F16E64567CF3D852728720FE154A4B" box="[334,463,563,590]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">3.97 (3.32)</td>
<td id="76F16E64567CF3D8513F8720FD5F4A4B" box="[515,645,563,590]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">2.33 (2.13)</td>
<td id="76F16E64567CF3D851858720FCE14A4B" box="[697,827,563,590]" gridcol="3" gridrow="3" pageId="11" pageNumber="12">2.97 (2.30)</td>
<td id="76F16E64567CF3D850528720FC2A4A4B" box="[878,1008,563,590]" gridcol="4" gridrow="3" pageId="11" pageNumber="12">3.13 (2.02)</td>
<td id="76F16E64567CF3D857188720FB7C4A4B" box="[1060,1190,563,590]" gridcol="5" gridrow="3" pageId="11" pageNumber="12">1.93 (1.55)</td>
<td id="76F16E64567CF3D857F68720FAB04A4B" box="[1226,1386,563,590]" gridcol="6" gridrow="3" pageId="11" pageNumber="12">14.33 (11.32)</td>
</tr>
<tr id="35200718567CF3D853F4874FFAB04A72" box="[200,1386,604,631]" gridrow="4" pageId="11" pageNumber="12">
<th id="76F16E64567CF3D853F4874FFEC64A72" box="[200,284,604,631]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">Leg III</th>
<td id="76F16E64567CF3D85272874FFE154A72" box="[334,463,604,631]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">3.20 (2.93)</td>
<td id="76F16E64567CF3D8513F874FFD5F4A72" box="[515,645,604,631]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">1.77 (1.83)</td>
<td id="76F16E64567CF3D85185874FFCE14A72" box="[697,827,604,631]" gridcol="3" gridrow="4" pageId="11" pageNumber="12">2.47 (1.85)</td>
<td id="76F16E64567CF3D85052874FFC2A4A72" box="[878,1008,604,631]" gridcol="4" gridrow="4" pageId="11" pageNumber="12">3.25 (2.75)</td>
<td id="76F16E64567CF3D85718874FFB7C4A72" box="[1060,1190,604,631]" gridcol="5" gridrow="4" pageId="11" pageNumber="12">1.93 (1.62)</td>
<td id="76F16E64567CF3D857F6874FFAB04A72" box="[1226,1386,604,631]" gridcol="6" gridrow="4" pageId="11" pageNumber="12">12.62 (10.61)</td>
</tr>
<tr id="35200718567CF3D853F48795FAB04AA4" box="[200,1386,646,673]" gridrow="5" pageId="11" pageNumber="12">
<th id="76F16E64567CF3D853F48795FEC64AA4" box="[200,284,646,673]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">Leg IV</th>
<td id="76F16E64567CF3D852728795FE154AA4" box="[334,463,646,673]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">4.35 (3.80)</td>
<td id="76F16E64567CF3D8513F8795FD5F4AA4" box="[515,645,646,673]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">2.17 (2.25)</td>
<td id="76F16E64567CF3D851858795FCE14AA4" box="[697,827,646,673]" gridcol="3" gridrow="5" pageId="11" pageNumber="12">3.45 (2.82)</td>
<td id="76F16E64567CF3D850528795FC2A4AA4" box="[878,1008,646,673]" gridcol="4" gridrow="5" pageId="11" pageNumber="12">4.37 (3.55)</td>
<td id="76F16E64567CF3D857188795FB7C4AA4" box="[1060,1190,646,673]" gridcol="5" gridrow="5" pageId="11" pageNumber="12">2.27 (1.87)</td>
<td id="76F16E64567CF3D857F68795FAB04AA4" box="[1226,1386,646,673]" gridcol="6" gridrow="5" pageId="11" pageNumber="12">16.61 (14.29)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A567C0C2C53D687DEFE1F4AE2" blockId="11.[189,1191,717,783]" box="[234,453,717,743]" pageId="11" pageNumber="12">
<emphasis id="B964D948567C0C2C53D687DEFE9F4AE2" bold="true" box="[234,325,717,743]" pageId="11" pageNumber="12">Female</emphasis>
(paratype)
</paragraph>
<paragraph id="8BAF055A567C0C2C538187E7FB7D4B0B" blockId="11.[189,1191,717,783]" box="[189,1191,756,783]" pageId="11" pageNumber="12">MEASUREMENTS. TBL 13.65, CL 4.67, CW 4.50, LL 0.37, LW 0.97, SL 2.65, SW 2.33.</paragraph>
<paragraph id="8BAF055A567C0C2C5381862AFE714B56" blockId="11.[189,427,825,852]" box="[189,427,825,852]" pageId="11" pageNumber="12">COLOUR. As in male.</paragraph>
<paragraph id="8BAF055A567C0C2C5381866CFA8A4C0B" blockId="11.[189,1399,894,1039]" pageId="11" pageNumber="12">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF567C0C2C503E866CFC864B9C" box="[770,860,894,921]" captionStart="Figs 84–89" captionStartId="63.[189,241,1892,1918]" captionTargetBox="[189,1398,266,1873]" captionTargetId="figure@63.[189,1398,265,1878]" captionText="Figs 84–89. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163450/files/figure.png" pageId="11" pageNumber="12">Fig. 85</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF567C0C2C57EF866CFAE04B9C" box="[1235,1338,895,921]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="11" pageNumber="12">Fig. 118</figureCitation>
. Eye diameters and interdistances: AME 0.09(0.11), ALE 0.21, PLE 0.12, PME 0.10, AME–AME 0.18(0.16), ALE–AME 0.09(0.08), ALE–PLE 0.06, PLE–PME 0.04, PME–PME 0.39. Each cheliceral furrow with 10 promarginal teeth and 6–7 mesobasal denticles. Maxillae each with 5–7 relatively long cuspules.
</paragraph>
<paragraph id="8BAF055A567C0C2C53818129FE5A4CCC" blockId="11.[189,1398,1082,1226]" pageId="11" pageNumber="12">LEGS. Scopula: distal on metatarsi I–II, entire on tarsi I–II and palpal tarsus, widely divided on tarsus III, elsewhere absent. Trichobothria: 2 rows of 8–9 each on tibiae, 10–12 on metatarsi, 11–13 on tarsi, 9 on palpal tarsus. Paired tarsal claws: outer and inner margins with 5–6 teeth, palpal claw with 4 teeth on inner margin.</paragraph>
<paragraph id="8BAF055A567C0C2C538181E6FE464DFF" blockId="11.[189,1399,1269,1530]" pageId="11" pageNumber="12">SPINATION. Femora I–IV with one basodorsal spine and 3 dorsal bristles alongside midline; femur III also with two prodorsal and retrodorsal rows of 2–3 bristles each; femur IV without prodorsal bristle row; palpal patella and patellae I–II aspinose. Palp: femur pd1; tibia v2–2–3; tarsus d8(9). Leg I: femur d1, pd1; tibia v1–1–1(0)–2(3); metatarsus v2–2–2. Leg II: femur d1, pd1; tibia p1–1, v1–1–2(3); metatarsus v2–2–2. Leg III: femur d1, rd1; patella p1, r1; tibia d1, p1–1, r1–1–1, v2–2–3; metatarsus pd1–1, p1–1, r1–1, v2–2–3. Leg IV: femur d1, rd1; patella r1; tibia d1, p1–1, r1–1, v2–2–2; metatarsus d1, p1–1–1–1, r1–1–1–1, v2–2–2.</paragraph>
<paragraph id="8BAF055A567C0C2C53818335FB254E45" blockId="11.[189,1279,1574,1600]" box="[189,1279,1574,1600]" pageId="11" pageNumber="12">
SPERMATHECAE. Broad, large and symmetrical, with receptacles close to each other (
<figureCitation id="132B19DF567C0C2C57B78335FB284E45" box="[1163,1266,1574,1600]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="11" pageNumber="12">Fig. 199</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A567C0C2C53818378FB794EA9" blockId="11.[189,1398,1643,1709]" pageId="11" pageNumber="12">SPINNERETS. PMS: length 0.50; diameter 0.22. PLS: maximal diameter 0.52; length of basal, medial and apical segments 0.90, 0.64, 0.73; total length 2.27; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A567C0C2C538183CBFEE94EF7" blockId="11.[189,307,1752,1778]" box="[189,307,1752,1778]" pageId="11" pageNumber="12">
<heading id="D0E7B236567C0C2C538183CBFEE94EF7" bold="true" box="[189,307,1752,1778]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B964D948567C0C2C538183CBFEE94EF7" bold="true" box="[189,307,1752,1778]" pageId="11" pageNumber="12">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567C0C2C53818219FCC34F4E" blockId="11.[189,1399,1802,1868]" pageId="11" pageNumber="12">Carapace length in males varies from 5.15 to 5.38, in females from 4.55 to 5.20. The copulative organs of both sexes do not show any significant variation.</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567C0C2C53818264FDE54FEE" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="8BAF055A567C0C2C53818264FEFA4F94" blockId="11.[189,288,1911,1937]" box="[189,288,1911,1937]" pageId="11" pageNumber="12">
<heading id="D0E7B236567C0C2C53818264FEFA4F94" bold="true" box="[189,288,1911,1937]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B964D948567C0C2C53818264FEFA4F94" bold="true" box="[189,288,1911,1937]" pageId="11" pageNumber="12">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567C0C2C538182BAFDE54FEE" blockId="11.[189,1398,1961,2027]" pageId="11" pageNumber="12">
The type series was collected in a low open oak forest (
<taxonomicName id="4C107ED9567C0C2C505782B9FC174FC1" box="[875,973,1962,1988]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="B964D948567C0C2C505782B9FC174FC1" box="[875,973,1962,1988]" italics="true" pageId="11" pageNumber="12">Quercus</emphasis>
</taxonomicName>
sp. mixed with
<taxonomicName id="4C107ED9567C0C2C57AA82B9FACA4FC1" box="[1174,1296,1962,1988]" class="Magnoliopsida" family="Rosaceae" genus="Crataegus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="B964D948567C0C2C57AA82B9FACA4FC1" box="[1174,1296,1962,1988]" italics="true" pageId="11" pageNumber="12">Crataegus</emphasis>
</taxonomicName>
sp.); all spiders were found under stones.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567B0C2B53818418FE1B497F" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BAF055A567B0C2B53818418FE8C4920" blockId="12.[189,342,267,293]" box="[189,342,267,293]" pageId="12" pageNumber="13">
<heading id="D0E7B236567B0C2B53818418FE8C4920" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B964D948567B0C2B53818418FE8C4920" bold="true" box="[189,342,267,293]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567B0C2B5381842FFE1B497F" blockId="12.[189,1399,315,378]" pageId="12" pageNumber="13">
Eastern part of the Caucasus Minor (eastern Armenia, south-western Azerbaijan, far north-western Iran). See
<figureCitation id="132B19DF567B0C2B53D38473FE8F497F" box="[239,341,352,378]" captionStart="Figure 220" captionStartId="86.[189,272,1534,1560]" captionTargetBox="[190,1398,265,1471]" captionTargetId="figure@86.[189,1398,265,1471]" captionText="Figure 220. Localities of Raveniola spp. in the eastern part of the Caucasus: R. dunini sp. nov. (), R. hyrcanica Dunin, 1988 () and R. zaitzevi (Charitonov, 1948) ().Abbreviation:TU = Turkey." httpUri="https://zenodo.org/record/1163496/files/figure.png" pageId="12" pageNumber="13">Figs 220</figureCitation>
and
<figureCitation id="132B19DF567B0C2B52B28473FE64497F" box="[398,446,352,378]" captionStart="Figure 222" captionStartId="87.[189,272,1855,1881]" captionTargetBox="[189,1398,1034,1819]" captionTargetId="figure@87.[189,1398,1033,1819]" captionText="Figure 222. Localities of Raveniola spp. in Iran: R. dunini sp. nov. (), R. marusiki sp. nov. (). R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (), R. niedermeyeri (Brignoli, 1972) () andR.vonwickiZonstein,2000().Abbreviations:AJ=Azerbaijan;AM =Armenia;UZ =Uzbekistan." httpUri="https://zenodo.org/record/1163500/files/figure.png" pageId="12" pageNumber="13">222</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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