243 lines
41 KiB
XML
243 lines
41 KiB
XML
<document id="786317FBCC420204AA24AD9375D93231" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. & Yağmur, Ersen A." docDate="2018" docId="03B9B44C56680C3853818418FAAC4F21" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola niedermeyeri" docType="key" docVersion="12" lastPageNumber="32" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="32" updateTime="1698518020641" updateUser="ExternalLinkService">
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<mods:title id="5D42DC41C4DE48F257E3B46C40038803">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="1A4273B6C524CFA18BFCCF00D081392A">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="F87B167DC79751B977519A3F65FE6FD0">Kunt, Kadir B.</mods:namePart>
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<mods:namePart id="77D148E05670FC844712DE7D8FBFC0E1">Yağmur, Ersen A.</mods:namePart>
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<mods:title id="ECA4A8577279943CECEA0066BBCEF4AE">European Journal of Taxonomy</mods:title>
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<mods:part id="DF545D86CD86C3AF76A34E131B67A143">
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<mods:date id="21B48DB44D52C7691D3256F6138A9F77">2018</mods:date>
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<mods:number id="81775CE7E5C3C936514AE686BB4E3D3E">2018-01-30</mods:number>
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<mods:number id="B5DCEF692EC5DA57F010BFB3C845661D">399</mods:number>
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<treatment id="03B9B44C56680C3853818418FAAC4F21" ID-DOI="http://doi.org/10.5281/zenodo.5980302" ID-GBIF-Taxon="140561343" ID-Zenodo-Dep="5980302" LSID="urn:lsid:plazi:treatment:03B9B44C56680C3853818418FAAC4F21" httpUri="http://treatment.plazi.org/id/03B9B44C56680C3853818418FAAC4F21" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="C30A56D156680C3853818418FD83492D" box="[189,601,267,296]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BAF055A56680C3853818418FD83492D" blockId="31.[189,601,267,296]" box="[189,601,267,296]" pageId="31" pageNumber="32">
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<emphasis id="B964D94856680C3853818418FD83492D" bold="true" box="[189,601,267,296]" pageId="31" pageNumber="32">
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<taxonomicName id="4C107ED956680C3853818418FDD9492D" ID-CoL="6WT99" baseAuthorityName="Brignoli" baseAuthorityYear="1972" box="[189,515,267,296]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="niedermeyeri">Raveniola niedermeyeri</taxonomicName>
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<emphasis id="B964D94856680C385137841FFD83492D" bold="true" box="[523,601,268,296]" italics="true" pageId="31" pageNumber="32">group</emphasis>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A56D156680C3853818451FC374A07" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="8BAF055A56680C3853818451FE164958" blockId="31.[189,460,322,349]" box="[189,460,322,349]" pageId="31" pageNumber="32">
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<heading id="D0E7B23656680C3853818451FE164958" bold="true" box="[189,460,322,349]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis id="B964D94856680C3853818451FE164958" bold="true" box="[189,460,322,349]" pageId="31" pageNumber="32">Diagnostic characters</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAF055A56680C3853818467FC374A07" blockId="31.[189,1399,372,515]" pageId="31" pageNumber="32">
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The subapical part of the male palpal tibia dilated retrolaterally and provided with a bare flattened area carrying a ventral sensilla (
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<figureCitation id="132B19DF56680C3851398488FD7549B0" box="[517,687,411,437]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 189–197</figureCitation>
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). Embolus relatively long, curved or gently twisted, with a rudimentary subapical keel (
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<figureCitation id="132B19DF56680C38512984D2FD6549D9" box="[533,703,449,476]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 191–194</figureCitation>
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), or vestiges of the embolic keel are lost (
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<figureCitation id="132B19DF56680C3857F584D2FEF24A06" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 189–190, 195–197</figureCitation>
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). Spermathecae relatively narrow, F-shaped (
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<figureCitation id="132B19DF56680C38500584FBFC3A4A07" box="[825,992,488,514]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="31" pageNumber="32">Figs 208–212</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A56D156680C385381873EFBB04AA5" pageId="31" pageNumber="32" type="synonymic_list">
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<paragraph id="8BAF055A56680C385381873EFE564A42" blockId="31.[189,396,557,583]" box="[189,396,557,583]" pageId="31" pageNumber="32">
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<heading id="D0E7B23656680C385381873EFE564A42" bold="true" box="[189,396,557,583]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis id="B964D94856680C385381873EFE564A42" bold="true" box="[189,396,557,583]" pageId="31" pageNumber="32">Species included</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAF055A56680C385381874CFBB04AA5" blockId="31.[189,1399,607,672]" pageId="31" pageNumber="32">
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<taxonomicName id="4C107ED956680C385381874CFE794A7C" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[189,419,607,633]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
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<emphasis id="B964D94856680C385381874CFE794A7C" box="[189,419,607,633]" italics="true" pageId="31" pageNumber="32">Raveniola marusiki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A257643356680C385296874CFDD24A7C" box="[426,520,607,633]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C107ED956680C3851288773FB754A7C" authority="Marusik, Zamani & Mirshamsi, 2014" authorityName="Marusik, Zamani & Mirshamsi" authorityYear="2014" box="[532,1199,607,633]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mazandaranica">
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<emphasis id="B964D94856680C3851288773FD364A7C" box="[532,748,607,633]" italics="true" pageId="31" pageNumber="32">R. mazandaranica</emphasis>
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<bibRefCitation id="EF8178AB56680C3851CF874CFB754A7C" author="Marusik Y. M. & Zamani A. & Mirshamsi O." box="[755,1199,607,633]" journalOrPublisher="ZooKeys" pageId="31" pageNumber="32" pagination="1 - 10" part="463" publicationUrl="https://doi.org/10.3897/zookeys.463.8692" refString="Marusik Y. M., Zamani A. & Mirshamsi O. 2014. Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae). ZooKeys 463: 1 - 10. https: // doi. org / 10.3897 / zookeys. 463.8692" title="Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae)" type="journal article" year="2014">Marusik, Zamani & Mirshamsi, 2014</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C107ED956680C3857878773FEA54AA5" authority="(Brignoli, 1972)" baseAuthorityName="Brignoli" baseAuthorityYear="1972" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="niedermeyeri">
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<emphasis id="B964D94856680C3857878773FAAD4A7C" box="[1211,1399,607,633]" italics="true" pageId="31" pageNumber="32">R. niedermeyeri</emphasis>
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(
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<bibRefCitation id="EF8178AB56680C3853F98796FEAC4AA5" author="Brignoli P. M." box="[197,374,645,672]" journalOrPublisher="Revue suisse de Zoologie" pageId="31" pageNumber="32" pagination="409 - 413" part="79" refString="Brignoli P. M. 1972. Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae). Revue suisse de Zoologie 79: 409 - 413." title="Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae)" type="journal article" year="1972">Brignoli, 1972</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName id="4C107ED956680C3852B68795FD114AA5" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[394,715,645,672]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vonwicki">
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<emphasis id="B964D94856680C3852B68795FDC94A9A" box="[394,531,645,671]" italics="true" pageId="31" pageNumber="32">R. vonwicki</emphasis>
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<bibRefCitation id="EF8178AB56680C3851268796FD114AA5" author="Zonstein S. L." box="[538,715,645,672]" journalOrPublisher="Tethys Entomological Research" pageId="31" pageNumber="32" pagination="49 - 52" part="2" refString="Zonstein S. L. 2000. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C107ED956680C38503F8795FBBC4AA5" authority="(Charitonov, 1948)" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[771,1126,645,672]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zaitzevi">
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<emphasis id="B964D94856680C38503F8795FCA64A9A" box="[771,892,646,671]" italics="true" pageId="31" pageNumber="32">R. zaitzevi</emphasis>
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(
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<bibRefCitation id="EF8178AB56680C3850B78796FB874AA5" author="Charitonov D. E." box="[907,1117,645,672]" journalOrPublisher="Soobshcheniya Akademii Nauk Gruzinskoi SSR" pageId="31" pageNumber="32" pagination="135 - 139" part="9" refString="Charitonov D. E. 1948. A new species of Brachythele from Georgia. Soobshcheniya Akademii Nauk Gruzinskoi SSR 9: 135 - 139. [In Russian and Latin]" title="A new species of Brachythele from Georgia" type="journal article" year="1948">Charitonov, 1948</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A56D156680C38538187DFFAAC4F21" pageId="31" pageNumber="32" type="key">
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<paragraph id="8BAF055A56680C38538187DFFD704AED" blockId="31.[189,682,715,745]" box="[189,682,715,745]" pageId="31" pageNumber="32">
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<emphasis id="B964D94856680C38538187DFFD704AED" bold="true" box="[189,682,715,745]" pageId="31" pageNumber="32">
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<emphasis id="B964D94856680C38538187DFFE4D4AED" bold="true" box="[189,407,715,744]" italics="true" pageId="31" pageNumber="32">Key to species of</emphasis>
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<taxonomicName id="4C107ED956680C3852A387DFFD8E4AEC" ID-CoL="6WT99" baseAuthorityName="Brignoli" baseAuthorityYear="1972" box="[415,596,716,745]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="niedermeyeri">niedermeyeri</taxonomicName>
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<emphasis id="B964D94856680C38516087DFFD704AED" bold="true" box="[604,682,716,744]" italics="true" pageId="31" pageNumber="32">group</emphasis>
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</emphasis>
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</paragraph>
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<paragraph id="8BAF055A56680C3853D68610FEEF4B18" blockId="31.[189,1399,771,1352]" box="[234,309,771,797]" pageId="31" pageNumber="32">
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<emphasis id="B964D94856680C3853D68610FEEF4B18" bold="true" box="[234,309,771,797]" pageId="31" pageNumber="32">Males</emphasis>
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</paragraph>
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<paragraph id="8BAF055A56680C3853818639FAAC4B41" blockId="31.[189,1399,771,1352]" box="[189,1398,810,837]" pageId="31" pageNumber="32">
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1. Subapical part of embolus corkscrew-shaped (
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<figureCitation id="132B19DF56680C3850298639FC614B40" box="[789,955,810,837]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 189–196</figureCitation>
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) ……………………………………2
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</paragraph>
|
||
<paragraph id="8BAF055A56680C3853FD8642FAAC4B6E" blockId="31.[189,1399,771,1352]" box="[193,1398,848,875]" pageId="31" pageNumber="32">
|
||
– Subapical part of embolus gradually curved (
|
||
<figureCitation id="132B19DF56680C38503D8643FCBC4B6E" box="[769,870,848,875]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Fig. 197</figureCitation>
|
||
) …………
|
||
<taxonomicName id="4C107ED956680C3850D18642FAAC4B6E" ID-CoL="6WT99" authority="(Brignoli, 1972)" baseAuthorityName="Brignoli" baseAuthorityYear="1972" box="[1005,1398,848,875]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="niedermeyeri">
|
||
<emphasis id="B964D94856680C3850D18642FB744B6E" bold="true" box="[1005,1198,848,875]" italics="true" pageId="31" pageNumber="32">R. niedermeyeri</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56680C3857838643FAB44B6E" author="Brignoli P. M." box="[1215,1390,848,875]" journalOrPublisher="Revue suisse de Zoologie" pageId="31" pageNumber="32" pagination="409 - 413" part="79" refString="Brignoli P. M. 1972. Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae). Revue suisse de Zoologie 79: 409 - 413." title="Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae)" type="journal article" year="1972">Brignoli, 1972</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853818686FAAD4BAA" blockId="31.[189,1399,771,1352]" box="[189,1399,917,943]" pageId="31" pageNumber="32">
|
||
2. Embolus with toothed keel vestige (
|
||
<figureCitation id="132B19DF56680C3851A38686FC9F4BAA" box="[671,837,917,943]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 191–194</figureCitation>
|
||
) ………………………………………………3
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853FD86AFFAAD4BD3" blockId="31.[189,1399,771,1352]" box="[193,1399,956,983]" pageId="31" pageNumber="32">
|
||
– Embolus smooth, without keel vestige (
|
||
<figureCitation id="132B19DF56680C3851EE86AFFC294BD2" box="[722,1011,956,983]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 189–190, 195–196</figureCitation>
|
||
) ………………………………4
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853818112FAAD4C47" blockId="31.[189,1399,771,1352]" pageId="31" pageNumber="32">
|
||
3. Keel vestige high and pectinate, with two large and a few smaller teeth (
|
||
<figureCitation id="132B19DF56680C3857AB8112FA984C1E" box="[1175,1346,1025,1051]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 191–192</figureCitation>
|
||
) … …………………………………………………………………………………
|
||
<taxonomicName id="4C107ED956680C385744813BFAD14C47" ID-CoL="4RMTY" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[1144,1291,1063,1090]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
|
||
<emphasis id="B964D94856680C385744813BFAD14C47" bold="true" box="[1144,1291,1063,1090]" italics="true" pageId="31" pageNumber="32">R. marusiki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356680C38562B813BFAAD4C47" box="[1303,1399,1064,1090]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853FD815DFAAC4C8A" blockId="31.[189,1399,771,1352]" pageId="31" pageNumber="32">
|
||
– Keel vestige low and densely dentate, with several tiny, uniform and equal-sized teeth (
|
||
<figureCitation id="132B19DF56680C3857C3815DFECD4C8A" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 193– 194</figureCitation>
|
||
) …………………………………………………………………
|
||
<taxonomicName id="4C107ED956680C3857358166FAAC4C8A" ID-CoL="6WT8W" authority="(Charitonov, 1948)" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[1033,1398,1140,1167]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zaitzevi">
|
||
<emphasis id="B964D94856680C3857358166FB524C8B" bold="true" box="[1033,1160,1140,1167]" italics="true" pageId="31" pageNumber="32">R. zaitzevi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56680C3857A78167FAB44C8A" author="Charitonov D. E." box="[1179,1390,1140,1167]" journalOrPublisher="Soobshcheniya Akademii Nauk Gruzinskoi SSR" pageId="31" pageNumber="32" pagination="135 - 139" part="9" refString="Charitonov D. E. 1948. A new species of Brachythele from Georgia. Soobshcheniya Akademii Nauk Gruzinskoi SSR 9: 135 - 139. [In Russian and Latin]" title="A new species of Brachythele from Georgia" type="journal article" year="1948">Charitonov, 1948</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C38538181AAFAAD4CFF" blockId="31.[189,1399,771,1352]" pageId="31" pageNumber="32">
|
||
4. Embolus gently spiraled; proximal part of embolus (preceding basal bend) equal in length to following apical part (
|
||
<figureCitation id="132B19DF56680C3852CA81F3FD464CFF" box="[502,668,1248,1274]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 189–190</figureCitation>
|
||
) …
|
||
<taxonomicName id="4C107ED956680C3851F081F3FAAD4CFF" ID-CoL="4RMTZ" authority="Marusik, Zamani & Mirshamsi, 2014" authorityName="Marusik, Zamani & Mirshamsi" authorityYear="2014" box="[716,1399,1247,1274]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mazandaranica">
|
||
<emphasis id="B964D94856680C3851F081F3FC714CFF" bold="true" box="[716,939,1247,1274]" italics="true" pageId="31" pageNumber="32">R. mazandaranica</emphasis>
|
||
<bibRefCitation id="EF8178AB56680C38508A81F3FAAD4CFF" author="Marusik Y. M. & Zamani A. & Mirshamsi O." box="[950,1399,1248,1274]" journalOrPublisher="ZooKeys" pageId="31" pageNumber="32" pagination="1 - 10" part="463" publicationUrl="https://doi.org/10.3897/zookeys.463.8692" refString="Marusik Y. M., Zamani A. & Mirshamsi O. 2014. Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae). ZooKeys 463: 1 - 10. https: // doi. org / 10.3897 / zookeys. 463.8692" title="Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae)" type="journal article" year="2014">Marusik, Zamani & Mirshamsi, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853FD8014FAAD4D42" blockId="31.[189,1399,771,1352]" pageId="31" pageNumber="32">
|
||
– Embolus sharply spiraled; proximal part of embolus (preceding basal bend) considerably longer than following spiraled apical part (
|
||
<figureCitation id="132B19DF56680C3851A3803EFC9F4D4D" box="[671,837,1325,1352]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">Figs 195–196</figureCitation>
|
||
) …………………
|
||
<taxonomicName id="4C107ED956680C38571A803DFAAD4D42" ID-CoL="79599" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[1062,1399,1325,1352]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vonwicki">
|
||
<emphasis id="B964D94856680C38571A803DFB624D4D" bold="true" box="[1062,1208,1325,1352]" italics="true" pageId="31" pageNumber="32">R. vonwicki</emphasis>
|
||
<bibRefCitation id="EF8178AB56680C3857FF803EFAAD4D42" author="Zonstein S. L." box="[1219,1399,1325,1351]" journalOrPublisher="Tethys Entomological Research" pageId="31" pageNumber="32" pagination="49 - 52" part="2" refString="Zonstein S. L. 2000. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853D68061FE8B4D89" blockId="31.[189,1399,1394,1828]" box="[234,337,1394,1420]" pageId="31" pageNumber="32">
|
||
<emphasis id="B964D94856680C3853D68061FE8B4D89" bold="true" box="[234,337,1394,1420]" pageId="31" pageNumber="32">Females</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C385381808BFB9E4DB7" blockId="31.[189,1399,1394,1828]" box="[189,1092,1432,1459]" pageId="31" pageNumber="32">
|
||
Females of
|
||
<taxonomicName id="4C107ED956680C38527A808BFDF14DB7" ID-CoL="4RMTY" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[326,555,1432,1458]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
|
||
<emphasis id="B964D94856680C38527A808BFDF14DB7" box="[326,555,1432,1458]" italics="true" pageId="31" pageNumber="32">Raveniola marusiki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356680C38510E808AFD574DB6" box="[562,653,1433,1459]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C107ED956680C3851F9808AFC474DB7" ID-CoL="4RMTZ" authorityName="Marusik, Zamani & Mirshamsi" authorityYear="2014" box="[709,925,1432,1458]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mazandaranica">
|
||
<emphasis id="B964D94856680C3851F9808AFC474DB7" box="[709,925,1432,1458]" italics="true" pageId="31" pageNumber="32">R. mazandaranica</emphasis>
|
||
</taxonomicName>
|
||
are unknown.
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C38538180CEFAAC4E1B" blockId="31.[189,1399,1394,1828]" pageId="31" pageNumber="32">
|
||
1. Median and lateral receptacles subequal in size or median ones slightly larger than laterals (
|
||
<figureCitation id="132B19DF56680C3853CF8317FE4E4E1B" box="[243,404,1540,1566]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="31" pageNumber="32">Figs 210–212</figureCitation>
|
||
) ………………………………………………………………………………………2
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853FD8338FAAC4E6E" blockId="31.[189,1399,1394,1828]" pageId="31" pageNumber="32">
|
||
– Median receptacles considerably smaller than laterals (
|
||
<figureCitation id="132B19DF56680C3850408339FBF84E40" box="[892,1058,1578,1605]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="31" pageNumber="32">Figs 208–209</figureCitation>
|
||
) …………………………… ………………………………………………………………………
|
||
<taxonomicName id="4C107ED956680C38573E8341FAAC4E6E" ID-CoL="6WT8W" authority="(Charitonov, 1948)" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[1026,1398,1617,1644]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zaitzevi">
|
||
<emphasis id="B964D94856680C38573E8341FB594E6E" bold="true" box="[1026,1155,1617,1644]" italics="true" pageId="31" pageNumber="32">R. zaitzevi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56680C3857A58342FAB44E6E" author="Charitonov D. E." box="[1177,1390,1617,1643]" journalOrPublisher="Soobshcheniya Akademii Nauk Gruzinskoi SSR" pageId="31" pageNumber="32" pagination="135 - 139" part="9" refString="Charitonov D. E. 1948. A new species of Brachythele from Georgia. Soobshcheniya Akademii Nauk Gruzinskoi SSR 9: 135 - 139. [In Russian and Latin]" title="A new species of Brachythele from Georgia" type="journal article" year="1948">Charitonov, 1948</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853818385FAAD4ED2" blockId="31.[189,1399,1394,1828]" pageId="31" pageNumber="32">
|
||
2. Median and lateral receptacles subequal in size; lateral ones with distinct stalk (
|
||
<figureCitation id="132B19DF56680C38579B8385FAD74EB5" box="[1191,1293,1686,1712]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="31" pageNumber="32">Fig. 210</figureCitation>
|
||
) ……… …………………………………………………………………………
|
||
<taxonomicName id="4C107ED956680C38571C83AEFAAD4ED2" ID-CoL="79599" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[1056,1399,1724,1751]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vonwicki">
|
||
<emphasis id="B964D94856680C38571C83AEFB6E4ED2" bold="true" box="[1056,1204,1724,1751]" italics="true" pageId="31" pageNumber="32">R. vonwicki</emphasis>
|
||
<bibRefCitation id="EF8178AB56680C3857FD83AFFAAD4ED2" author="Zonstein S. L." box="[1217,1399,1724,1751]" journalOrPublisher="Tethys Entomological Research" pageId="31" pageNumber="32" pagination="49 - 52" part="2" refString="Zonstein S. L. 2000. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56680C3853FD83F7FAAC4F21" blockId="31.[189,1399,1394,1828]" pageId="31" pageNumber="32">
|
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– Median receptacles slightly larger than laterals; lateral heads sessile (
|
||
<figureCitation id="132B19DF56680C38577883F0FB364EF8" box="[1092,1260,1763,1789]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="31" pageNumber="32">Figs 211–212</figureCitation>
|
||
) ………… ……………………………………………………………………
|
||
<taxonomicName id="4C107ED956680C3850D98219FAAC4F21" ID-CoL="6WT99" authority="(Brignoli, 1972)" baseAuthorityName="Brignoli" baseAuthorityYear="1972" box="[997,1398,1801,1828]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="niedermeyeri">
|
||
<emphasis id="B964D94856680C3850D98219FB734F21" bold="true" box="[997,1193,1801,1828]" italics="true" pageId="31" pageNumber="32">R. niedermeyeri</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56680C3857808219FAB44F21" author="Brignoli P. M." box="[1212,1390,1802,1828]" journalOrPublisher="Revue suisse de Zoologie" pageId="31" pageNumber="32" pagination="409 - 413" part="79" refString="Brignoli P. M. 1972. Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae). Revue suisse de Zoologie 79: 409 - 413." title="Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae)" type="journal article" year="1972">Brignoli, 1972</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |