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<document id="69AA187E512BA37A9362B04652860A4D" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56640C3353818729FAAD4949" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola micropa in Uludag Mts" docType="key" docVersion="13" lastPageNumber="21" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="20" updateTime="1698518020641" updateUser="ExternalLinkService">
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<mods:title id="8949FA501101439BDEDAE70218E83C15">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="59598A2C48F0B44734CEB6929DAE20A9">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="AD65E472522D68EC8E1CFDECB5930757">Kunt, Kadir B.</mods:namePart>
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<mods:namePart id="E02173A126AF8252D5CCCC53B141803F">Yağmur, Ersen A.</mods:namePart>
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<mods:title id="ED1999B78BE61BD2B358FED839B20AA8">European Journal of Taxonomy</mods:title>
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<mods:date id="100D475890B880B2639F224369C4B762">2018</mods:date>
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<mods:number id="A757B2932276198241FDC6D96182544F">2018-01-30</mods:number>
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<treatment id="03B9B44C56640C3353818729FAAD4949" ID-DOI="http://doi.org/10.5281/zenodo.5980288" ID-GBIF-Taxon="140561346" ID-Zenodo-Dep="5980288" LSID="urn:lsid:plazi:treatment:03B9B44C56640C3353818729FAAD4949" httpUri="http://treatment.plazi.org/id/03B9B44C56640C3353818729FAAD4949" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="C30A56D156640C3453818729FDCF4A52" box="[189,533,570,599]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BAF055A56640C3453818729FDCF4A52" blockId="19.[189,533,570,599]" box="[189,533,570,599]" pageId="19" pageNumber="20">
<emphasis id="B964D94856640C3453818729FDCF4A52" bold="true" box="[189,533,570,599]" pageId="19" pageNumber="20">
<taxonomicName id="4C107ED956640C3453818729FE654A52" ID-CoL="6WSX9" authorityName="in Uludag Mts" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[189,447,570,599]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="micropa">Raveniola micropa</taxonomicName>
<emphasis id="B964D94856640C3452FB8728FDCF4A52" bold="true" box="[455,533,571,599]" italics="true" pageId="19" pageNumber="20">group</emphasis>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156640C345381877CFB674B22" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BAF055A56640C345381877CFE164A8F" blockId="19.[189,460,623,650]" box="[189,460,623,650]" pageId="19" pageNumber="20">
<heading id="D0E7B23656640C345381877CFE164A8F" bold="true" box="[189,460,623,650]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B964D94856640C345381877CFE164A8F" bold="true" box="[189,460,623,650]" pageId="19" pageNumber="20">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56640C345381878CFB674B22" blockId="19.[189,1399,671,807]" pageId="19" pageNumber="20">
Male palpal tibia moderately short and subcylindrical, without subapical sensilla (
<figureCitation id="132B19DF56640C345781878CFAB04ABF" box="[1213,1386,671,698]" captionStart-0="Figs 153–156" captionStart-1="Figs 157–160" captionStartId-0="74.[189,241,1894,1920]" captionStartId-1="75.[189,241,1894,1920]" captionTargetBox-0="[189,1398,265,1868]" captionTargetBox-1="[189,1395,266,1870]" captionTargetId-0="figure@74.[189,1398,265,1868]" captionTargetId-1="figure@75.[189,1398,265,1875]" captionTargetPageId-0="74" captionText-0="Figs 153–156. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 153. R. birecikensis sp. nov. (holotype). 154. Same species as preceding (paratype, Birecik, revolved bulbshowninventralposition).155.Samespeciesaspreceding(dwarf conspecific♂?). 156.R.anadolu sp. nov. (holotype). Scale bars: 0.5 mm." captionText-1="Figs 157–160. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 157. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 158. R. turcica sp. nov. (holotype). 159. R. micropa (Ausserer, 1871) (Uludağ). 160. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/1163472/files/figure.png" httpUri-1="https://zenodo.org/record/1163474/files/figure.png" pageId="19" pageNumber="20">Figs 153–159</figureCitation>
). Embolus moderately long, hooked or spiraled, with more or less developed vertically-oriented keel (as in
<figureCitation id="132B19DF56640C34523487FBFE744B07" box="[264,430,744,770]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 180–182</figureCitation>
), or with its rudiments (
<figureCitation id="132B19DF56640C3451F387FBFCED4B07" box="[719,823,744,770]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Fig. 183</figureCitation>
), or without keel (
<figureCitation id="132B19DF56640C34572887FBFB674B07" box="[1044,1213,744,770]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 184–188</figureCitation>
). Spermathecae moderately wide, with lateral receptacles situated proximal to medians (
<figureCitation id="132B19DF56640C345735861EFB6B4B22" box="[1033,1201,781,807]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="19" pageNumber="20">Figs 204–207</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156640C345381865CFC804BBB" pageId="19" pageNumber="20" type="synonymic_list">
<paragraph id="8BAF055A56640C345381865CFE564B6C" blockId="19.[189,396,847,873]" box="[189,396,847,873]" pageId="19" pageNumber="20">
<heading id="D0E7B23656640C345381865CFE564B6C" bold="true" box="[189,396,847,873]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B964D94856640C345381865CFE564B6C" bold="true" box="[189,396,847,873]" pageId="19" pageNumber="20">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56640C345381866CFC804BBB" blockId="19.[189,1399,895,958]" pageId="19" pageNumber="20">
<taxonomicName id="4C107ED956640C345381866CFE454B9C" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[189,415,895,921]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="anadolu" status="sp. nov.">
<emphasis id="B964D94856640C345381866CFE454B9C" box="[189,415,895,921]" italics="true" pageId="19" pageNumber="20">Raveniola anadolu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3452978693FDD74B9F" box="[427,525,896,922]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C107ED956640C3451218693FC6F4B9C" authority="Kunt &amp; Yagmur, 2010" authorityName="Kunt &amp; Yagmur" authorityYear="2010" box="[541,949,895,922]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B964D94856640C3451218693FD4E4B9C" box="[541,660,895,921]" italics="true" pageId="19" pageNumber="20">R. arthuri</emphasis>
<bibRefCitation id="EF8178AB56640C34519C8693FC6F4B9C" author="Kunt K. B. &amp; Yagmur E. A." box="[672,949,895,922]" journalOrPublisher="Turkish Journal of Zoology" pageId="19" pageNumber="20" pagination="305 - 309" part="34" publicationUrl="https://doi.org/10.3906/zoo-0906-40" refString="Kunt K. B. &amp; Yagmur E. A. 2010. A new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae; Nemesiidae) from Turkey. Turkish Journal of Zoology 34: 305 - 309. https: // doi. org / 10.3906 / zoo- 0906 - 40" title="A new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae; Nemesiidae) from Turkey" type="journal article" year="2010">Kunt &amp; Yağmur, 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C107ED956640C3450F88693FBAE4B9C" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[964,1140,895,921]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="birecikensis" status="sp. nov.">
<emphasis id="B964D94856640C3450F88693FBAE4B9C" box="[964,1140,895,921]" italics="true" pageId="19" pageNumber="20">R. birecikensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3457BC8693FB3B4B9F" box="[1152,1249,896,922]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C107ED956640C3457CD8693FE5E4BBB" authority="(Ausserer, 1871)" authorityName="in Uludag Mts" authorityYear="1871" baseAuthorityName="Ausserer" baseAuthorityYear="1871" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="micropa">
<emphasis id="B964D94856640C3457CD8693FAAD4B9C" box="[1265,1399,896,921]" italics="true" pageId="19" pageNumber="20">R. micropa</emphasis>
(
<bibRefCitation id="EF8178AB56640C3453F986B7FEA04BBB" author="Ausserer A." box="[197,378,932,958]" journalOrPublisher="Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien" pageId="19" pageNumber="20" pagination="177 - 224" part="21" refString="Ausserer A. 1871. Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 21: 177 - 224." title="Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor)" type="journal article" year="1871">Ausserer, 1871</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C107ED956640C3452AC86B6FE314BBB" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[400,491,933,958]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
<emphasis id="B964D94856640C3452AC86B6FE314BBB" box="[400,491,933,958]" italics="true" pageId="19" pageNumber="20">R. nana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3452CE86B7FD974BBB" box="[498,589,932,958]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C107ED956640C3451B986B6FD224BBB" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[645,760,933,958]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="turcica" status="sp. nov.">
<emphasis id="B964D94856640C3451B986B6FD224BBB" box="[645,760,933,958]" italics="true" pageId="19" pageNumber="20">R. turcica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3451C386B7FC804BBB" box="[767,858,932,958]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156640C33538186FAFAAD4949" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="key">
<paragraph id="8BAF055A56640C34538186FAFDBC4C00" blockId="19.[189,614,1000,1029]" box="[189,614,1000,1029]" pageId="19" pageNumber="20">
<emphasis id="B964D94856640C34538186FAFDBC4C00" bold="true" box="[189,614,1000,1029]" pageId="19" pageNumber="20">
<emphasis id="B964D94856640C34538186FAFE4D4C00" bold="true" box="[189,407,1000,1029]" italics="true" pageId="19" pageNumber="20">Key to species of</emphasis>
<taxonomicName id="4C107ED956640C3452A386FBFDCA4C00" ID-CoL="6WSX9" authorityName="in Uludag Mts" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[415,528,1000,1029]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="micropa">micropa</taxonomicName>
<emphasis id="B964D94856640C34512486FAFDBC4C00" bold="true" box="[536,614,1001,1029]" italics="true" pageId="19" pageNumber="20">group</emphasis>
</emphasis>
</paragraph>
<paragraph id="8BAF055A56640C3453D6810DFEEF4C3D" blockId="19.[189,695,1054,1116]" box="[234,309,1054,1080]" pageId="19" pageNumber="20">
<emphasis id="B964D94856640C3453D6810DFEEF4C3D" bold="true" box="[234,309,1054,1080]" pageId="19" pageNumber="20">Males</emphasis>
</paragraph>
<paragraph id="8BAF055A56640C3453818151FD6D4C59" blockId="19.[189,695,1054,1116]" box="[189,695,1090,1116]" pageId="19" pageNumber="20">
Male of
<taxonomicName id="4C107ED956640C34521C8151FE004C59" ID-CoL="6WT8X" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[288,474,1090,1116]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
<emphasis id="B964D94856640C34521C8151FE004C59" box="[288,474,1090,1116]" italics="true" pageId="19" pageNumber="20">Raveniola nana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3452DD8151FDE14C59" box="[481,571,1090,1116]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
unknown.
</paragraph>
<paragraph id="8BAF055A56640C3453818196FAAD4CC1" blockId="19.[189,1399,1156,1748]" pageId="19" pageNumber="20">
1. Proximal part of embolus short and flattened; embolic keel well-developed (
<figureCitation id="132B19DF56640C3457BF8197FAF34C9A" box="[1155,1321,1156,1183]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 180–182</figureCitation>
) …… ………………………………………………………………………………
<taxonomicName id="4C107ED956640C34576681BAFACA4CC1" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[1114,1296,1193,1220]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="biresecensis" status="sp. nov.">
<emphasis id="B964D94856640C34576681BAFACA4CC1" bold="true" box="[1114,1296,1193,1220]" pageId="19" pageNumber="20">
<emphasis id="B964D94856640C34576681BAFBB44CC6" bold="true" box="[1114,1134,1193,1219]" italics="true" pageId="19" pageNumber="20">R</emphasis>
.
<emphasis id="B964D94856640C34574381BAFACA4CC1" bold="true" box="[1151,1296,1193,1220]" italics="true" pageId="19" pageNumber="20">biresecensis</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C34562681B9FAAD4CC1" box="[1306,1399,1194,1220]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BAF055A56640C3453FD81DDFAAC4D09" blockId="19.[189,1399,1156,1748]" pageId="19" pageNumber="20">
– Proximal part of embolus longer, gradually tapering and curved; keel vestigial or absent (
<figureCitation id="132B19DF56640C34563C81DEFECD4D09" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 183– 188</figureCitation>
) …………………………………………………………………………………………………2
</paragraph>
<paragraph id="8BAF055A56640C3453818026FAAD4D4A" blockId="19.[189,1399,1156,1748]" box="[189,1399,1332,1359]" pageId="19" pageNumber="20">
2. Embolus hooked (
<figureCitation id="132B19DF56640C3452F58026FD6A4D4A" box="[457,688,1332,1359]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 183, 185–186</figureCitation>
) ……………………………………………………………3
</paragraph>
<paragraph id="8BAF055A56640C3453FD8049FAAD4D76" blockId="19.[189,1399,1156,1748]" box="[193,1399,1369,1396]" pageId="19" pageNumber="20">
– Embolus corkscrew-shaped (
<figureCitation id="132B19DF56640C34517F804AFCF24D76" box="[579,808,1369,1395]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 184, 187–188</figureCitation>
) …………………………………………………4
</paragraph>
<paragraph id="8BAF055A56640C345381808FFAAD4DDE" blockId="19.[189,1399,1156,1748]" pageId="19" pageNumber="20">
3. Embolus with small keel vestige, subequal in length to tegulum, bent medially (
<figureCitation id="132B19DF56640C3457FC808FFAF24DB3" box="[1216,1320,1436,1462]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Fig. 183</figureCitation>
) …… …………………………………………………………………………………
<taxonomicName id="4C107ED956640C34574480D2FADF4DDE" ID-CoL="4RMTG" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[1144,1285,1472,1499]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="anadolu" status="sp. nov.">
<emphasis id="B964D94856640C34574480D2FADF4DDE" bold="true" box="[1144,1285,1472,1499]" italics="true" pageId="19" pageNumber="20">R. anadolu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C34562880D2FAAD4DDE" box="[1300,1399,1473,1499]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BAF055A56640C3453FD80F6FAAD4E21" blockId="19.[189,1399,1156,1748]" pageId="19" pageNumber="20">
– Embolus without keel vestige, considerably longer than tegulum, bent subapically (
<figureCitation id="132B19DF56640C3457F580F6FAB44E05" box="[1225,1390,1509,1536]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 185–186</figureCitation>
) ……………………………………………………………………………………
<taxonomicName id="4C107ED956640C3457A98319FAD54E26" ID-CoL="794X9" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[1173,1295,1545,1572]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="turcica" status="sp. nov.">
<emphasis id="B964D94856640C3457A98319FAD54E26" bold="true" box="[1173,1295,1545,1572]" italics="true" pageId="19" pageNumber="20">R. turcica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3456258319FAAD4E21" box="[1305,1399,1546,1572]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BAF055A56640C345381835FFAAD4E8E" blockId="19.[189,1399,1156,1748]" pageId="19" pageNumber="20">
4. Proximal part of embolus (preceding basal bend) equal in length to spiraled apical part (
<figureCitation id="132B19DF56640C345634835FFAB44E63" box="[1288,1390,1612,1638]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Fig. 184</figureCitation>
) ……………………………………………………………………
<taxonomicName id="4C107ED956640C3450D98362FAAD4E8E" ID-CoL="4RMTJ" authority="Kunt &amp; Yagmur, 2010" authorityName="Kunt &amp; Yagmur" authorityYear="2010" box="[997,1399,1648,1675]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B964D94856640C3450D98362FBBA4E8E" bold="true" box="[997,1120,1648,1675]" italics="true" pageId="19" pageNumber="20">R. arthuri</emphasis>
<bibRefCitation id="EF8178AB56640C3457568362FAAD4E8E" author="Kunt K. B. &amp; Yagmur E. A." box="[1130,1399,1649,1675]" journalOrPublisher="Turkish Journal of Zoology" pageId="19" pageNumber="20" pagination="305 - 309" part="34" publicationUrl="https://doi.org/10.3906/zoo-0906-40" refString="Kunt K. B. &amp; Yagmur E. A. 2010. A new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae; Nemesiidae) from Turkey. Turkish Journal of Zoology 34: 305 - 309. https: // doi. org / 10.3906 / zoo- 0906 - 40" title="A new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae; Nemesiidae) from Turkey" type="journal article" year="2010">Kunt &amp; Yağmur, 2010</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BAF055A56640C3453FD8386FAAD4ED6" blockId="19.[189,1399,1156,1748]" pageId="19" pageNumber="20">
– Proximal part of embolus (preceding basal bend) considerably longer than spiraled apical part (
<figureCitation id="132B19DF56640C3453CF83AAFE414ED6" box="[243,411,1721,1747]" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt &amp; Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="19" pageNumber="20">Figs 187–188</figureCitation>
) ………………………………………………………
<taxonomicName id="4C107ED956640C34572B83A9FAAD4ED6" ID-CoL="6WSX9" authority="(Ausserer, 1871)" authorityName="in Uludag Mts" authorityYear="1871" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[1047,1399,1721,1748]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="micropa">
<emphasis id="B964D94856640C34572B83A9FB7B4ED6" bold="true" box="[1047,1185,1721,1748]" italics="true" pageId="19" pageNumber="20">R. micropa</emphasis>
(
<bibRefCitation id="EF8178AB56640C34578A83AAFAB44ED6" author="Ausserer A." box="[1206,1390,1721,1747]" journalOrPublisher="Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien" pageId="19" pageNumber="20" pagination="177 - 224" part="21" refString="Ausserer A. 1871. Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 21: 177 - 224." title="Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor)" type="journal article" year="1871">Ausserer, 1871</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BAF055A56640C3453D683EFFE8B4F13" blockId="19.[189,1227,1788,1851]" box="[234,337,1788,1814]" pageId="19" pageNumber="20">
<emphasis id="B964D94856640C3453D683EFFE8B4F13" bold="true" box="[234,337,1788,1814]" pageId="19" pageNumber="20">Females</emphasis>
</paragraph>
<paragraph id="8BAF055A56640C3453818233FB114F3F" blockId="19.[189,1227,1788,1851]" box="[189,1227,1824,1851]" pageId="19" pageNumber="20">
Females of
<taxonomicName id="4C107ED956640C34527A8232FDFE4F3E" ID-CoL="4RMTG" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[326,548,1825,1851]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="anadolu" status="sp. nov.">
<emphasis id="B964D94856640C34527A8232FDFE4F3E" box="[326,548,1825,1851]" italics="true" pageId="19" pageNumber="20">Raveniola anadolu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3451108232FD534F3E" box="[556,649,1825,1851]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C107ED956640C3451A88232FCD24F3E" ID-CoL="4RMTJ" authorityName="Kunt &amp; Yagmur" authorityYear="2010" box="[660,776,1825,1851]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B964D94856640C3451A88232FCD24F3E" box="[660,776,1825,1851]" italics="true" pageId="19" pageNumber="20">R. arthuri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C107ED956640C34507D8232FC374F3E" ID-CoL="4RMTL" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[833,1005,1825,1851]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="birecikensis" status="sp. nov.">
<emphasis id="B964D94856640C34507D8232FC374F3E" box="[833,1005,1825,1851]" italics="true" pageId="19" pageNumber="20">R. birecikensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356640C3450C88232FB954F3E" box="[1012,1103,1825,1851]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
unknown.
</paragraph>
<paragraph id="8BAF055A56640C3453818270FAAD4FEE" blockId="19.[189,1399,1891,2028]" pageId="19" pageNumber="20">
1. Spermathecae moderately long: distance between median heads two times as long as spermathecae length (
<figureCitation id="132B19DF56640C345278829BFE324FA6" box="[324,488,1928,1955]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="19" pageNumber="20">Figs 204–205</figureCitation>
) ………………………………………………………………………………2 – Spermathecae short: distance between median heads three times as long as spermathecae length (
<figureCitation id="132B19DF56640C3453CF82C2FE824FE9" box="[243,344,2001,2028]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="19" pageNumber="20">Fig. 206</figureCitation>
) ………………………………………………………………
<taxonomicName id="4C107ED956640C34571A82C2FAAD4FEE" ID-CoL="6WSX9" authority="(Ausserer, 1871)" authorityName="in Uludag Mts" authorityYear="1871" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[1062,1399,2001,2027]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="micropa">
<emphasis id="B964D94856640C34571A82C2FB714FEE" bold="true" box="[1062,1195,2001,2027]" italics="true" pageId="19" pageNumber="20">R. micropa</emphasis>
(
<bibRefCitation id="EF8178AB56640C34578782C2FAB44FEE" author="Ausserer A." box="[1211,1390,2001,2027]" journalOrPublisher="Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien" pageId="19" pageNumber="20" pagination="177 - 224" part="21" refString="Ausserer A. 1871. Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 21: 177 - 224." title="Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor)" type="journal article" year="1871">Ausserer, 1871</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BAF055A56630C3353818418FAAD4923" blockId="20.[189,1399,267,332]" box="[189,1399,267,294]" pageId="20" pageNumber="21">
2. Median and lateral receptacles located close to each other (
<figureCitation id="132B19DF56630C3350948418FBD44920" box="[936,1038,267,293]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="20" pageNumber="21">Fig. 204</figureCitation>
) ……………
<taxonomicName id="4C107ED956630C33578D841FFAC94923" ID-CoL="6WT8X" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[1201,1299,268,294]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
<emphasis id="B964D94856630C33578D841FFAC94923" bold="true" box="[1201,1299,268,294]" italics="true" pageId="20" pageNumber="21">R. nana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356630C335627841FFAAD4923" box="[1307,1399,268,294]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BAF055A56630C3353FD8421FAAD4949" blockId="20.[189,1399,267,332]" box="[193,1399,305,332]" pageId="20" pageNumber="21">
– Median and lateral receptacles wide apart (
<figureCitation id="132B19DF56630C3351C98422FC864949" box="[757,860,305,332]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="20" pageNumber="21">Fig. 205</figureCitation>
) …………………………
<taxonomicName id="4C107ED956630C3357A98421FAD5494E" ID-CoL="794X9" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[1173,1295,305,332]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="turcica" status="sp. nov.">
<emphasis id="B964D94856630C3357A98421FAD5494E" bold="true" box="[1173,1295,305,332]" italics="true" pageId="20" pageNumber="21">R. turcica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356630C3356258421FAAD4949" box="[1305,1399,306,332]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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