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<document id="768D6870DD8DE8EB37ADF23624DF54A3" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56560C035106862BFD224B39" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola zaitzevi" docType="treatment" docVersion="12" lastPageNumber="37" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="34" updateTime="1698518020641" updateUser="ExternalLinkService">
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<mods:title id="227BDE3C1F0FFB9388D4EE86E762B6C5">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="530EADCF3367AC802828048EF20A73E6">Yağmur, Ersen A.</mods:namePart>
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<taxonomicName id="4C107ED956560C065106862BFC234B57" ID-CoL="6WT8W" authority="(Charitonov, 1948)" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[570,1017,824,851]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C065106862BFCC94B57" bold="true" box="[570,787,824,851]" italics="true" pageId="33" pageNumber="34">Raveniola zaitzevi</emphasis>
(
<bibRefCitation id="EF8178AB56560C06501F862BFC2B4B57" author="Charitonov D. E." box="[803,1009,824,850]" journalOrPublisher="Soobshcheniya Akademii Nauk Gruzinskoi SSR" pageId="33" pageNumber="34" pagination="135 - 139" part="9" refString="Charitonov D. E. 1948. A new species of Brachythele from Georgia. Soobshcheniya Akademii Nauk Gruzinskoi SSR 9: 135 - 139. [In Russian and Latin]" title="A new species of Brachythele from Georgia" type="journal article" year="1948">Charitonov, 1948</bibRefCitation>
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<paragraph id="8BAF055A56560C0653E3864FFA8F4B73" blockId="33.[223,1365,824,887]" box="[223,1365,860,887]" pageId="33" pageNumber="34">
<figureCitation id="132B19DF56560C0653E3864FFEE34B73" box="[223,313,860,886]" captionStart="Figs 7–12" captionStartId="51.[189,241,1719,1745]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@51.[189,1399,265,1680]" captionText="Figs 7–12. Raveniola spp., ♀♀. Tarsal organ, legs I (7–8, 10, 12) and IV (9, 11), dorsal views. — 7–9. R. hyrcanica Dunin, 1988. 10. R. pontica (Spassky, 1937). 11. R. micropa (Ausserer, 1871). 12. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163426/files/figure.png" pageId="33" pageNumber="34">Figs 12</figureCitation>
,
<figureCitation id="132B19DF56560C065279864FFEBE4B73" box="[325,356,860,886]" captionStart="Figs 13–18" captionStartId="52.[189,241,1725,1751]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@52.[189,1399,265,1680]" captionText="Figs 13–18. Raveniola spp., ♀♀. Bases of trichobothria, legs I (13, 15–16, 18) and IV (14, 17), dorsal views.—13–14.R.hyrcanicaDunin,1988.15.R.pontica(Spassky,1937).16–17.R.micropa(Ausserer, 1871). 18. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163428/files/figure.png" pageId="33" pageNumber="34">18</figureCitation>
,
<figureCitation id="132B19DF56560C06524D864FFE674B73" box="[369,445,860,886]" captionStart="Figs 25–30" captionStartId="54.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@54.[189,1399,265,1680]" captionText="Figs 25–30. Raveniolaspp.,♀♀. Posteriormedian spinnerets (25–27),detailsof spigots (28–30);apical (25–26,28–30)andventral(27)views.—25,29–30.R.hyrcanicaDunin,1988.26.R.pontica(Spassky, 1937). 27–28. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163432/files/figure.png" pageId="33" pageNumber="34">27–28</figureCitation>
,
<figureCitation id="132B19DF56560C0652F5864FFDCF4B73" box="[457,533,860,886]" captionStart="Figs 31–36" captionStartId="55.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@55.[189,1399,265,1680]" captionText="Figs 31–36. Raveniola spp., ♀♀. Posterior lateral spinnerets (31–33), with spigots in close-up view (34–36); proventral (31), apical (32, 36) and ventral (33–35) views. — 31, 34. R. zaitzevi (Charitonov, 1948). 32, 36. R. micropa (Ausserer, 1871). 33, 35. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163434/files/figure.png" pageId="33" pageNumber="34">31, 34</figureCitation>
,
<figureCitation id="132B19DF56560C06511D864FFDB64B72" box="[545,620,860,887]" captionStart="Figs 52–58" captionStartId="58.[189,241,1833,1859]" captionTargetBox="[189,1398,265,1770]" captionTargetId="figure@58.[189,1398,265,1771]" captionText="Figs 52–58. Raveniola spp., habitus of ♂♂ (52–53, 55, 57) and ♀♀ (54, 56, 58), lateral (52) and dorsal (53–58) views. — 52. R. biresekensis sp. nov. (dwarf conspecific male?). 53–54. R. micropa (Ausserer, 1871). 55–56. R. zaizevi (Charitonov, 1948). 57–58. R. niedermeyeri (Brignoli, 1972). Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163440/files/figure.png" pageId="33" pageNumber="34">55–56</figureCitation>
,
<figureCitation id="132B19DF56560C065145864FFD2B4B73" box="[633,753,860,886]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="33" pageNumber="34">61, 63, 66</figureCitation>
,
<figureCitation id="132B19DF56560C0651C1864FFCC74B73" box="[765,797,860,886]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="33" pageNumber="34">80</figureCitation>
,
<figureCitation id="132B19DF56560C065015864FFC934B73" box="[809,841,860,886]" captionStart="Figs 90–95" captionStartId="64.[189,241,1891,1917]" captionTargetBox="[189,1398,265,1846]" captionTargetId="figure@64.[189,1398,265,1849]" captionText="Figs 90–95. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 90. R. turcica sp. nov. (paratype). 91. R. micropa (Ausserer, 1871) (holotype). 92. Same species as preceding (Uludağ). 93. R. zaitzevi(Charitonov, 1948)(Lagodekhi).94.R.vonwickiZonstein, 2000(paratype).95.R.niedermeyeri (Brignoli, 1972) (paratype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163452/files/figure.png" pageId="33" pageNumber="34">93</figureCitation>
,
<figureCitation id="132B19DF56560C065069864FFC584B73" box="[853,898,860,886]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="33" pageNumber="34">113</figureCitation>
,
<figureCitation id="132B19DF56560C0650B2864FFC674B73" box="[910,957,860,886]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="33" pageNumber="34">126</figureCitation>
,
<figureCitation id="132B19DF56560C0650F5864FFC2D4B73" box="[969,1015,860,886]" captionStart="Figs 141–144" captionStartId="71.[189,241,1858,1884]" captionTargetBox="[239,1352,266,1791]" captionTargetId="figure@71.[206,1378,265,1793]" captionText="Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163466/files/figure.png" pageId="33" pageNumber="34">141</figureCitation>
,
<figureCitation id="132B19DF56560C06573F864FFBEB4B73" box="[1027,1073,860,886]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="33" pageNumber="34">161</figureCitation>
,
<figureCitation id="132B19DF56560C065702864FFB7D4B73" box="[1086,1191,860,886]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="33" pageNumber="34">193–194</figureCitation>
,
<figureCitation id="132B19DF56560C06578F864FFAC74B73" box="[1203,1309,860,886]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="33" pageNumber="34">208–209</figureCitation>
,
<figureCitation id="132B19DF56560C065615864FFA8F4B73" box="[1321,1365,860,886]" captionStart="Figure 220" captionStartId="86.[189,272,1534,1560]" captionTargetBox="[190,1398,265,1471]" captionTargetId="figure@86.[189,1398,265,1471]" captionText="Figure 220. Localities of Raveniola spp. in the eastern part of the Caucasus: R. dunini sp. nov. (), R. hyrcanica Dunin, 1988 () and R. zaitzevi (Charitonov, 1948) ().Abbreviation:TU = Turkey." httpUri="https://zenodo.org/record/1163496/files/figure.png" pageId="33" pageNumber="34">220</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156560C065381868DFB714C8E" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="8BAF055A56560C065381868DFD2A4BBD" blockId="33.[189,898,926,989]" box="[189,752,926,953]" pageId="33" pageNumber="34">
<taxonomicName id="4C107ED956560C065381868DFD624BBC" authority="Charitonov, 1948: 135" authorityName="Charitonov" authorityPageNumber="135" authorityYear="1948" box="[189,696,926,953]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C065381868DFE704BBD" box="[189,426,926,952]" italics="true" pageId="33" pageNumber="34">Brachythele zaitzevi</emphasis>
<bibRefCitation id="EF8178AB56560C06528D868DFD5B4BBD" author="Charitonov D. E." box="[433,641,926,952]" journalOrPublisher="Soobshcheniya Akademii Nauk Gruzinskoi SSR" pageId="33" pageNumber="34" pagination="135 - 139" part="9" refString="Charitonov D. E. 1948. A new species of Brachythele from Georgia. Soobshcheniya Akademii Nauk Gruzinskoi SSR 9: 135 - 139. [In Russian and Latin]" title="A new species of Brachythele from Georgia" type="journal article" year="1948">Charitonov, 1948</bibRefCitation>
: 135
</taxonomicName>
(♀).
</paragraph>
<paragraph id="8BAF055A56560C06538186D1FC584BD8" blockId="33.[189,898,926,989]" box="[189,898,962,989]" pageId="33" pageNumber="34">
<taxonomicName id="4C107ED956560C06538186D1FD544BD9" authority="Mcheidze, 1983: 122" authorityName="Mcheidze" authorityPageNumber="122" authorityYear="1983" box="[189,654,962,988]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="recki">
<emphasis id="B964D94856560C06538186D1FE564BD9" box="[189,396,962,988]" italics="true" pageId="33" pageNumber="34">Brachythele recki</emphasis>
<bibRefCitation id="EF8178AB56560C0652AF86D1FD8E4BD9" author="Mcheidze T. S." box="[403,596,962,988]" editor="Eliashvili T." journalOrPublisher="Metzniereba Publishing House, Tbilisi. [In Russian]" pageId="33" pageNumber="34" pagination="155 - 159" refString="Mcheidze T. S. 1983. A new spider species, Brachythele recki sp. nov., from Georgia. In: Eliashvili T. (ed.) Fauna i Ekologia Bespozvonochnykh Gruzii: 155 - 159. Metzniereba Publishing House, Tbilisi. [In Russian]" title="A new spider species, Brachythele recki sp. nov., from Georgia" type="book chapter" volumeTitle="Fauna i Ekologia Bespozvonochnykh Gruzii" year="1983">Mcheidze, 1983</bibRefCitation>
: 122
</taxonomicName>
, fig. 1 (♀).
<emphasis id="B964D94856560C06502A86D0FC584BD8" bold="true" box="[790,898,963,989]" pageId="33" pageNumber="34">syn. nov.</emphasis>
</paragraph>
<paragraph id="8BAF055A56560C0653818116FC1C4C1B" blockId="33.[189,1399,1028,1163]" box="[189,966,1028,1055]" pageId="33" pageNumber="34">
<taxonomicName id="4C107ED956560C0653818116FE704C1B" authorityName="Charitonov" authorityYear="1948" box="[189,426,1029,1055]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C0653818116FE704C1B" box="[189,426,1029,1055]" italics="true" pageId="33" pageNumber="34">Brachythele zaitzevi</emphasis>
</taxonomicName>
–
<bibRefCitation id="EF8178AB56560C0652FB8116FDB14C1A" author="Brignoli P. M." box="[455,619,1029,1055]" journalOrPublisher="Manchester University Press, Manchester" pageId="33" pageNumber="34" refString="Brignoli P. M. 1983. A Catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester." title="A Catalogue of the Araneae described between 1940 and 1981" type="book" year="1983">Brignoli 1983</bibRefCitation>
: 123. —
<bibRefCitation id="EF8178AB56560C0651E68116FC594C1A" author="Zonstein S. L." box="[730,899,1028,1055]" journalOrPublisher="Trudy zoologicheskogo Instituta AN SSSR" pageId="33" pageNumber="34" pagination="156 - 161" part="139" refString="Zonstein S. L. 1985. Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR. Trudy zoologicheskogo Instituta AN SSSR 139: 156 - 161. [In Russian with English summary]" title="Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR" type="journal article" year="1985">Zonstein 1985</bibRefCitation>
: 160.
</paragraph>
<paragraph id="8BAF055A56560C065381813AFAAD4C62" blockId="33.[189,1399,1028,1163]" pageId="33" pageNumber="34">
<taxonomicName id="4C107ED956560C065381813AFE4C4C47" ID-CoL="6WT8W" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[189,406,1065,1091]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C065381813AFE4C4C47" box="[189,406,1065,1091]" italics="true" pageId="33" pageNumber="34">Raveniola zaitzevi</emphasis>
</taxonomicName>
–
<bibRefCitation id="EF8178AB56560C06528B813AFDB94C46" author="Zonstein S. L." box="[439,611,1065,1091]" journalOrPublisher="Zoologicheskii Zhurnal" pageId="33" pageNumber="34" pagination="1013 - 1019" part="66" refString="Zonstein S. L. 1987. A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna. Zoologicheskii Zhurnal 66 (7): 1013 - 1019. [In Russian with English summary]" title="A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna" type="journal article" year="1987">Zonstein 1987</bibRefCitation>
: 1015. —
<bibRefCitation id="EF8178AB56560C0651DB813AFCAF4C46" author="Dunin P. M." box="[743,885,1065,1091]" journalOrPublisher="Zoologicheskii Zhurnal" pageId="33" pageNumber="34" pagination="1245 - 1248" part="67" refString="Dunin P. M. 1988. Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan. Zoologicheskii Zhurnal 67 (8): 1245 - 1248. [In Russian with English summary]" title="Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan" type="journal article" year="1988">Dunin 1988</bibRefCitation>
: 1247; 1989: 33. —
<bibRefCitation id="EF8178AB56560C065749813AFAC34C46" author="Platnick N. I." box="[1141,1305,1065,1091]" journalOrPublisher="A Supplement to Brignoli's A Catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester and New York" pageId="33" pageNumber="34" pagination="1981 - 1987" refString="Platnick N. I. 1989. Advances in Spider Taxonomy 1981 - 1987: A Supplement to Brignoli's A Catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester and New York." title="Advances in Spider Taxonomy" type="book" year="1989">Platnick 1989</bibRefCitation>
: 91. —
<bibRefCitation id="EF8178AB56560C0653D6815EFE7C4C62" author="Mikhailov K. G." box="[234,422,1101,1127]" journalOrPublisher="Zoological Museum of the Moscow State University, Moscow" pageId="33" pageNumber="34" refString="Mikhailov K. G. 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum of the Moscow State University, Moscow." title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)" type="book" year="1997">Mikhailov 1997</bibRefCitation>
: 20. —
<bibRefCitation id="EF8178AB56560C065138815EFC704C62" author="Guseinov E. F. &amp; Aliev H. A. &amp; Snegovaya N. Y." box="[516,938,1101,1127]" journalOrPublisher="Materials of the 1 st Congress of Azerbaijan Society of Zoologists" pageId="33" pageNumber="34" pagination="191 - 195" refString="Guseinov E. F., Aliev H. A. &amp; Snegovaya N. Y. 2003. To the knowledge of the spider fauna of Ismailly Reserve. Materials of the 1 st Congress of Azerbaijan Society of Zoologists: 191 - 195. [In Azerbaijani with English summary]" title="To the knowledge of the spider fauna of Ismailly Reserve" type="proceedings" year="2003">Guseinov, Aliev &amp; Snegovaya 2003</bibRefCitation>
: 192. —
<bibRefCitation id="EF8178AB56560C06572E815EFAEF4C62" author="Guseinov E. F. &amp; Alieva T. V." box="[1042,1333,1101,1127]" journalOrPublisher="Proceedings of the Azerbaijan Society of Zoologists" pageId="33" pageNumber="34" pagination="281 - 287" part="1" refString="Guseinov E. F. &amp; Alieva T. V. 2008. The fauna of spiders (Arachnida: Araneae) of Pirkuli Reserve, Azerbaijan. Proceedings of the Azerbaijan Society of Zoologists 1: 281 - 287. [In Russian with English summary]" title="The fauna of spiders (Arachnida: Araneae) of Pirkuli Reserve, Azerbaijan" type="proceedings paper" year="2008">Guseinov &amp; Alieva 2008</bibRefCitation>
: 281.
</paragraph>
<paragraph id="8BAF055A56560C0653818162FB714C8E" blockId="33.[189,1399,1028,1163]" box="[189,1195,1136,1163]" pageId="33" pageNumber="34">
<taxonomicName id="4C107ED956560C0653818162FEAF4C8E" ID-CoL="6X58X" baseAuthorityName="Mcheidze" baseAuthorityYear="1983" box="[189,373,1137,1163]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="recki">
<emphasis id="B964D94856560C0653818162FEAF4C8E" box="[189,373,1137,1163]" italics="true" pageId="33" pageNumber="34">Raveniola recki</emphasis>
</taxonomicName>
–
<bibRefCitation id="EF8178AB56560C0652AE8162FDE14C8E" author="Zonstein S. L." box="[402,571,1137,1163]" journalOrPublisher="Zoologicheskii Zhurnal" pageId="33" pageNumber="34" pagination="1013 - 1019" part="66" refString="Zonstein S. L. 1987. A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna. Zoologicheskii Zhurnal 66 (7): 1013 - 1019. [In Russian with English summary]" title="A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna" type="journal article" year="1987">Zonstein 1987</bibRefCitation>
: 1015. —
<bibRefCitation id="EF8178AB56560C0651848162FC814C8E" author="Platnick N. I." box="[696,859,1137,1163]" journalOrPublisher="A Supplement to Brignoli's A Catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester and New York" pageId="33" pageNumber="34" pagination="1981 - 1987" refString="Platnick N. I. 1989. Advances in Spider Taxonomy 1981 - 1987: A Supplement to Brignoli's A Catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester and New York." title="Advances in Spider Taxonomy" type="book" year="1989">Platnick 1989</bibRefCitation>
: 90. —
<bibRefCitation id="EF8178AB56560C0650878162FBAD4C8E" author="Mikhailov K. G." box="[955,1143,1137,1163]" journalOrPublisher="Zoological Museum of the Moscow State University, Moscow" pageId="33" pageNumber="34" refString="Mikhailov K. G. 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum of the Moscow State University, Moscow." title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)" type="book" year="1997">Mikhailov 1997</bibRefCitation>
: 20.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156560C06538181A0FE0A4E60" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BAF055A56560C06538181A0FEEC4CC8" blockId="33.[189,310,1203,1229]" box="[189,310,1203,1229]" pageId="33" pageNumber="34">
<heading id="D0E7B23656560C06538181A0FEEC4CC8" bold="true" box="[189,310,1203,1229]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B964D94856560C06538181A0FEEC4CC8" bold="true" box="[189,310,1203,1229]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56560C06538181F0FE0A4E60" blockId="33.[189,1399,1250,1637]" pageId="33" pageNumber="34">
In the general shape of its embolus
<taxonomicName id="4C107ED956560C06516181F1FCE94CF9" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[605,819,1250,1276]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C06516181F1FCE94CF9" box="[605,819,1250,1276]" italics="true" pageId="33" pageNumber="34">Raveniola zaitzevi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C107ED956560C0650F081F0FB8F4CF9" authorityName="Zonstein" authorityYear="2000" box="[972,1109,1250,1276]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vonwicki">
<emphasis id="B964D94856560C0650F081F0FB8F4CF9" box="[972,1109,1250,1276]" italics="true" pageId="33" pageNumber="34">R. vonwicki</emphasis>
</taxonomicName>
and
<taxonomicName id="4C107ED956560C0657B281F0FACF4CF9" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[1166,1301,1250,1276]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
<emphasis id="B964D94856560C0657B281F0FACF4CF9" box="[1166,1301,1250,1276]" italics="true" pageId="33" pageNumber="34">R. marusiki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356560C06562081F0FAAD4CF8" box="[1308,1399,1251,1277]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="132B19DF56560C0653F98015FEB14D25" box="[197,363,1286,1312]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="33" pageNumber="34">Figs 193–194</figureCitation>
; cf.
<figureCitation id="132B19DF56560C0652A28015FD984D24" box="[414,578,1286,1313]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="33" pageNumber="34">Figs 195–196</figureCitation>
), but differs from the former in possessing an embolic keel, and from the latter by having completely different configuration of that keel (very low and evenly micro-serrate in
<taxonomicName id="4C107ED956560C065381805CFEEF4D6D" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[189,309,1359,1384]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C065381805CFEEF4D6D" box="[189,309,1359,1384]" italics="true" pageId="33" pageNumber="34">R. zaitzevi</emphasis>
</taxonomicName>
, and high heterogeneous in
<taxonomicName id="4C107ED956560C065143805CFCDC4D6D" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[639,774,1358,1384]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
<emphasis id="B964D94856560C065143805CFCDC4D6D" box="[639,774,1358,1384]" italics="true" pageId="33" pageNumber="34">R. marusiki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356560C065031805CFCBC4D6C" box="[781,870,1359,1385]" pageId="33" pageNumber="34" rank="species">sp. nov</taxonomicNameLabel>
). However, in the shape of the spermathecae,
<taxonomicName id="4C107ED956560C0653818060FEEF4D89" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[189,309,1395,1420]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C0653818060FEEF4D89" box="[189,309,1395,1420]" italics="true" pageId="33" pageNumber="34">R. zaitzevi</emphasis>
</taxonomicName>
shows some resemblance to
<taxonomicName id="4C107ED956560C0651B68060FCD14D89" box="[650,779,1395,1420]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="micropa">
<emphasis id="B964D94856560C0651B68060FCD14D89" box="[650,779,1395,1420]" italics="true" pageId="33" pageNumber="34">R. micropa</emphasis>
</taxonomicName>
from another species group, because they both share relatively short and conical spermathecal bases (but it should be noted that females of two closer relatives of
<taxonomicName id="4C107ED956560C0653E280A8FE824DD1" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[222,344,1467,1492]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C0653E280A8FE824DD1" box="[222,344,1467,1492]" italics="true" pageId="33" pageNumber="34">R. zaitzevi</emphasis>
</taxonomicName>
, namely
<taxonomicName id="4C107ED956560C0652FB80A8FD454DD1" authorityName="Marusik, Zamani &amp; Mirshamsi" authorityYear="2014" box="[455,671,1466,1492]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="mazandaranica">
<emphasis id="B964D94856560C0652FB80A8FD454DD1" box="[455,671,1466,1492]" italics="true" pageId="33" pageNumber="34">R. mazandaranica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C107ED956560C0651E780A8FCB94DD1" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[731,867,1466,1492]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
<emphasis id="B964D94856560C0651E780A8FCB94DD1" box="[731,867,1466,1492]" italics="true" pageId="33" pageNumber="34">R. marusiki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356560C06505080A8FC104DD0" box="[876,970,1467,1493]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
, are as yet unknown). Nevertheless, females of
<taxonomicName id="4C107ED956560C06527480CCFE1E4DFD" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[328,452,1503,1528]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856560C06527480CCFE1E4DFD" box="[328,452,1503,1528]" italics="true" pageId="33" pageNumber="34">R. zaitzevi</emphasis>
</taxonomicName>
are well-distinguishable from those of
<taxonomicName id="4C107ED956560C06509680CCFBEA4DFD" box="[938,1072,1503,1528]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="micropa">
<emphasis id="B964D94856560C06509680CCFBEA4DFD" box="[938,1072,1503,1528]" italics="true" pageId="33" pageNumber="34">R. micropa</emphasis>
</taxonomicName>
, because they have longer spermathecae with large globular lateral receptacles which are considerably wider than the medians (in
<taxonomicName id="4C107ED956560C0653818334FEE54E45" box="[189,319,1575,1600]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="micropa">
<emphasis id="B964D94856560C0653818334FEE54E45" box="[189,319,1575,1600]" italics="true" pageId="33" pageNumber="34">R. micropa</emphasis>
</taxonomicName>
spermathecae wider and shorter, and lateral receptacles are subequal to medians,
<figureCitation id="132B19DF56560C06563D8335FF374E60" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="33" pageNumber="34">Figs 208– 209</figureCitation>
; cf.
<figureCitation id="132B19DF56560C0652228359FE1E4E60" box="[286,452,1610,1637]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="33" pageNumber="34">Figs 206–207</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156560C055381839EFEBC4AAB" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BAF055A56560C065381839EFE704EA2" blockId="33.[189,426,1677,1703]" box="[189,426,1677,1703]" pageId="33" pageNumber="34">
<heading id="D0E7B23656560C065381839EFE704EA2" bold="true" box="[189,426,1677,1703]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B964D94856560C065381839EFE704EA2" bold="true" box="[189,426,1677,1703]" pageId="33" pageNumber="34">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B780F0756560C0653D683AFFC064EFF" ID-GBIF-Occurrence="1806244359" collectingDate="1937-06-11" collectionCode="ZMPU" collectorName="A. F. Zaitzev" country="Georgia" location="Lagodekhi" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<paragraph id="8BAF055A56560C0653D683AFFE864ED3" blockId="33.[189,988,1724,1787]" box="[234,348,1724,1750]" pageId="33" pageNumber="34">
<typeStatus id="54ABBBF856560C0653D683AFFE864ED3" box="[234,348,1724,1750]" pageId="33" pageNumber="34" type="holotype">
<emphasis id="B964D94856560C0653D683AFFE864ED3" bold="true" box="[234,348,1724,1750]" pageId="33" pageNumber="34">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BAF055A56560C06538183F3FC064EFF" blockId="33.[189,988,1724,1787]" box="[189,988,1760,1787]" pageId="33" pageNumber="34">
<collectingCountry id="F30745CA56560C06538183F3FE9C4EFF" box="[189,326,1760,1786]" name="Georgia" pageId="33" pageNumber="34">GEORGIA</collectingCountry>
:
<specimenCount id="9D16CED356560C06526C83F2FEBC4EFE" box="[336,358,1761,1787]" pageId="33" pageNumber="34" type="female">♀</specimenCount>
,
<location id="8ECF538156560C06524983F3FE2C4EFF" LSID="urn:lsid:plazi:treatment:03B9B44C56560C035106862BFD224B39:8ECF538156560C06524983F3FE2C4EFF" box="[373,502,1760,1786]" country="Georgia" name="Lagodekhi" pageId="33" pageNumber="34">Lagodekhi</location>
,
<collectingDate id="EFEADA7256560C06513D83F3FD434EFF" box="[513,665,1760,1787]" pageId="33" pageNumber="34" value="1937-06-11">
<date id="FFAE239A56560C06513D83F3FD434EFF" box="[513,665,1760,1787]" pageId="33" pageNumber="34" value="1937-06-11">11 Jun. 1937</date>
</collectingDate>
,
<collectorName id="26E5608C56560C06519883F3FCEF4EFF" box="[676,821,1760,1786]" pageId="33" pageNumber="34">A.F. Zaitzev</collectorName>
leg. (
<collectionCode id="ED019D9F56560C06504B83F3FC154EFF" box="[887,975,1760,1786]" collectionName="ZMPU" pageId="33" pageNumber="34">ZMPU</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BAF055A56560C0653D68230FEBD4F38" blockId="33.[189,888,1827,1889]" box="[234,359,1827,1853]" pageId="33" pageNumber="34">
<emphasis id="B964D94856560C0653D68230FEBD4F38" bold="true" box="[234,359,1827,1853]" pageId="33" pageNumber="34">Paratypes</emphasis>
</paragraph>
<paragraph id="8BAF055A56560C0653818254FCA24F64" blockId="33.[189,888,1827,1889]" box="[189,888,1862,1889]" pageId="33" pageNumber="34">
<materialsCitation id="3B780F0756560C0653818254FCA24F64" ID-GBIF-Occurrence="1806244366" box="[189,888,1862,1889]" pageId="33" pageNumber="34">GEORGIA: 2 ♀♀, collection data as for holotype (ZMPU).</materialsCitation>
</paragraph>
<paragraph id="8BAF055A56560C0653D6829BFCD84FA1" blockId="33.[189,1399,1928,2028]" box="[234,770,1928,1956]" pageId="33" pageNumber="34">
<emphasis id="B964D94856560C0653D6829BFE3B4FA7" bold="true" box="[234,481,1928,1954]" pageId="33" pageNumber="34">Additional material</emphasis>
(10 ♂♂, 62 ♀♀, 8 juvs)
</paragraph>
<paragraph id="8BAF055A56560C05538182BEFE064A42" blockId="33.[189,1399,1928,2028]" lastBlockId="34.[189,1399,265,584]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<materialsCitation id="3B780F0756560C06538182BEFAE34FC2" ID-GBIF-Occurrence="1806244367" box="[189,1337,1963,1991]" pageId="33" pageNumber="34">AZERBAIJAN: 1 ♀, Qakh, 41°25′ N, 46°55′ E, 700 m, 19 Jun. 1977, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756560C06567D82BEFBD44FEE" ID-GBIF-Occurrence="1806244386" pageId="33" pageNumber="34">1 ♀, Bucaq, 41°05′ N, 47°22′ E, 600 m, 7 Jul. 1978, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756560C05572982C2FCB84920" ID-GBIF-Occurrence="1806244375" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">2 ♀♀, Car (Dzhar), 41°40′ N, 46°41′ E, 800 m, 6–14 Jul. 1981, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C055055841FFD47494F" ID-GBIF-Occurrence="1806244384" pageId="34" pageNumber="35">7 ♂♂, 2 ♀♀, same locality but 900–1100 m, 17 Sep. 1987, S.L. Zonstein leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C05519F8423FE45496B" ID-GBIF-Occurrence="1806244383" pageId="34" pageNumber="35">1 ♀, Zaqatala, 41°38′ N, 46°38′ E, 650 m, 10 Jul. 1981, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C0552948447FECF4997" ID-GBIF-Occurrence="1806244372" pageId="34" pageNumber="35">12 ♀♀, Qalaybugurt, 40°45′ N, 48°34′ E, 900 m, 12 Jul. 1982, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C055227846BFE9249B3" ID-GBIF-Occurrence="1806244371" pageId="34" pageNumber="35">1 ♂, 1 ♀, Pirkuli Reserve, 40°46′ N, 48°35′ E, 1200–1400 m, 25–29 May 1984, D.V. Logunov leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C055273848FFC0149B3" ID-GBIF-Occurrence="1806244373" box="[335,987,412,439]" pageId="34" pageNumber="35">2 ♂♂, same collection data but 5–9 Sept. 1984 (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C0550DE848FFC8049DE" ID-GBIF-Occurrence="1806244368" pageId="34" pageNumber="35">8 ♀♀, 2 juv., Tazakand, 40°53′ N, 49°11′ E, 250 m, 23 May 1986, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C05505E84D2FD2A49FA" ID-GBIF-Occurrence="1806244393" pageId="34" pageNumber="35">12 ♀♀, 4 juvs, Vandam, 40°56′ N, 47°46′ E, 700 m, 17 Jun. 1986, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C0551C684F6FD494A26" ID-GBIF-Occurrence="1806244400" pageId="34" pageNumber="35">3 ♀♀, 2 juvs, Qusanca, 40°44′ N, 48°06′ E, 900 m, 20 Jun. 1986, P.M. Dunin leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756550C0551A1871AFE064A42" ID-GBIF-Occurrence="1835183919" pageId="34" pageNumber="35">12 ♀♀, Ismayilli, 40°49′ N, 48°10′ E, 700 m, 5 Oct. 1988, I. Pashenko leg. (TAU).</materialsCitation>
</paragraph>
<paragraph id="8BAF055A56550C0553818763FEBC4AAB" blockId="34.[189,1399,622,686]" pageId="34" pageNumber="35">GEORGIA: 7 ♀♀, Lagodekhi Reserve, 41°51′ N, 46°17′ E, 600–1200 m, 24 Jul. 1982, Y.M. Marusik leg. (ZMMU).</paragraph>
</subSubSection>
<subSubSection id="C30A56D156550C04538187C5FC144FEE" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="8BAF055A56550C05538187C5FE944AF5" blockId="34.[189,334,726,752]" box="[189,334,726,752]" pageId="34" pageNumber="35">
<heading id="D0E7B23656550C05538187C5FE944AF5" bold="true" box="[189,334,726,752]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B964D94856550C05538187C5FE944AF5" bold="true" box="[189,334,726,752]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56550C0552218615FCFB4B24" blockId="34.[189,801,774,837]" box="[285,801,774,801]" pageId="34" pageNumber="35">
<emphasis id="B964D94856550C0552218615FE874B25" bold="true" box="[285,349,774,800]" pageId="34" pageNumber="35">Male</emphasis>
(Car; described here for the first time)
</paragraph>
<paragraph id="8BAF055A56550C0553818638FE6D4B40" blockId="34.[189,801,774,837]" box="[189,439,810,837]" pageId="34" pageNumber="35">
HABITUS. See
<figureCitation id="132B19DF56550C0552618639FE6D4B40" box="[349,439,810,837]" captionStart="Figs 52–58" captionStartId="58.[189,241,1833,1859]" captionTargetBox="[189,1398,265,1770]" captionTargetId="figure@58.[189,1398,265,1771]" captionText="Figs 52–58. Raveniola spp., habitus of ♂♂ (52–53, 55, 57) and ♀♀ (54, 56, 58), lateral (52) and dorsal (53–58) views. — 52. R. biresekensis sp. nov. (dwarf conspecific male?). 53–54. R. micropa (Ausserer, 1871). 55–56. R. zaizevi (Charitonov, 1948). 57–58. R. niedermeyeri (Brignoli, 1972). Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163440/files/figure.png" pageId="34" pageNumber="35">Fig. 55.</figureCitation>
</paragraph>
<paragraph id="8BAF055A56550C055381867EFB7D4B82" blockId="34.[189,1191,877,904]" box="[189,1191,877,904]" pageId="34" pageNumber="35">MEASUREMENTS. TBL 13.80, CL 5.40, CW 4.67, LL 0.36, LW 0.87, SL 2.42, SW 2.28.</paragraph>
<paragraph id="8BAF055A56550C05538186BCFECB4C7B" blockId="34.[189,1399,943,1150]" pageId="34" pageNumber="35">COLOUR. Carapace, palps and legs dorsally brownish orange; eye tubercle with three brownish black spots surrounding AMEs and each of two groups including PME and lateral eyes; chelicerae orangered; sternum, labium, maxillae, palps and legs ventrally pale reddish brown; entire abdomen light greyish brown, with darker brown dorsal pattern represented by interrupted median lanceolate spot and few pairs of also interrupted transverse fasciae located posteriorly, spinnerets pale yellowish brown.</paragraph>
<paragraph id="8BAF055A56550C05538181BBFEB74D57" blockId="34.[189,1399,1191,1362]" pageId="34" pageNumber="35">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF56550C05502181B4FCA64CC4" box="[797,892,1191,1217]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="34" pageNumber="35">Fig. 80</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF56550C05563A81B4FAA94CC4" box="[1286,1395,1191,1217]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="34" pageNumber="35">Fig. 113</figureCitation>
. Eye diameters and interdistances: AME 0.13 (0.18), ALE 0.23, PLE 0.13, PME 0.13, AME– AME 0.12(0.07), ALE–AME 0.10(0.08), ALE–PLE 0.09, PLE–PME 0.03, PME–PME 0.35. Each cheliceral furrow with 10–11 promarginal teeth and 8–10 mesobasal denticles. Maxillae with 8–9 cuspules each.
</paragraph>
<paragraph id="8BAF055A56550C0553818068FEE64E04" blockId="34.[189,1399,1402,1537]" pageId="34" pageNumber="35">
LEGS. Tibia and metatarsus I as shown in
<figureCitation id="132B19DF56550C0551E08069FC934D90" box="[732,841,1402,1429]" captionStart="Figs 141–144" captionStartId="71.[189,241,1858,1884]" captionTargetBox="[239,1352,266,1791]" captionTargetId="figure@71.[206,1378,265,1793]" captionText="Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163466/files/figure.png" pageId="34" pageNumber="35">Fig. 141</figureCitation>
. Scopula: distal on metatarsi I–II, entire on tarsi I, narrowly divided on tarsi II, elsewhere absent. Trichobothria: 2 rows of 9–11 each on tibiae, 10–11 on metatarsi, 9–11 on tarsi, 8–9 on cymbium. Paired claws: inner and outer margins with 6–8 teeth each.
</paragraph>
<paragraph id="8BAF055A56550C0553818339FBAB4EFC" blockId="34.[189,1398,1578,1786]" pageId="34" pageNumber="35">SPINATION. Palp: femur d1–1–1, pd1, rd1; tibia d1–1–1, p1–1–1, r1, v2–3–2; cymbium d7(9). Leg I: femur d1–1–1–1(0), pd1–1, dr1–1; tibia p1–1, pv2–1(2), rv1–1–m–m; metatarsus p1, v1–2–2(3). Leg II: femur d1–1–1–1(0), pd1–1; patella p1–1; tibia p1–1–1, v2–2–3; metatarsus p1–1(0)–1, v2–2–3. Leg III: femur d1–1–1–1, pd1–1–1, rd1–1; patella p1–1, r1; tibia d1, p1–1, r1–1–1, v2–2–3; metatarsus d1–1, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur d1–1–1–0, pd1–1–1, rd1; patella p1(0)–1, r1; tibia d1–1, p1–1, r1–1–1, v2–2–3; metatarsus d1–1–2–1, p1–1–1, r1–1(0)–1, v2–1–1–3. Patella I aspinose.</paragraph>
<paragraph id="8BAF055A56550C0553818231FD8E4F81" blockId="34.[189,1398,1825,1924]" pageId="34" pageNumber="35">
PALP. Tibia, cymbium and palpal organ as shown in
<figureCitation id="132B19DF56550C05501D8232FC5C4F3E" box="[801,902,1825,1851]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="34" pageNumber="35">Fig. 161</figureCitation>
. Embolic keel vestige represented by very low micro-serrate edge formed by numerous uniformly-shaped denticles; subapical part of embolus corkscrew-shaped (
<figureCitation id="132B19DF56550C05529C8279FD9D4F81" box="[416,583,1898,1924]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="34" pageNumber="35">Figs 193–194</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56550C05538182BEFB794FEE" blockId="34.[189,1398,1964,2027]" pageId="34" pageNumber="35">SPINNERETS. PMS: length 0.35; diameter 0.18. PLS: maximal diameter 0.37; length of basal, medial and apical segments 0.78, 0.63, 0.60; total length 2.01; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A56540C045381841FFE354920" blockId="35.[189,495,267,294]" box="[189,495,267,294]" pageId="35" pageNumber="36">LEG MEASUREMENTS. ♂(♀).</paragraph>
<paragraph id="8BAF055A56540C0452568458FAB14A33" pageId="35" pageNumber="36">
<table id="F910F7FA5654F3D853F48459FAB14A33" box="[200,1387,330,566]" colsContinueFrom="24.[200,1341,913,1148]" colsContinueIn="37.[200,1339,463,698]" gridcols="7" gridrows="6" pageId="35" pageNumber="36">
<tr id="352007185654F3D853F48459FAB14960" box="[200,1387,330,357]" gridrow="0" pageId="35" pageNumber="36" rowspan-0="1">
<th id="76F16E645654F3D8526C8459FE0B4960" box="[336,465,330,357]" gridcol="1" gridrow="0" pageId="35" pageNumber="36">Femur</th>
<th id="76F16E645654F3D851398459FD5D4960" box="[517,647,330,357]" gridcol="2" gridrow="0" pageId="35" pageNumber="36">Patella</th>
<th id="76F16E645654F3D851878459FCE64960" box="[699,828,330,357]" gridcol="3" gridrow="0" pageId="35" pageNumber="36">Tibia</th>
<th id="76F16E645654F3D8504C8459FC284960" box="[880,1010,330,357]" gridcol="4" gridrow="0" pageId="35" pageNumber="36">Metatarsus</th>
<th id="76F16E645654F3D8571A8459FB7D4960" box="[1062,1191,330,357]" gridcol="5" gridrow="0" pageId="35" pageNumber="36">Tarsus</th>
<th id="76F16E645654F3D857F08459FAB14960" box="[1228,1387,330,357]" gridcol="6" gridrow="0" pageId="35" pageNumber="36">Total</th>
</tr>
<tr id="352007185654F3D853F48467FAB1498A" box="[200,1387,372,399]" gridrow="1" pageId="35" pageNumber="36">
<th id="76F16E645654F3D853F48467FEC6498A" box="[200,284,372,399]" gridcol="0" gridrow="1" pageId="35" pageNumber="36">Palp</th>
<td id="76F16E645654F3D8526C8467FE0B498A" box="[336,465,372,399]" gridcol="1" gridrow="1" pageId="35" pageNumber="36">3.10 (3.32)</td>
<td id="76F16E645654F3D851398467FD5D498A" box="[517,647,372,399]" gridcol="2" gridrow="1" pageId="35" pageNumber="36">1.27 (1.85)</td>
<td id="76F16E645654F3D851878467FCE6498A" box="[699,828,372,399]" gridcol="3" gridrow="1" pageId="35" pageNumber="36">2.05 (2.17)</td>
<td id="76F16E645654F3D8504C8467FC28498A" box="[880,1010,372,399]" gridcol="4" gridrow="1" pageId="35" pageNumber="36">–</td>
<td id="76F16E645654F3D8571A8467FB7D498A" box="[1062,1191,372,399]" gridcol="5" gridrow="1" pageId="35" pageNumber="36">0.99 (1.95)</td>
<td id="76F16E645654F3D857F08467FAB1498A" box="[1228,1387,372,399]" gridcol="6" gridrow="1" pageId="35" pageNumber="36">7.41 (9.29)</td>
</tr>
<tr id="352007185654F3D853F4848DFAB149BC" box="[200,1387,414,441]" gridrow="2" pageId="35" pageNumber="36">
<th id="76F16E645654F3D853F4848DFEC649BC" box="[200,284,414,441]" gridcol="0" gridrow="2" pageId="35" pageNumber="36">Leg I</th>
<td id="76F16E645654F3D8526C848DFE0B49BC" box="[336,465,414,441]" gridcol="1" gridrow="2" pageId="35" pageNumber="36">4.77 (4.10)</td>
<td id="76F16E645654F3D85139848DFD5D49BC" box="[517,647,414,441]" gridcol="2" gridrow="2" pageId="35" pageNumber="36">2.58 (2.47)</td>
<td id="76F16E645654F3D85187848DFCE649BC" box="[699,828,414,441]" gridcol="3" gridrow="2" pageId="35" pageNumber="36">3.62 (3.03)</td>
<td id="76F16E645654F3D8504C848DFC2849BC" box="[880,1010,414,441]" gridcol="4" gridrow="2" pageId="35" pageNumber="36">3.02 (2.30)</td>
<td id="76F16E645654F3D8571A848DFB7D49BC" box="[1062,1191,414,441]" gridcol="5" gridrow="2" pageId="35" pageNumber="36">1.90 (1.83)</td>
<td id="76F16E645654F3D857F0848DFAB149BC" box="[1228,1387,414,441]" gridcol="6" gridrow="2" pageId="35" pageNumber="36">15.89 (13.73)</td>
</tr>
<tr id="352007185654F3D853F484D4FAB149E7" box="[200,1387,455,482]" gridrow="3" pageId="35" pageNumber="36">
<th id="76F16E645654F3D853F484D4FEC649E7" box="[200,284,455,482]" gridcol="0" gridrow="3" pageId="35" pageNumber="36">Leg II</th>
<td id="76F16E645654F3D8526C84D4FE0B49E7" box="[336,465,455,482]" gridcol="1" gridrow="3" pageId="35" pageNumber="36">4.05 (3.65)</td>
<td id="76F16E645654F3D8513984D4FD5D49E7" box="[517,647,455,482]" gridcol="2" gridrow="3" pageId="35" pageNumber="36">2.27 (2.38)</td>
<td id="76F16E645654F3D8518784D4FCE649E7" box="[699,828,455,482]" gridcol="3" gridrow="3" pageId="35" pageNumber="36">2.97 (2.88)</td>
<td id="76F16E645654F3D8504C84D4FC2849E7" box="[880,1010,455,482]" gridcol="4" gridrow="3" pageId="35" pageNumber="36">2.98 (2.32)</td>
<td id="76F16E645654F3D8571A84D4FB7D49E7" box="[1062,1191,455,482]" gridcol="5" gridrow="3" pageId="35" pageNumber="36">1.83 (1.77)</td>
<td id="76F16E645654F3D857F084D4FAB149E7" box="[1228,1387,455,482]" gridcol="6" gridrow="3" pageId="35" pageNumber="36">14.10 (13.00)</td>
</tr>
<tr id="352007185654F3D853F484E2FAB14A09" box="[200,1387,497,524]" gridrow="4" pageId="35" pageNumber="36">
<th id="76F16E645654F3D853F484E2FEC64A09" box="[200,284,497,524]" gridcol="0" gridrow="4" pageId="35" pageNumber="36">Leg III</th>
<td id="76F16E645654F3D8526C84E2FE0B4A09" box="[336,465,497,524]" gridcol="1" gridrow="4" pageId="35" pageNumber="36">3.43 (3.17)</td>
<td id="76F16E645654F3D8513984E2FD5D4A09" box="[517,647,497,524]" gridcol="2" gridrow="4" pageId="35" pageNumber="36">1.83 (1.95)</td>
<td id="76F16E645654F3D8518784E2FCE64A09" box="[699,828,497,524]" gridcol="3" gridrow="4" pageId="35" pageNumber="36">2.32 (2.17)</td>
<td id="76F16E645654F3D8504C84E2FC284A09" box="[880,1010,497,524]" gridcol="4" gridrow="4" pageId="35" pageNumber="36">3.30 (2.75)</td>
<td id="76F16E645654F3D8571A84E2FB7D4A09" box="[1062,1191,497,524]" gridcol="5" gridrow="4" pageId="35" pageNumber="36">1.86 (1.83)</td>
<td id="76F16E645654F3D857F084E2FAB14A09" box="[1228,1387,497,524]" gridcol="6" gridrow="4" pageId="35" pageNumber="36">12.72 (11.87)</td>
</tr>
<tr id="352007185654F3D853F48708FAB14A33" box="[200,1387,539,566]" gridrow="5" pageId="35" pageNumber="36">
<th id="76F16E645654F3D853F48708FEC64A33" box="[200,284,539,566]" gridcol="0" gridrow="5" pageId="35" pageNumber="36">Leg IV</th>
<td id="76F16E645654F3D8526C8708FE0B4A33" box="[336,465,539,566]" gridcol="1" gridrow="5" pageId="35" pageNumber="36">4.62 (4.05)</td>
<td id="76F16E645654F3D851398708FD5D4A33" box="[517,647,539,566]" gridcol="2" gridrow="5" pageId="35" pageNumber="36">2.00 (2.37)</td>
<td id="76F16E645654F3D851878708FCE64A33" box="[699,828,539,566]" gridcol="3" gridrow="5" pageId="35" pageNumber="36">3.53 (3.25)</td>
<td id="76F16E645654F3D8504C8708FC284A33" box="[880,1010,539,566]" gridcol="4" gridrow="5" pageId="35" pageNumber="36">4.47 (3.97)</td>
<td id="76F16E645654F3D8571A8708FB7D4A33" box="[1062,1191,539,566]" gridcol="5" gridrow="5" pageId="35" pageNumber="36">2.12 (2.13)</td>
<td id="76F16E645654F3D857F08708FAB14A33" box="[1228,1387,539,566]" gridcol="6" gridrow="5" pageId="35" pageNumber="36">16.74 (15.77)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A56540C0453D68770FDC44A7B" blockId="35.[189,542,611,678]" box="[234,542,611,638]" pageId="35" pageNumber="36">
<emphasis id="B964D94856540C0453D68770FE9F4A78" bold="true" box="[234,325,611,637]" pageId="35" pageNumber="36">Female</emphasis>
(from Lagodekhi)
</paragraph>
<paragraph id="8BAF055A56540C045381879FFE6D4AA3" blockId="35.[189,542,611,678]" box="[189,439,651,678]" pageId="35" pageNumber="36">
HABITUS. See
<figureCitation id="132B19DF56540C0452618798FE6D4AA3" box="[349,439,651,678]" captionStart="Figs 52–58" captionStartId="58.[189,241,1833,1859]" captionTargetBox="[189,1398,265,1770]" captionTargetId="figure@58.[189,1398,265,1771]" captionText="Figs 52–58. Raveniola spp., habitus of ♂♂ (52–53, 55, 57) and ♀♀ (54, 56, 58), lateral (52) and dorsal (53–58) views. — 52. R. biresekensis sp. nov. (dwarf conspecific male?). 53–54. R. micropa (Ausserer, 1871). 55–56. R. zaizevi (Charitonov, 1948). 57–58. R. niedermeyeri (Brignoli, 1972). Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163440/files/figure.png" pageId="35" pageNumber="36">Fig. 56.</figureCitation>
</paragraph>
<paragraph id="8BAF055A56540C04538187C0FB7D4AE9" blockId="35.[189,1191,722,749]" box="[189,1191,722,749]" pageId="35" pageNumber="36">MEASUREMENTS. TBL 17.20, CL 6.22, CW 4.97, LL 0.57, LW 1.10, SL 2.87, SW 2.60.</paragraph>
<paragraph id="8BAF055A56540C045381860AFAFC4B36" blockId="35.[189,1318,793,820]" box="[189,1318,793,820]" pageId="35" pageNumber="36">COLOUR. As in male, but chelicerae slightly darker, cherry-red; clypeus lighter, yellowish brown.</paragraph>
<paragraph id="8BAF055A56540C0453818673FDA94C19" blockId="35.[189,1399,864,1053]" pageId="35" pageNumber="36">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF56540C04503E8673FC864B7F" box="[770,860,864,890]" captionStart="Figs 90–95" captionStartId="64.[189,241,1891,1917]" captionTargetBox="[189,1398,265,1846]" captionTargetId="figure@64.[189,1398,265,1849]" captionText="Figs 90–95. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 90. R. turcica sp. nov. (paratype). 91. R. micropa (Ausserer, 1871) (holotype). 92. Same species as preceding (Uludağ). 93. R. zaitzevi(Charitonov, 1948)(Lagodekhi).94.R.vonwickiZonstein, 2000(paratype).95.R.niedermeyeri (Brignoli, 1972) (paratype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163452/files/figure.png" pageId="35" pageNumber="36">Fig. 93</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF56540C0457EE8673FAE04B7F" box="[1234,1338,864,890]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="35" pageNumber="36">Fig. 126</figureCitation>
. Eye diameters and interdistances: AME 0.12(0.16), ALE 0.26, PLE 0.12, PME 0.10, AME–AME 0.17(0.13), ALE–AME 0.11(0.09), ALE–PLE 0.05, PLE–PME 0.08, PME–PME 0.40. Each cheliceral furrow with 10–11 promarginal teeth and 6–7 denticles. Maxillae with 9–12 cuspules each. Sternal sigilla small, oval and submarginal as shown in
<figureCitation id="132B19DF56540C0451248111FDA94C19" box="[536,627,1026,1052]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="35" pageNumber="36">Fig. 63.</figureCitation>
</paragraph>
<paragraph id="8BAF055A56540C0453818159FE704CD8" blockId="35.[189,1399,1097,1245]" pageId="35" pageNumber="36">LEGS. Scopula: distal on metatarsi I–II, entire on tarsus I and palpal tarsus, narrowly divided on tarsus II, elsewhere absent. Trichobothria: 2 rows of 9–10 each on tibiae, 11–15 on metatarsi, 10–13 on tarsi, 8 on palpal tarsus. Paired tarsal claws: 6–7 teeth on inner margin, 5–6 on outer margin. Palpal claw with 4 promarginal teeth.</paragraph>
<paragraph id="8BAF055A56540C0453818019FC964DEA" blockId="35.[189,1399,1290,1519]" pageId="35" pageNumber="36">SPINATION. All femora dorsally with one basodorsal spine and 3–6 bristles (undeveloped spines) located medially and distally; palpal patella and patellae I–II aspinose. Palp: femur pd1; tibia v2–2–3; tarsus v8(9). Leg I: femur pd1; tibia v2–2–3; metatarsus v2–2–1. Leg II: femur d1, pd1; tibia p1–1, v2–2– 3; metatarsus v2–2–2. Leg III: femur pd1–1, rd1–1; patella p1–1, r1; tibia d1, p1–1, r1–1, v2–2–3; metatarsus pd1–1, p1–1–1, r1–1, v2–2–2(3). Leg IV: femur pd 1(0), rd1; patella r1; tibia p1–1, r1(0)–1– 1, v2–2–3; metatarsus p1–1–1, r1–1–1–1, v2–2–2–2(3).</paragraph>
<paragraph id="8BAF055A56540C045381830FFAAD4E33" blockId="35.[189,1399,1563,1590]" box="[189,1399,1563,1590]" pageId="35" pageNumber="36">
SPERMATHECAE. Moderately narrow and relatively short, with large globular lateral receptacles (
<figureCitation id="132B19DF56540C0456388308FAB04E33" box="[1284,1386,1563,1590]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="35" pageNumber="36">Fig. 208</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56540C0453818371FAF64EA0" blockId="35.[189,1398,1634,1702]" pageId="35" pageNumber="36">
SPINNERETS. See
<figureCitation id="132B19DF56540C0452478371FE0B4E79" box="[379,465,1634,1660]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="35" pageNumber="36">Fig. 66</figureCitation>
. PMS: length 0.50; diameter 0.21. PLS: maximal diameter 0.60; length of basal, medial and apical segments 0.85, 0.53, 0.60; total length 1.98; apical segment shortly digitiform.
</paragraph>
<paragraph id="8BAF055A56540C04538183C1FEE94EE9" blockId="35.[189,307,1746,1772]" box="[189,307,1746,1772]" pageId="35" pageNumber="36">
<heading id="D0E7B23656540C04538183C1FEE94EE9" bold="true" box="[189,307,1746,1772]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B964D94856540C04538183C1FEE94EE9" bold="true" box="[189,307,1746,1772]" pageId="35" pageNumber="36">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56540C0453818215FC144FEE" blockId="35.[189,1399,1797,2027]" pageId="35" pageNumber="36">
Carapace length in males varies from 4.42 to 5.57, in females from 4.70 to 6.45. The dorsal abdominal pattern in specimens collected from eastern, less humid part of the geographical range, is weakly developed to indistinct. In some females, independent of the localities they were taken from, the posterior half of the carapace looks somewhat lighter in colour than the anterior one (as in specimens T.S. Mcheidze used to describe as
<taxonomicName id="4C107ED956540C04516B82BBFCFC4FC7" authorityName="Mcheidze" authorityYear="1983" box="[599,806,1960,1986]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="recki">
<emphasis id="B964D94856540C04516B82BBFCFC4FC7" box="[599,806,1960,1986]" italics="true" pageId="35" pageNumber="36">Brachythele recki</emphasis>
</taxonomicName>
). The structure of the spermathecae in all females examined shows a quite insignificant variation (see
<figureCitation id="132B19DF56540C04502082C2FC184FEE" box="[796,962,2001,2027]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="35" pageNumber="36">Figs 208–209</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156530C0353818418FDFD499B" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="8BAF055A56530C0353818418FEFA4920" blockId="36.[189,288,267,293]" box="[189,288,267,293]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C0353818418FEFA4920" bold="true" box="[189,288,267,293]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B964D94856530C0353818418FEFA4920" bold="true" box="[189,288,267,293]" pageId="36" pageNumber="37">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56530C0353818428FDFD499B" blockId="36.[189,1398,315,414]" pageId="36" pageNumber="37">
The spiders were found inhabiting soil cavities under stones and logs in broad-leaved forests of the low mountain zone and of the midlands. These forests are dominated by
<taxonomicName id="4C107ED956530C0350E88473FB2C497F" authority="M. Bieb." authorityName="M.Bieb." box="[980,1270,352,378]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="iberica">
<emphasis id="B964D94856530C0350E88473FB55497F" box="[980,1167,352,378]" italics="true" pageId="36" pageNumber="37">Quercus iberica</emphasis>
M.Bieb.
</taxonomicName>
and
<taxonomicName id="4C107ED956530C0356118472FE52499B" authority="Lipsky" authorityName="Lipsky" authorityYear="1898" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B964D94856530C0356118472FEF7499B" italics="true" pageId="36" pageNumber="37">Fagus orientalis</emphasis>
Lipsky
</taxonomicName>
, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156530C03538184D4FEFC4A33" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BAF055A56530C03538184D4FE8C49E4" blockId="36.[189,342,455,481]" box="[189,342,455,481]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C03538184D4FE8C49E4" bold="true" box="[189,342,455,481]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B964D94856530C03538184D4FE8C49E4" bold="true" box="[189,342,455,481]" pageId="36" pageNumber="37">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56530C03538184E4FEFC4A33" blockId="36.[189,1398,503,566]" pageId="36" pageNumber="37">
Southern slope of Caucasus Major east of 46° E (north-eastern Georgia, north-western Azerbaijan). See
<figureCitation id="132B19DF56530C035381870FFEF94A33" box="[189,291,540,566]" captionStart="Figure 220" captionStartId="86.[189,272,1534,1560]" captionTargetBox="[190,1398,265,1471]" captionTargetId="figure@86.[189,1398,265,1471]" captionText="Figure 220. Localities of Raveniola spp. in the eastern part of the Caucasus: R. dunini sp. nov. (), R. hyrcanica Dunin, 1988 () and R. zaitzevi (Charitonov, 1948) ().Abbreviation:TU = Turkey." httpUri="https://zenodo.org/record/1163496/files/figure.png" pageId="36" pageNumber="37">Fig. 220</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156530C035381874CFD224B39" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BAF055A56530C035381874CFED94A7C" blockId="36.[189,259,607,633]" box="[189,259,607,633]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C035381874CFED94A7C" bold="true" box="[189,259,607,633]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B964D94856530C035381874CFED94A7C" bold="true" box="[189,259,607,633]" pageId="36" pageNumber="37">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56530C035381879CFD224B39" blockId="36.[189,1399,655,829]" pageId="36" pageNumber="37">
<bibRefCitation id="EF8178AB56530C035381879CFE504AAF" author="Mcheidze T. S." box="[189,394,655,682]" editor="Eliashvili T." journalOrPublisher="Metzniereba Publishing House, Tbilisi. [In Russian]" pageId="36" pageNumber="37" pagination="155 - 159" refString="Mcheidze T. S. 1983. A new spider species, Brachythele recki sp. nov., from Georgia. In: Eliashvili T. (ed.) Fauna i Ekologia Bespozvonochnykh Gruzii: 155 - 159. Metzniereba Publishing House, Tbilisi. [In Russian]" title="A new spider species, Brachythele recki sp. nov., from Georgia" type="book chapter" volumeTitle="Fauna i Ekologia Bespozvonochnykh Gruzii" year="1983">Mcheidze (1983)</bibRefCitation>
considered some differences in the colouration of the carapace, in the tarsal claw dentition, and in the number of cuspules sufficient to consider two females from Lagodekhi, Georgia as representatives of a distinct species,
<taxonomicName id="4C107ED956530C0351B387CAFC844AF6" authorityName="Mcheidze" authorityYear="1983" box="[655,862,729,755]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="recki">
<emphasis id="B964D94856530C0351B387CAFC844AF6" box="[655,862,729,755]" italics="true" pageId="36" pageNumber="37">Brachythele recki</emphasis>
</taxonomicName>
. Having examined the available material, we found that all these differences lay within the limits of intraspecific variability. We thus regard
<taxonomicName id="4C107ED956530C03562087EDFAAD4B1D" authorityName="Mcheidze" authorityYear="1983" box="[1308,1399,766,792]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="recki">
<emphasis id="B964D94856530C03562087EDFAAD4B1D" box="[1308,1399,766,792]" italics="true" pageId="36" pageNumber="37">B. recki</emphasis>
</taxonomicName>
to be a synonym of
<taxonomicName id="4C107ED956530C0352998630FD2E4B39" authority="Charitonov, 1948" authorityName="Charitonov" authorityYear="1948" box="[421,756,802,828]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856530C0352998630FDC44B39" box="[421,542,803,828]" italics="true" pageId="36" pageNumber="37">B. zaitzevi</emphasis>
<bibRefCitation id="EF8178AB56530C0351198631FD2E4B39" author="Charitonov D. E." box="[549,756,802,828]" journalOrPublisher="Soobshcheniya Akademii Nauk Gruzinskoi SSR" pageId="36" pageNumber="37" pagination="135 - 139" part="9" refString="Charitonov D. E. 1948. A new species of Brachythele from Georgia. Soobshcheniya Akademii Nauk Gruzinskoi SSR 9: 135 - 139. [In Russian and Latin]" title="A new species of Brachythele from Georgia" type="journal article" year="1948">Charitonov, 1948</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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