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<document id="D1DBE255D58A2E69DD76065E77E92AA1" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56530C02514E866FFD834E0C" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola marusiki Zonstein &amp; Kunt &amp; Yağmur 2018, sp. nov." docType="treatment" docUuid="C48F164E-60BD-4454-A24D-FC7459D30BDF" docUuidSource="ZooBank" docVersion="14" lastPageNumber="38" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="37" updateTime="1698518020641" updateUser="ExternalLinkService">
<mods:mods id="2011400A6E25CE8C104E5010909B174F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="532EEEC37847EE85B89AA7921A7ACCA3">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:name id="33BEA80970BF73D888EDCD7FDBE37FC3" type="personal">
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<mods:namePart id="8E2D99D741F2A4A44E45D8A26B206978">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="7C4F85E9271BC9CF805B3AB0C5D6B157">Kunt, Kadir B.</mods:namePart>
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<mods:namePart id="2144B2B3AC9D771AA2AC3764D518BBEC">Yağmur, Ersen A.</mods:namePart>
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<mods:title id="0B831F758B0E70461C9044DE060DE7C1">European Journal of Taxonomy</mods:title>
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<mods:part id="2F4A677AB999B413A390EF86992672A8">
<mods:date id="67DF947D2A69C139291942FDAF6A8797">2018</mods:date>
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<mods:number id="6BF19A5C077E06EA112F093572E87E93">2018-01-30</mods:number>
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<mods:number id="B5FB236DB6200CC5DB2D6F77D11BFA0C">399</mods:number>
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<mods:identifier id="582CC9318439726A8F0BF2CC3B4EFD0A" type="DOI">10.5852/ejt.2018.399</mods:identifier>
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<treatment id="03B9B44C56530C02514E866FFD834E0C" ID-DOI="http://doi.org/10.5281/zenodo.5980306" ID-GBIF-Taxon="140561351" ID-Zenodo-Dep="5980306" LSID="urn:lsid:zoobank.org:act:C48F164E-60BD-4454-A24D-FC7459D30BDF" httpUri="http://treatment.plazi.org/id/03B9B44C56530C02514E866FFD834E0C" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection id="C30A56D156530C03514E866FFB124BBE" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BAF055A56530C03514E866FFC1B4B92" blockId="36.[363,1224,892,992]" box="[626,961,892,919]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C03514E866FFC1B4B92" box="[626,961,892,919]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
<taxonomicName id="4C107ED956530C03514E866FFCBA4B92" ID-CoL="4RMTY" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[626,864,892,919]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
<emphasis id="B964D94856530C03514E866FFCBA4B92" bold="true" box="[626,864,892,919]" italics="true" pageId="36" pageNumber="37">Raveniola marusiki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356530C03505B866EFC1B4B92" box="[871,961,893,919]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BAF055A56530C03525786B3FB124BBE" blockId="36.[363,1224,892,992]" box="[363,1224,928,955]" pageId="36" pageNumber="37">urn:lsid:zoobank.org:act:C48F164E-60BD-4454-A24D-FC7459D30BDF</paragraph>
</subSubSection>
<subSubSection id="C30A56D156530C03516086D6FC0D4BE5" box="[604,983,965,992]" pageId="36" pageNumber="37" type="description">
<paragraph id="8BAF055A56530C03516086D6FC0D4BE5" blockId="36.[363,1224,892,992]" box="[604,983,965,992]" pageId="36" pageNumber="37">
<figureCitation id="132B19DF56530C03516086D6FD6F4BE5" box="[604,693,965,992]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="36" pageNumber="37">Figs 81</figureCitation>
,
<figureCitation id="132B19DF56530C0351FE86D5FD2A4BE5" box="[706,752,966,992]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="36" pageNumber="37">114</figureCitation>
,
<figureCitation id="132B19DF56530C0351C786D5FCF04BE5" box="[763,810,966,992]" captionStart="Figs 141–144" captionStartId="71.[189,241,1858,1884]" captionTargetBox="[239,1352,266,1791]" captionTargetId="figure@71.[206,1378,265,1793]" captionText="Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163466/files/figure.png" pageId="36" pageNumber="37">142</figureCitation>
,
<figureCitation id="132B19DF56530C03500A86D6FCBF4BDA" box="[822,869,965,991]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="36" pageNumber="37">162</figureCitation>
,
<figureCitation id="132B19DF56530C03504D86D5FC0D4BE5" box="[881,983,966,992]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="36" pageNumber="37">191–192</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156530C035381811BFC0A4C99" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BAF055A56530C035381811BFEEC4C27" blockId="36.[189,310,1032,1058]" box="[189,310,1032,1058]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C035381811BFEEC4C27" bold="true" box="[189,310,1032,1058]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B964D94856530C035381811BFEEC4C27" bold="true" box="[189,310,1032,1058]" pageId="36" pageNumber="37">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56530C035381812AFC0A4C99" blockId="36.[189,1398,1080,1181]" pageId="36" pageNumber="37">
In the shape of the distally spiraled embolus, the holotype of
<taxonomicName id="4C107ED956530C035094812AFB4A4C56" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[936,1168,1081,1107]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
<emphasis id="B964D94856530C035094812AFB4A4C56" box="[936,1168,1081,1107]" italics="true" pageId="36" pageNumber="37">Raveniola marusiki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356530C0357A6812AFB2D4C56" box="[1178,1271,1081,1107]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
resembles males of
<taxonomicName id="4C107ED956530C035210814DFE6D4C72" authorityName="Zonstein" authorityYear="2000" box="[300,439,1117,1143]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="vonwicki">
<emphasis id="B964D94856530C035210814DFE6D4C72" box="[300,439,1117,1143]" italics="true" pageId="36" pageNumber="37">R. vonwicki</emphasis>
</taxonomicName>
and
<taxonomicName id="4C107ED956530C0352C8814DFDB54C72" authorityName="Charitonov" authorityYear="1948" box="[500,623,1118,1143]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856530C0352C8814DFDB54C72" box="[500,623,1118,1143]" italics="true" pageId="36" pageNumber="37">R. zaitzevi</emphasis>
</taxonomicName>
, but differ from them, as well as from other species of the same group, by the unique shape of its pectinate embolic keel (
<figureCitation id="132B19DF56530C0350608191FC194C99" box="[860,963,1154,1180]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="36" pageNumber="37">Fig. 191</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156530C03538181D6FC734D31" pageId="36" pageNumber="37" type="etymology">
<paragraph id="8BAF055A56530C03538181D6FE9E4CDA" blockId="36.[189,324,1221,1247]" box="[189,324,1221,1247]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C03538181D6FE9E4CDA" bold="true" box="[189,324,1221,1247]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B964D94856530C03538181D6FE9E4CDA" bold="true" box="[189,324,1221,1247]" pageId="36" pageNumber="37">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56530C03538181E6FC734D31" blockId="36.[189,1398,1269,1332]" pageId="36" pageNumber="37">The specific epithet is given in honour of our friend and colleague Yuri Marusik for his immense contribution to the study of the spider fauna of Turkey and Iran.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156530C035381804EFC044DF4" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BAF055A56530C035381804EFE704D72" blockId="36.[189,426,1373,1399]" box="[189,426,1373,1399]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C035381804EFE704D72" bold="true" box="[189,426,1373,1399]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B964D94856530C035381804EFE704D72" bold="true" box="[189,426,1373,1399]" pageId="36" pageNumber="37">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B780F0756530C0353D6809EFC044DF4" ID-GBIF-Occurrence="1806244353" collectingDate="1903-10-07" collectingDateMax="1903-10-09" collectingDateMin="1903-10-07" collectionCode="ZMMU" collectorName="N. A. Zarudny" country="Iran" elevation="750" latitude="36.8" location="Elburs Mts, Germabdasht Valley 15 – 20 km SE of Gorgan" longLatPrecision="1185" longitude="54.566666" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Golestan Province" typeStatus="holotype">
<paragraph id="8BAF055A56530C0353D6809EFE864DA2" blockId="36.[189,1399,1421,1521]" box="[234,348,1421,1447]" pageId="36" pageNumber="37">
<typeStatus id="54ABBBF856530C0353D6809EFE864DA2" box="[234,348,1421,1447]" pageId="36" pageNumber="37" type="holotype">
<emphasis id="B964D94856530C0353D6809EFE864DA2" bold="true" box="[234,348,1421,1447]" pageId="36" pageNumber="37">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BAF055A56530C03538180A1FC044DF4" blockId="36.[189,1399,1421,1521]" pageId="36" pageNumber="37">
<collectingCountry id="F30745CA56530C03538180A1FED34DC9" box="[189,265,1458,1484]" name="Iran" pageId="36" pageNumber="37">IRAN</collectingCountry>
:
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,
<collectingRegion id="49D4CBB856530C03527C80A1FDC54DC9" box="[320,543,1458,1484]" country="Iran" name="Golestan" pageId="36" pageNumber="37">Golestan Province</collectingRegion>
,
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Elburs Mts, Germabdasht Valley
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SE of Gorgan
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,
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,
<geoCoordinate id="EE24639D56530C03538180C6FEFF4DF4" box="[189,293,1493,1521]" direction="east" orientation="longitude" pageId="36" pageNumber="37" precision="925" value="54.566666">54°34′ E</geoCoordinate>
,
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<quantity id="4CE8A8BF56530C03520E80C5FE1D4DF4" box="[306,455,1494,1521]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="36" pageNumber="37" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">500–1000 m</quantity>
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,
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</collectingDate>
,
<collectorName id="26E5608C56530C0351B680C4FCF74DF5" box="[650,813,1494,1521]" pageId="36" pageNumber="37">N.A. Zarudny</collectorName>
leg. (
<collectionCode id="ED019D9F56530C03505380C4FC0B4DF4" box="[879,977,1495,1521]" collectionName="ZMMU" pageId="36" pageNumber="37">ZMMU</collectionCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C30A56D156530C025381830AFEE34D06" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="8BAF055A56530C035381830AFE944E36" blockId="36.[189,334,1561,1587]" box="[189,334,1561,1587]" pageId="36" pageNumber="37">
<heading id="D0E7B23656530C035381830AFE944E36" bold="true" box="[189,334,1561,1587]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B964D94856530C035381830AFE944E36" bold="true" box="[189,334,1561,1587]" pageId="36" pageNumber="37">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56530C0353D68359FE714E61" blockId="36.[189,1172,1609,1673]" box="[234,427,1609,1636]" pageId="36" pageNumber="37">
<emphasis id="B964D94856530C0353D68359FEF04E61" bold="true" box="[234,298,1610,1636]" pageId="36" pageNumber="37">Male</emphasis>
(holotype)
</paragraph>
<paragraph id="8BAF055A56530C035381837CFB4E4E8C" blockId="36.[189,1172,1609,1673]" box="[189,1172,1646,1673]" pageId="36" pageNumber="37">MEASUREMENTS. TL 14.50, CL 5.64, CW 4.96, LL 0.49, LW 0.97, SL 2.58, SW 2.53.</paragraph>
<paragraph id="8BAF055A56530C03538183A2FD434F3C" blockId="36.[189,1399,1713,1849]" pageId="36" pageNumber="37">COLOUR. Carapace, palps and legs dorsally brownish rufous; eye tubercle anteriorly brownish black, chelicerae reddish brown; sternum, labium, maxillae, palps and legs ventrally yellowish brown; abdomen dorsally brown, with darker pattern consisting of partially fused dark brown irregular spots; ventral surface of abdomen light greyish brown.</paragraph>
<paragraph id="8BAF055A56530C0353818270FAC04FEE" blockId="36.[189,1399,1890,2027]" pageId="36" pageNumber="37">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF56530C03503E8271FC814F78" box="[770,859,1890,1917]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="36" pageNumber="37">Fig. 81</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF56530C0357EF8271FAE04F78" box="[1235,1338,1890,1917]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="36" pageNumber="37">Fig. 114</figureCitation>
. Eye diameters and interdistances: AME 0.15(0.19), ALE 0.25, PLE 0.18, PME 0.13, AME–AME 0.10(0.05), ALE–AME 0.08(0.06), ALE–PLE 0.11, PLE–PME 0.04, PME–PME 0.37. Each cheliceral furrow with 9 promarginal teeth and 6 mesobasal denticles. Maxillae: each with 9 relatively small cuspules.
</paragraph>
<paragraph id="8BAF055A56520C025381841FFC334968" blockId="37.[189,1398,267,365]" pageId="37" pageNumber="38">
LEGS. Tibia and metatarsus I as shown in
<figureCitation id="132B19DF56520C0251928418FCCE4920" box="[686,788,267,293]" captionStart="Figs 141–144" captionStartId="71.[189,241,1858,1884]" captionTargetBox="[239,1352,266,1791]" captionTargetId="figure@71.[206,1378,265,1793]" captionText="Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163466/files/figure.png" pageId="37" pageNumber="38">Fig. 142</figureCitation>
. Scopula: distal on metatarsi I–II, entire on tarsi I– II; absent on tarsi III–IV. Trichobothria: 2 rows of 9–10 each on tibiae, 10–14 on metatarsi, 10–12 on tarsi, 7–8 on cymbium. Paired claws with 8–11 teeth on each margin.
</paragraph>
<paragraph id="8BAF055A56520C0253818486FE7A49AA" blockId="37.[189,416,405,431]" box="[189,416,405,431]" pageId="37" pageNumber="38">LEG MEASUREMENTS.</paragraph>
<paragraph id="8BAF055A56520C02525484DCFAE14ABC" pageId="37" pageNumber="38">
<table id="F910F7FA5652F3D853F484DCFAE14ABF" box="[200,1339,463,698]" colsContinueFrom="35.[200,1387,330,566]" colsContinueIn="38.[200,1383,1781,2016]" gridcols="7" gridrows="6" pageId="37" pageNumber="38">
<tr id="352007185652F3D853F484DCFAE149EC" box="[200,1339,463,489]" gridrow="0" pageId="37" pageNumber="38" rowspan-0="1">
<th id="76F16E645652F3D8525484DCFE6F49EC" box="[360,437,463,489]" gridcol="1" gridrow="0" pageId="37" pageNumber="38">Femur</th>
<th id="76F16E645652F3D8512084DCFDB649EC" box="[540,620,463,489]" gridcol="2" gridrow="0" pageId="37" pageNumber="38">Patella</th>
<th id="76F16E645652F3D851E784DCFCC249EC" box="[731,792,463,489]" gridcol="3" gridrow="0" pageId="37" pageNumber="38">Tibia</th>
<th id="76F16E645652F3D8505384DCFC2A49EC" box="[879,1008,463,489]" gridcol="4" gridrow="0" pageId="37" pageNumber="38">Metatarsus</th>
<th id="76F16E645652F3D8570284DCFB5049EC" box="[1086,1162,463,489]" gridcol="5" gridrow="0" pageId="37" pageNumber="38">Tarsus</th>
<th id="76F16E645652F3D857C584DCFAE149EC" box="[1273,1339,463,489]" gridcol="6" gridrow="0" pageId="37" pageNumber="38">Total</th>
</tr>
<tr id="352007185652F3D853F484EBFAE14A16" box="[200,1339,504,531]" gridrow="1" pageId="37" pageNumber="38">
<th id="76F16E645652F3D853F484EBFF264A16" box="[200,252,504,531]" gridcol="0" gridrow="1" pageId="37" pageNumber="38">Palp</th>
<td id="76F16E645652F3D8525484EBFE6F4A16" box="[360,437,504,531]" gridcol="1" gridrow="1" pageId="37" pageNumber="38">3.01</td>
<td id="76F16E645652F3D8512084EBFDB64A16" box="[540,620,504,531]" gridcol="2" gridrow="1" pageId="37" pageNumber="38">1.72</td>
<td id="76F16E645652F3D851E784EBFCC24A16" box="[731,792,504,531]" gridcol="3" gridrow="1" pageId="37" pageNumber="38">2.37</td>
<td id="76F16E645652F3D8505384EBFC2A4A16" box="[879,1008,504,531]" gridcol="4" gridrow="1" pageId="37" pageNumber="38">–</td>
<td id="76F16E645652F3D8570284EBFB504A16" box="[1086,1162,504,531]" gridcol="5" gridrow="1" pageId="37" pageNumber="38">0.95</td>
<td id="76F16E645652F3D857C584EBFAE14A16" box="[1273,1339,504,531]" gridcol="6" gridrow="1" pageId="37" pageNumber="38">8.05</td>
</tr>
<tr id="352007185652F3D853F48731FAE14A38" box="[200,1339,546,573]" gridrow="2" pageId="37" pageNumber="38">
<th id="76F16E645652F3D853F48731FF264A38" box="[200,252,546,573]" gridcol="0" gridrow="2" pageId="37" pageNumber="38">I</th>
<td id="76F16E645652F3D852548731FE6F4A38" box="[360,437,546,573]" gridcol="1" gridrow="2" pageId="37" pageNumber="38">4.59</td>
<td id="76F16E645652F3D851208731FDB64A38" box="[540,620,546,573]" gridcol="2" gridrow="2" pageId="37" pageNumber="38">2.76</td>
<td id="76F16E645652F3D851E78731FCC24A38" box="[731,792,546,573]" gridcol="3" gridrow="2" pageId="37" pageNumber="38">3.67</td>
<td id="76F16E645652F3D850538731FC2A4A38" box="[879,1008,546,573]" gridcol="4" gridrow="2" pageId="37" pageNumber="38">3.36</td>
<td id="76F16E645652F3D857028731FB504A38" box="[1086,1162,546,573]" gridcol="5" gridrow="2" pageId="37" pageNumber="38">2.18</td>
<td id="76F16E645652F3D857C58731FAE14A38" box="[1273,1339,546,573]" gridcol="6" gridrow="2" pageId="37" pageNumber="38">16.56</td>
</tr>
<tr id="352007185652F3D853F4875FFAE14A63" box="[200,1339,588,614]" gridrow="3" pageId="37" pageNumber="38">
<th id="76F16E645652F3D853F4875FFF264A63" box="[200,252,588,614]" gridcol="0" gridrow="3" pageId="37" pageNumber="38">II</th>
<td id="76F16E645652F3D85254875FFE6F4A63" box="[360,437,588,614]" gridcol="1" gridrow="3" pageId="37" pageNumber="38">3.90</td>
<td id="76F16E645652F3D85120875FFDB64A63" box="[540,620,588,614]" gridcol="2" gridrow="3" pageId="37" pageNumber="38">2.14</td>
<td id="76F16E645652F3D851E7875FFCC24A63" box="[731,792,588,614]" gridcol="3" gridrow="3" pageId="37" pageNumber="38">3.03</td>
<td id="76F16E645652F3D85053875FFC2A4A63" box="[879,1008,588,614]" gridcol="4" gridrow="3" pageId="37" pageNumber="38">3.24</td>
<td id="76F16E645652F3D85702875FFB504A63" box="[1086,1162,588,614]" gridcol="5" gridrow="3" pageId="37" pageNumber="38">2.00</td>
<td id="76F16E645652F3D857C5875FFAE14A63" box="[1273,1339,588,614]" gridcol="6" gridrow="3" pageId="37" pageNumber="38">14.31</td>
</tr>
<tr id="352007185652F3D853F48766FAE14A95" box="[200,1339,629,656]" gridrow="4" pageId="37" pageNumber="38">
<th id="76F16E645652F3D853F48766FF264A95" box="[200,252,629,656]" gridcol="0" gridrow="4" pageId="37" pageNumber="38">III</th>
<td id="76F16E645652F3D852548766FE6F4A95" box="[360,437,629,656]" gridcol="1" gridrow="4" pageId="37" pageNumber="38">3.76</td>
<td id="76F16E645652F3D851208766FDB64A95" box="[540,620,629,656]" gridcol="2" gridrow="4" pageId="37" pageNumber="38">1.88</td>
<td id="76F16E645652F3D851E78766FCC24A95" box="[731,792,629,656]" gridcol="3" gridrow="4" pageId="37" pageNumber="38">2.51</td>
<td id="76F16E645652F3D850538766FC2A4A95" box="[879,1008,629,656]" gridcol="4" gridrow="4" pageId="37" pageNumber="38">3.57</td>
<td id="76F16E645652F3D857028766FB504A95" box="[1086,1162,629,656]" gridcol="5" gridrow="4" pageId="37" pageNumber="38">2.01</td>
<td id="76F16E645652F3D857C58766FAE14A95" box="[1273,1339,629,656]" gridcol="6" gridrow="4" pageId="37" pageNumber="38">13.73</td>
</tr>
<tr id="352007185652F3D853F4878CFAE14ABF" box="[200,1339,671,698]" gridrow="5" pageId="37" pageNumber="38">
<th id="76F16E645652F3D853F4878CFF264ABF" box="[200,252,671,698]" gridcol="0" gridrow="5" pageId="37" pageNumber="38">IV</th>
<td id="76F16E645652F3D85254878CFE6F4ABF" box="[360,437,671,698]" gridcol="1" gridrow="5" pageId="37" pageNumber="38">4.75</td>
<td id="76F16E645652F3D85120878CFDB64ABF" box="[540,620,671,698]" gridcol="2" gridrow="5" pageId="37" pageNumber="38">2.25</td>
<td id="76F16E645652F3D851E7878CFCC24ABF" box="[731,792,671,698]" gridcol="3" gridrow="5" pageId="37" pageNumber="38">3.70</td>
<td id="76F16E645652F3D85053878CFC2A4ABF" box="[879,1008,671,698]" gridcol="4" gridrow="5" pageId="37" pageNumber="38">5.12</td>
<td id="76F16E645652F3D85702878CFB504ABF" box="[1086,1162,671,698]" gridcol="5" gridrow="5" pageId="37" pageNumber="38">2.28</td>
<td id="76F16E645652F3D857C5878CFAE14ABF" box="[1273,1339,671,698]" gridcol="6" gridrow="5" pageId="37" pageNumber="38">18.10</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A56520C02538187F1FAAD4BAA" blockId="37.[189,1399,738,943]" pageId="37" pageNumber="38">SPINATION. Patella I aspinose. Palp: femur d0–1–1–1, pd1; patella p1; tibia d1–1–0, p2–2–2, r1–1, pv0–0–1, rv1–1–0; cymbium d6. Leg I: femur d1–1–1–1, pd1–1–1, rd0–0–1; tibia p1–1–0, pv v1–0–1–0; rv0–1–0– m–m; metatarsus v0–0–2. Leg II: femur d1–1–1–1, pd0–1–1; tibia p1–1–1, v2–2–2; metatarsus p0–1–0; v2–2–2. Leg III: femur d1–1–1–1, pd1–1–1, rd1–1–1; patella p1–1(0), r1; tibia d1–1–0, p1–1–1, r1–1– 1, v2–2–3; metatarsus d1–0–1, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur d1–1–1–1, pd1–1–1, rd0–0–1; patella r1; tibia d1–1–0, p1–1–1, r1–1–1, v2–2–3; metatarsus d1–0–1–0, p2–2–1–1, r1–1–1–1, v2–2–2–3.</paragraph>
<paragraph id="8BAF055A56520C02538186C4FDD54C3D" blockId="37.[189,1399,983,1080]" pageId="37" pageNumber="38">
PALP. Tibia, cymbium and palpal organ as shown in
<figureCitation id="132B19DF56520C02500386C4FC7D4BF4" box="[831,935,983,1009]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="37" pageNumber="38">Fig. 162</figureCitation>
. Short pectinate embolic keel consists of two large and a few smaller teeth; embolus gently corkscrew-shaped, with proximal bend located submedially (
<figureCitation id="132B19DF56520C025260810DFDD84C3D" box="[348,514,1054,1080]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="37" pageNumber="38">Figs 191–192</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56520C0253818173FB794C9B" blockId="37.[189,1398,1120,1182]" pageId="37" pageNumber="38">SPINNERETS. PMS: length 0.48; diameter 0.21. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.71, 0.55, 0.58; total length 1.84; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A56520C0253D681D5FE9F4CE5" blockId="37.[189,325,1222,1283]" box="[234,325,1222,1248]" pageId="37" pageNumber="38">
<emphasis id="B964D94856520C0253D681D5FE9F4CE5" bold="true" box="[234,325,1222,1248]" pageId="37" pageNumber="38">Female</emphasis>
</paragraph>
<paragraph id="8BAF055A56520C02538181FAFEE34D06" blockId="37.[189,325,1222,1283]" box="[189,313,1257,1283]" pageId="37" pageNumber="38">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156520C0253818038FDE24D9D" pageId="37" pageNumber="38" type="biology_ecology">
<paragraph id="8BAF055A56520C0253818038FEFA4D40" blockId="37.[189,288,1323,1349]" box="[189,288,1323,1349]" pageId="37" pageNumber="38">
<heading id="D0E7B23656520C0253818038FEFA4D40" bold="true" box="[189,288,1323,1349]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B964D94856520C0253818038FEFA4D40" bold="true" box="[189,288,1323,1349]" pageId="37" pageNumber="38">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56520C0253818049FDE24D9D" blockId="37.[189,1398,1370,1433]" pageId="37" pageNumber="38">Judging from the label data, the holotypic male was collected in the broad-leaved forest zone of Alborz Mts. Other details are unknown.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156520C02538180D3FD834E0C" box="[189,601,1472,1546]" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BAF055A56520C02538180D3FE8C4DDF" blockId="37.[189,342,1472,1498]" box="[189,342,1472,1498]" pageId="37" pageNumber="38">
<heading id="D0E7B23656520C02538180D3FE8C4DDF" bold="true" box="[189,342,1472,1498]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B964D94856520C02538180D3FE8C4DDF" bold="true" box="[189,342,1472,1498]" pageId="37" pageNumber="38">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56520C02538180E3FD834E0C" blockId="37.[189,601,1519,1546]" box="[189,601,1519,1546]" pageId="37" pageNumber="38">Known only from the type locality.</paragraph>
</subSubSection>
</treatment>
</document>
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