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<mods:title id="68DECF22412358D0870BC0D39A1B444B">Occurrence of the milliped Sinocallipus simplipodicus Zhang, 1993 in Laos, with reviews of the Southeast Asian and global callipodidan faunas, and remarks on the phylogenetic position of the order (Callipodida: Sinocallipodidea: Sinocallipodidae).</mods:title>
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<paragraph id="8BB920469D502C2BFE89F8B8FCE40546" blockId="3.[264,776,1804,1830]" box="[264,776,1804,1830]" pageId="3" pageNumber="4">
<heading id="D0F1972A9D502C2BFE89F8B8FCE40546" bold="true" box="[264,776,1804,1830]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B972FC549D502C2BFE89F8B8FCE40546" bold="true" box="[264,776,1804,1830]" pageId="3" pageNumber="4">
The Order
<taxonomicName id="4C065BC59D502C2BFE14F8B8FDC40546" ID-CoL="7NF8C" box="[405,552,1804,1830]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
in Southeast Asia
</emphasis>
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</paragraph>
<paragraph id="8BB920469D502C2CFE89F8E3FD430016" blockId="3.[264,1322,1879,1905]" lastBlockId="4.[264,1324,284,1510]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Prior to 1981, the easternmost record of the
<taxonomicName id="4C065BC59D502C2BFC94F8E3FC720511" box="[789,926,1879,1905]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
in Eurasia was that of
<taxonomicName id="4C065BC59D502C2CFB31F8E3FDDD0356" authority="Attems, 1936" authorityName="Attems" authorityYear="1936" class="Diplopoda" family="Caspiopetalidae" genus="Bollmania" kingdom="Animalia" lastPageId="4" lastPageNumber="5" order="Callipodida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kohalana">
<emphasis id="B972FC549D502C2CFB31F8E3FE990355" italics="true" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">Bollmania kohalana</emphasis>
(Attems, 1936)
</taxonomicName>
, from Kohala, Punjab Province,
<collectingCountry id="F31160D69D572C2CFC2FFEA8FBFD0356" box="[942,1041,284,310]" name="Pakistan" pageId="4" pageNumber="5">Pakistan</collectingCountry>
(Attems, 1936). In that year, Golovatch (1981) discovered two unidentifiable females from Jiangsu (=Kiangsu) Province,
<collectingCountry id="F31160D69D572C2CFEFFFED8FE2B03E6" box="[382,455,364,390]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
, in the collection at the Institute of Zoology, Polish Academy of Sciences, Warsaw. He assigned them to
<taxonomicName id="4C065BC59D572C2CFDF7FE20FD1C03CD" box="[630,752,404,429]" class="Diplopoda" family="Caspiopetalidae" genus="Bollmania" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFDF7FE20FD1C03CD" box="[630,752,404,429]" italics="true" pageId="4" pageNumber="5">Bollmania</emphasis>
</taxonomicName>
, then the most geographically proximate genus, although the
<collectingCountry id="F31160D69D572C2CFE21FE08FDE803B6" box="[416,516,444,470]" name="Pakistan" pageId="4" pageNumber="5">Pakistan</collectingCountry>
locality is on the other (south) side of the Himalayas and some
<quantity id="4CFE8DA39D572C2CFB68FE08FECB039E" metricMagnitude="3" metricUnit="m" metricValue="4.1038272" pageId="4" pageNumber="5" unit="mi" value="2.55">2,550 mi</quantity>
(
<quantity id="4CFE8DA39D572C2CFEB9FE50FE41039E" box="[312,429,484,510]" metricMagnitude="3" metricUnit="m" metricValue="4.08" pageId="4" pageNumber="5" unit="km" value="4.08">4,080 km</quantity>
) west of the Chinese site; he (Golovatch, 1983, 1991) also referenced this record. We think it unlikely that this genus and family occurs in eastern
<collectingCountry id="F31160D69D572C2CFBD0FDB8FB7B0046" box="[1105,1175,524,550]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
not far from Shanghai, so we assign this record to the most common Chinese family,
<taxonomicName id="4C065BC59D572C2CFBDDFD80FAE4002E" box="[1116,1288,564,590]" class="Diplopoda" family="Paracortinidae" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Paracortinidae</taxonomicName>
, as an unidentifiable genus and species.
</paragraph>
<paragraph id="8BB920469D572C2CFEB9FD30FD5706C6" blockId="4.[264,1324,284,1510]" pageId="4" pageNumber="5">
Zhang (1993) was the first to propose a new taxon for Chinese callipodidans when he erected the family
<taxonomicName id="4C065BC59D572C2CFE6CFD18FD5E00A6" box="[493,690,684,710]" class="Diplopoda" family="Sinocallipodidae" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Sinocallipodidae</taxonomicName>
for
<taxonomicName id="4C065BC59D572C2CFD68FD18FBC400A5" box="[745,1064,684,709]" class="Diplopoda" family="Sinocallipodidae" genus="Sinocallipus" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="simplipodicus" status="gen. nov.">
<emphasis id="B972FC549D572C2CFD68FD18FBC400A5" box="[745,1064,684,709]" italics="true" pageId="4" pageNumber="5">Sinocallipus simplipodicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A241412F9D572C2CFBB8FD18FB6200A6" box="[1081,1166,684,710]" pageId="4" pageNumber="5" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp., from Xian Cave, Yunnan Province. Later that year, Wang and Zhang (1993) established the family
<taxonomicName id="4C065BC59D572C2CFEDEFD48FDE40176" box="[351,520,764,790]" class="Diplopoda" family="Paracortinidae" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Paracortinidae</taxonomicName>
for three new genera and seven new species —
<taxonomicName id="4C065BC59D572C2CFBDAFD48FB060175" box="[1115,1258,764,789]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFBDAFD48FB060175" box="[1115,1258,764,789]" italics="true" pageId="4" pageNumber="5">Paracortina</emphasis>
</taxonomicName>
(two species),
<taxonomicName id="4C065BC59D572C2CFEF7FC90FE38015D" box="[374,468,804,829]" class="Actinopterygii" family="Cyprinidae" genus="Relictus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="B972FC549D572C2CFEF7FC90FE38015D" box="[374,468,804,829]" italics="true" pageId="4" pageNumber="5">Relictus</emphasis>
</taxonomicName>
(two species), and
<taxonomicName id="4C065BC59D572C2CFD3DFC90FCEE015D" box="[700,770,804,829]" class="Diplopoda" family="Paracortinidae" genus="Altum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFD3DFC90FCEE015D" box="[700,770,804,829]" italics="true" pageId="4" pageNumber="5">Altum</emphasis>
</taxonomicName>
(three species) — and all the new generic and familial taxa were subsequently cited by Shelley
<emphasis id="B972FC549D572C2CFCCCFCF8FC640105" box="[845,904,844,869]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(2000). Wang and Mauriès (1996) likewise listed all these taxa, reducing
<taxonomicName id="4C065BC59D572C2CFD4EFCC0FCC201ED" box="[719,814,884,909]" class="Actinopterygii" family="Cyprinidae" genus="Relictus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="B972FC549D572C2CFD4EFCC0FCC201ED" box="[719,814,884,909]" italics="true" pageId="4" pageNumber="5">Relictus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C065BC59D572C2CFCE6FCC0FC4101ED" box="[871,941,884,909]" class="Diplopoda" family="Paracortinidae" genus="Altum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFCE6FCC0FC4101ED" box="[871,941,884,909]" italics="true" pageId="4" pageNumber="5">Altum</emphasis>
</taxonomicName>
to subgenera under
<taxonomicName id="4C065BC59D572C2CFB1AFCC0FAC601ED" box="[1179,1322,884,909]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFB1AFCC0FAC601ED" box="[1179,1322,884,909]" italics="true" pageId="4" pageNumber="5">Paracortina</emphasis>
</taxonomicName>
and transferring their species into this genus, but Wang (1996), in a subsequent paper in this volume, recognized three full genera, which technically is the latest word on their statuses. Our analysis of the gonopod drawings of Wang and Zhang (1993) and our experience with distantly related callipodidans in North
<collectingCountry id="F31160D69D572C2CFCDAFBA0FC2C064E" box="[859,960,1044,1070]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
indicate that there is probably only one valid genus, so we favor the arrangement of Wang and Mauriès (1996). However, the status of
<taxonomicName id="4C065BC59D572C2CFE6BFBD0FDA5061D" box="[490,585,1124,1149]" class="Actinopterygii" family="Cyprinidae" genus="Relictus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="B972FC549D572C2CFE6BFBD0FDA5061D" box="[490,585,1124,1149]" italics="true" pageId="4" pageNumber="5">Relictus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C065BC59D572C2CFD09FBD0FD22061D" box="[648,718,1124,1149]" class="Diplopoda" family="Paracortinidae" genus="Altum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFD09FBD0FD22061D" box="[648,718,1124,1149]" italics="true" pageId="4" pageNumber="5">Altum</emphasis>
</taxonomicName>
as subgenera of
<taxonomicName id="4C065BC59D572C2CFC21FBD0FBC3061D" box="[928,1071,1124,1149]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFC21FBD0FBC3061D" box="[928,1071,1124,1149]" italics="true" pageId="4" pageNumber="5">Paracortina</emphasis>
</taxonomicName>
, rather than outright synonyms, remains open to question.
</paragraph>
<paragraph id="8BB920469D572C2CFEB9FB00FB260786" blockId="4.[264,1324,284,1510]" pageId="4" pageNumber="5">
Shear (2000
<emphasis id="B972FC549D572C2CFE4DFB00FE3706AD" box="[460,475,1204,1229]" italics="true" pageId="4" pageNumber="5">a</emphasis>
) proposed
<taxonomicName id="4C065BC59D572C2CFDEBFB00FCE606AD" box="[618,778,1204,1229]" class="Diplopoda" family="Paracortinidae" genus="Scotopetalum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFDEBFB00FCE606AD" box="[618,778,1204,1229]" italics="true" pageId="4" pageNumber="5">Scotopetalum</emphasis>
</taxonomicName>
for a new species collected in a cave near Hanoi,
<collectingCountry id="F31160D69D572C2CFEE2FB68FE250696" box="[355,457,1244,1270]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
, and assigned it to the family
<taxonomicName id="4C065BC59D572C2CFCCBFB68FBEB0696" box="[842,1031,1244,1270]" class="Diplopoda" family="Schizopetalidae" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Schizopetalidae</taxonomicName>
, otherwise known from Europe, the mid­East, and, at least for now, western North
<collectingCountry id="F31160D69D572C2CFC40FAB0FBC6077E" box="[961,1066,1284,1310]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
. In addition to erecting the suborder Sinocallipodidea, Shear also suggested that the
<taxonomicName id="4C065BC59D572C2CFB8FFA98FB5B0726" box="[1038,1207,1324,1350]" class="Diplopoda" family="Paracortinidae" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Paracortinidae</taxonomicName>
probably should be considered only a subfamily of the
<taxonomicName id="4C065BC59D572C2CFCB2FAE0FC00070E" box="[819,1004,1364,1390]" class="Diplopoda" family="Schizopetalidae" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Schizopetalidae</taxonomicName>
but did not formalize this status without examining authentic specimens. Thus, the composition of the
<taxonomicName id="4C065BC59D572C2CFB1EFAC8FAC407F6" box="[1183,1320,1404,1430]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
in southeast Asia is as follows; taxa are listed in chronological order by genera and species along with
<typeStatus id="54BD9EE49D572C2CFE0DFA78FE520786" box="[396,446,1484,1510]" pageId="4" pageNumber="5">type</typeStatus>
localities and other occurrences, all of which are mapped in fig. 2.
</paragraph>
<paragraph id="8BB920469D572C2CFE89F9F0FE31043E" blockId="4.[264,1323,1604,1870]" box="[264,477,1604,1630]" pageId="4" pageNumber="5">
Order
<taxonomicName id="4C065BC59D572C2CFED5F9F0FE31043E" box="[340,477,1604,1630]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
</paragraph>
<paragraph id="8BB920469D572C2CFEB9F9D8FCFC04E5" blockId="4.[264,1323,1604,1870]" box="[312,784,1644,1670]" pageId="4" pageNumber="5">
Suborder Sinocallipodidea Shear, 2000
<emphasis id="B972FC549D572C2CFC80F9D8FCFC04E5" box="[769,784,1644,1669]" italics="true" pageId="4" pageNumber="5">a</emphasis>
</paragraph>
<paragraph id="8BB920469D572C2CFEE9F920FCCF04CE" blockId="4.[264,1323,1604,1870]" box="[360,803,1684,1710]" pageId="4" pageNumber="5">
Family
<taxonomicName id="4C065BC59D572C2CFE43F920FCCF04CE" authority="Zhang, 1993" authorityName="Zhang" authorityYear="1993" box="[450,803,1684,1710]" class="Diplopoda" family="Sinocallipodidae" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Sinocallipodidae Zhang, 1993</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C31C73CD9D572C2FFE19F908FD2201EE" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="description">
<paragraph id="8BB920469D572C2CFE19F908FCFB04B6" blockId="4.[264,1323,1604,1870]" box="[408,791,1724,1750]" pageId="4" pageNumber="5">
Genus
<taxonomicName id="4C065BC59D572C2CFE6BF908FCFB04B6" authority="Zhang, 1993" authorityName="Zhang" authorityYear="1993" box="[490,791,1724,1750]" class="Diplopoda" family="Sinocallipodidae" genus="Sinocallipus" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D572C2CFE6BF908FD9704B5" box="[490,635,1724,1749]" italics="true" pageId="4" pageNumber="5">Sinocallipus</emphasis>
Zhang, 1993
</taxonomicName>
</paragraph>
<paragraph id="8BB920469D572C2CFE49F950FB0C052E" blockId="4.[264,1323,1604,1870]" pageId="4" pageNumber="5">
<taxonomicName id="4C065BC59D572C2CFE49F950FCCB049E" authority="Zhang, 1993" authorityName="Zhang" authorityYear="1993" box="[456,807,1764,1790]" class="Diplopoda" family="Sinocallipodidae" genus="Sinocallipus" kingdom="Animalia" order="Callipodida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="simplipodicus">
<emphasis id="B972FC549D572C2CFE49F950FD64049D" box="[456,648,1764,1789]" italics="true" pageId="4" pageNumber="5">S. simplipodicus</emphasis>
Zhang, 1993
</taxonomicName>
.
<collectingCountry id="F31160D69D572C2CFCB8F950FC6E049E" box="[825,898,1764,1790]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
, Yunnan Prov., Hekou Yaozu Autonomous County, Xiao Cave (Zhang, 1993). Also occurring in
<collectingCountry id="F31160D69D572C2CFE79F880FDD8052E" box="[504,564,1844,1870]" name="Laos" pageId="4" pageNumber="5">Laos</collectingCountry>
, Champasak/Attapu Provs., Bolavens Plateau (this paper).
</paragraph>
<paragraph id="8BB920469D562C2DFEB9FEA8FCF70356" blockId="5.[312,1324,284,1710]" box="[312,795,284,310]" pageId="5" pageNumber="6">Suborder Schizopetalidea Verhoeff, 1909</paragraph>
<paragraph id="8BB920469D562C2DFEE9FEF0FC68033E" blockId="5.[312,1324,284,1710]" box="[360,900,324,350]" pageId="5" pageNumber="6">
Family
<taxonomicName id="4C065BC59D562C2DFE43FEF0FC68033E" authority="Wang and Zhang, 1993" authorityName="Wang and Zhang" authorityYear="1993" box="[450,900,324,350]" class="Diplopoda" family="Paracortinidae" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Paracortinidae Wang and Zhang, 1993</taxonomicName>
</paragraph>
<paragraph id="8BB920469D562C2DFE19FED8FC7E03E6" blockId="5.[312,1324,284,1710]" box="[408,914,364,390]" pageId="5" pageNumber="6">
Genus
<taxonomicName id="4C065BC59D562C2DFE6BFED8FC7E03E6" authority="Wang and Zhang, 1993" authorityName="Wang and Zhang" authorityYear="1993" box="[490,914,364,390]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D562C2DFE6BFED8FD9503E5" box="[490,633,364,389]" italics="true" pageId="5" pageNumber="6">Paracortina</emphasis>
Wang and Zhang, 1993
</taxonomicName>
</paragraph>
<paragraph id="8BB920469D562C2DFE49FE20FC0503CE" blockId="5.[312,1324,284,1710]" box="[456,1001,404,430]" pageId="5" pageNumber="6">
Subgenus
<taxonomicName id="4C065BC59D562C2DFDC0FE20FC0503CE" authority="Wang and Zhang, 1993" authorityName="Wang and Zhang" authorityYear="1993" box="[577,1001,404,430]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D562C2DFDC0FE20FD3C03CD" box="[577,720,404,429]" italics="true" pageId="5" pageNumber="6">Paracortina</emphasis>
Wang and Zhang, 1993
</taxonomicName>
</paragraph>
<paragraph id="8BB920469D562C2DFE79FE08FBAA039E" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<emphasis id="B972FC549D562C2DFE79FE08FD2903B5" box="[504,709,444,469]" italics="true" pageId="5" pageNumber="6">P. (P.) leptoclada</emphasis>
Wang and Zhang, 1993.
<collectingCountry id="F31160D69D562C2DFB80FE08FBA603B6" box="[1025,1098,444,470]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Yunnan Province, Zhong Dian County (Wang and Zhang, 1993).
</paragraph>
<paragraph id="8BB920469D562C2DFE79FDB8FC02002E" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<emphasis id="B972FC549D562C2DFE79FDB8FD7F0045" box="[504,659,524,549]" italics="true" pageId="5" pageNumber="6">P. (P.) voluta</emphasis>
Wang and Zhang, 1993.
<collectingCountry id="F31160D69D562C2DFC53FDB8FBF70046" box="[978,1051,524,550]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Sichuan Province, Ya Jang County (Wang and Zhang, 1993).
</paragraph>
<paragraph id="8BB920469D562C2DFE49FDE8FC540016" blockId="5.[312,1324,284,1710]" box="[456,952,604,630]" pageId="5" pageNumber="6">
Subgenus
<taxonomicName id="4C065BC59D562C2DFDC0FDE8FC540016" authority="Wang and Zhang, 1993" authorityName="Wang and Zhang" authorityYear="1993" box="[577,952,604,630]" class="Actinopterygii" family="Cyprinidae" genus="Relictus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B972FC549D562C2DFDC0FDE8FD730015" box="[577,671,604,629]" italics="true" pageId="5" pageNumber="6">Relictus</emphasis>
Wang and Zhang, 1993
</taxonomicName>
</paragraph>
<paragraph id="8BB920469D562C2DFE79FD30FBA900A6" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<emphasis id="B972FC549D562C2DFE79FD30FDA100FD" box="[504,589,644,669]" italics="true" pageId="5" pageNumber="6">P. (R..)</emphasis>
<taxonomicName id="4C065BC59D562C2DFDD8FD30FC3300FE" authority="Wang and Zhang, 1993" authorityName="Wang and Zhang" authorityYear="1993" box="[601,991,644,670]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="stimula">
<emphasis id="B972FC549D562C2DFDD8FD30FD4300FD" box="[601,687,644,669]" italics="true" pageId="5" pageNumber="6">stimula</emphasis>
Wang and Zhang, 1993
</taxonomicName>
.
<collectingCountry id="F31160D69D562C2DFC7DFD30FBA900FE" box="[1020,1093,644,670]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Yunnan Province, Zhong Dian County (Wang and Zhang, 1993).
</paragraph>
<paragraph id="8BB920469D562C2DFE79FD60FC5D015E" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<emphasis id="B972FC549D562C2DFE79FD60FD47008D" box="[504,683,724,749]" italics="true" pageId="5" pageNumber="6">P. (R..) thallina</emphasis>
Wang and Zhang, 1993.
<collectingCountry id="F31160D69D562C2DFC58FD60FBCE008E" box="[985,1058,724,750]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Sichuan Province, Ba Tang County. Also recorded from Yunnan Province, Zhong Dian County (Wang and Zhang, 1993).
</paragraph>
<paragraph id="8BB920469D562C2DFE45FCF8FC700106" blockId="5.[312,1324,284,1710]" box="[452,924,844,870]" pageId="5" pageNumber="6">
Subgenus
<taxonomicName id="4C065BC59D562C2DFDBCFCF8FC700106" authority="Wang and Zhang, 1993" authorityName="Wang and Zhang" authorityYear="1993" box="[573,924,844,870]" class="Diplopoda" family="Paracortinidae" genus="Altum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D562C2DFDBCFCF8FD6F0105" box="[573,643,844,869]" italics="true" pageId="5" pageNumber="6">Altum</emphasis>
Wang and Zhang, 1993
</taxonomicName>
</paragraph>
<paragraph id="8BB920469D562C2DFE79FCC0FC5D01BE" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<emphasis id="B972FC549D562C2DFE79FCC0FDA401ED" box="[504,584,884,909]" italics="true" pageId="5" pageNumber="6">P. (A.)</emphasis>
<taxonomicName id="4C065BC59D562C2DFDD4FCC0FC3601EE" authority="Wang and Zhang, 1993" authorityName="Wang and Zhang" authorityYear="1993" box="[597,986,884,910]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="viriosa">
<emphasis id="B972FC549D562C2DFDD4FCC0FD4A01ED" box="[597,678,884,909]" italics="true" pageId="5" pageNumber="6">viriosa</emphasis>
Wang and Zhang, 1993
</taxonomicName>
.
<collectingCountry id="F31160D69D562C2DFC78FCC0FBAE01EE" box="[1017,1090,884,910]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Yunnan Province, Zhong Dian County. Also recorded from Tibet, Mang Kang County (Wang and Zhang, 1993).
</paragraph>
<paragraph id="8BB920469D562C2DFE79FC58FD74061E" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<emphasis id="B972FC549D562C2DFE79FC58FD720665" box="[504,670,1004,1029]" italics="true" pageId="5" pageNumber="6">P. (A.) serrata</emphasis>
Wang and Zhang, 1993.
<collectingCountry id="F31160D69D562C2DFC4DFC58FBF90666" box="[972,1045,1004,1030]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Tibet, De Qin Autonomous Region (Wang and Zhang, 1993). This site is in the southeast periphery of Tibet, just across the border from Yunnan Province.
</paragraph>
<paragraph id="8BB920469D562C2DFE79FB38FBA906AE" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<emphasis id="B972FC549D562C2DFE79FB38FD5806C5" box="[504,692,1164,1189]" italics="true" pageId="5" pageNumber="6">P. (A.) carinata</emphasis>
Wang and Zhang, 1993.
<collectingCountry id="F31160D69D562C2DFC7CFB38FBAA06C6" box="[1021,1094,1164,1190]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Yunnan Province, Zhong Dian County (Wang and Zhang, 1993).
</paragraph>
<paragraph id="8BB920469D562C2DFE19FB68FAF90696" blockId="5.[312,1324,284,1710]" box="[408,1301,1244,1270]" pageId="5" pageNumber="6">
Unidentifiable genus and species.
<collectingCountry id="F31160D69D562C2DFCB1FB68FC950696" box="[816,889,1244,1270]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
, Jiangsu Province, Cisian­shan, ca.
</paragraph>
<paragraph id="8BB920469D562C2DFDA9FAB0FBA5077E" blockId="5.[312,1324,284,1710]" box="[552,1097,1284,1310]" pageId="5" pageNumber="6">
15.6 mi (
<quantity id="4CFE8DA39D562C2DFD10FAB0FD32077E" box="[657,734,1284,1310]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="6" unit="km" value="25.0">25 km</quantity>
) S Nanjing (Golovatch, 1981).
</paragraph>
<paragraph id="8BB920469D562C2DFEE9FA98FCDE0726" blockId="5.[312,1324,284,1710]" box="[360,818,1324,1350]" pageId="5" pageNumber="6">
Family
<taxonomicName id="4C065BC59D562C2DFE43FA98FCDE0726" authority="Verhoeff, 1909" authorityName="Verhoeff" authorityYear="1909" box="[450,818,1324,1350]" class="Diplopoda" family="Schizopetalidae" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Schizopetalidae Verhoeff, 1909</taxonomicName>
</paragraph>
<paragraph id="8BB920469D562C2DFE19FAE0FCC6070D" blockId="5.[312,1324,284,1710]" box="[408,810,1364,1390]" pageId="5" pageNumber="6">
Genus
<taxonomicName id="4C065BC59D562C2DFE6BFAE0FCF7070E" authority="Shear, 2000" authorityName="Shear" authorityYear="2000" box="[490,795,1364,1390]" class="Diplopoda" family="Paracortinidae" genus="Scotopetalum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972FC549D562C2DFE6BFAE0FD65070D" box="[490,649,1364,1389]" italics="true" pageId="5" pageNumber="6">Scotopetalum</emphasis>
Shear, 2000
</taxonomicName>
<emphasis id="B972FC549D562C2DFC9AFAE0FCC6070D" box="[795,810,1364,1389]" italics="true" pageId="5" pageNumber="6">a</emphasis>
</paragraph>
<paragraph id="8BB920469D562C2DFE45FAC8FAFD04CE" blockId="5.[312,1324,284,1710]" pageId="5" pageNumber="6">
<taxonomicName id="4C065BC59D562C2DFE45FAC8FD3C07F6" authority="Shear, 2000" authorityName="Shear" authorityYear="2000" box="[452,720,1404,1430]" class="Diplopoda" family="Paracortinidae" genus="Scotopetalum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="warreni">
<emphasis id="B972FC549D562C2DFE45FAC8FDD107F5" box="[452,573,1404,1429]" italics="true" pageId="5" pageNumber="6">S. warreni</emphasis>
Shear, 2000
</taxonomicName>
<emphasis id="B972FC549D562C2DFD4EFAC8FD0907F5" box="[719,741,1404,1429]" italics="true" pageId="5" pageNumber="6">a.</emphasis>
<collectingCountry id="F31160D69D562C2DFD75FAC8FCB607F6" box="[756,858,1404,1430]" name="Vietnam" pageId="5" pageNumber="6">Vietnam</collectingCountry>
, northwest of Hanoi, anunnamed cave at Hong Mat; also tentatively recorded from Huang Tua Cave northwest of Hanoi (Shear, 2000
<emphasis id="B972FC549D562C2DFCCCFA78FCB00785" box="[845,860,1484,1509]" italics="true" pageId="5" pageNumber="6">a</emphasis>
). The exact locations of these sites, which are south of the Tropic of Cancer, are unknown, and Hong (Huong, Hoang, Hung) Mat is not on any map available to us, nor can we find any town with the word, “Mat,” as part of its name. The coordinates given for the
<typeStatus id="54BD9EE49D562C2DFD6CF9D8FCF304E6" box="[749,799,1644,1670]" pageId="5" pageNumber="6">type</typeStatus>
locality are in fact southwest, not northwest, of Hanoi, and we have placed them on the map accordingly (fig. 2).
</paragraph>
<caption id="DF7970CE9D552C2EFE89FB2AFC3E07E6" httpUri="https://zenodo.org/record/294083/files/figure.png" pageId="6" pageNumber="7" targetBox="[287,1299,313,1172]" targetPageId="6">
<paragraph id="8BB920469D552C2EFE89FB2AFC3E07E6" blockId="6.[264,1324,1182,1414]" pageId="6" pageNumber="7">
<emphasis id="B972FC549D552C2EFE89FB2AFE6706D6" bold="true" box="[264,395,1182,1206]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
Distribution of the
<taxonomicName id="4C065BC59D552C2EFDE8FB2BFD0906D6" box="[617,741,1183,1206]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
in southeastern Asia. The Yangtze River is indicated in China; the dotted line, a smooth curve drawn around range extremes, shows the border of the projected ordinal distribution shaded in fig. 1; and the dashed line represents the Tropic of Cancer. Dots,
<taxonomicName id="4C065BC59D552C2EFECBFAB3FD87077E" box="[330,619,1287,1310]" class="Diplopoda" family="Sinocallipodidae" genus="Sinocallipus" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="simplipodicus">
<emphasis id="B972FC549D552C2EFECBFAB3FD87077E" box="[330,619,1287,1310]" italics="true" pageId="6" pageNumber="7">Sinocallipus simplipodicus</emphasis>
</taxonomicName>
; upright triangle,
<emphasis id="B972FC549D552C2EFCB2FAB3FBB1077E" box="[819,1117,1287,1310]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C065BC59D552C2EFCB2FAB3FC59077E" box="[819,949,1287,1310]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Paracortina</taxonomicName>
(P.) leptoclada
</emphasis>
; solid diamond,
<emphasis id="B972FC549D552C2EFA96FAB3FE900721" italics="true" pageId="6" pageNumber="7">P. (P.) voluta</emphasis>
; solid square,
<taxonomicName id="4C065BC59D552C2EFDA7FA9EFCEA0721" box="[550,774,1322,1345]" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="stimula" subGenus="Relictus">
<emphasis id="B972FC549D552C2EFDA7FA9EFCEA0721" box="[550,774,1322,1345]" italics="true" pageId="6" pageNumber="7">P. (Relictus) stimula</emphasis>
</taxonomicName>
; open squares in China,
<emphasis id="B972FC549D552C2EFBA7FA9EFB2D0721" box="[1062,1217,1322,1345]" italics="true" pageId="6" pageNumber="7">P.(R.) thallina</emphasis>
; stars,
<taxonomicName id="4C065BC59D552C2EFA96FA9EFE470703" class="Diplopoda" family="Paracortinidae" genus="Paracortina" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="viriosa" subGenus="Altum">
<emphasis id="B972FC549D552C2EFA96FA9EFE470703" italics="true" pageId="6" pageNumber="7">P. (Altum) viriosa</emphasis>
</taxonomicName>
; inverted triangle,
<emphasis id="B972FC549D552C2EFDF6FAF8FCE30703" box="[631,783,1356,1379]" italics="true" pageId="6" pageNumber="7">P. (A.) serrata</emphasis>
; circle,
<emphasis id="B972FC549D552C2EFCE7FAF8FBF90703" box="[870,1045,1356,1379]" italics="true" pageId="6" pageNumber="7">P. (A.) carinata;</emphasis>
open squares in Vietnam,
<taxonomicName id="4C065BC59D552C2EFE89FADBFE1F07E6" box="[264,499,1391,1414]" class="Diplopoda" family="Paracortinidae" genus="Scotopetalum" higherTaxonomySource="CoL" kingdom="Animalia" order="Callipodida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="warreni">
<emphasis id="B972FC549D552C2EFE89FADBFE1F07E6" box="[264,499,1391,1414]" italics="true" pageId="6" pageNumber="7">Scotopetalum warreni</emphasis>
</taxonomicName>
;?, unknown paracortinid genus and species.
</paragraph>
</caption>
<paragraph id="8BB920469D552C2FFEB9FA0EFD2201EE" blockId="6.[264,1323,1466,1892]" lastBlockId="7.[264,1325,284,910]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
The occurrence of identifiable callipodidans in Tibet, Sichuan, and Yunnan Provinces,
<collectingCountry id="F31160D69D552C2EFE89FA56FEBD079C" box="[264,337,1506,1532]" name="China" pageId="6" pageNumber="7">China</collectingCountry>
, northern
<collectingCountry id="F31160D69D552C2EFE4BFA56FDDC079C" box="[458,560,1506,1532]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
, and southern
<collectingCountry id="F31160D69D552C2EFD5CFA56FCF5079C" box="[733,793,1506,1532]" name="Laos" pageId="6" pageNumber="7">Laos</collectingCountry>
, and the discovery of unidentifiable females in eastern Jiangsu Province, ca.
<quantity id="4CFE8DA39D552C2EFD06F9BEFD370444" box="[647,731,1546,1572]" metricMagnitude="5" metricUnit="m" metricValue="2.8968192" pageId="6" pageNumber="7" unit="mi" value="180.0">180 mi</quantity>
(
<quantity id="4CFE8DA39D552C2EFD6DF9BEFCA60444" box="[748,842,1546,1572]" metricMagnitude="5" metricUnit="m" metricValue="2.88" pageId="6" pageNumber="7" unit="km" value="288.0">288 km</quantity>
) west of metropolitan Shanghai and not far from the mouth of the Yangtze River, is evidence of a widespread callipodidan fauna in southeastern Asia (
<figureCitation id="133D3CC39D552C2EFE66F9EEFDB40414" box="[487,600,1626,1652]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[264,368,1296,1320]" captionStartId-1="6.[264,368,1182,1206]" captionTargetBox-0="[283,1313,932,1285]" captionTargetBox-1="[287,1299,313,1172]" captionTargetId-0="figure@2.[274,1314,929,1289]" captionTargetId-1="figure@6.[285,1310,308,1172]" captionTargetPageId-0="2" captionTargetPageId-1="6" captionText-0="FIGURE 1. Approximate global distribution of the order Callipodida." captionText-1="FIGURE 2. Distribution of the Callipodida in southeastern Asia. The Yangtze River is indicated in China; the dotted line, a smooth curve drawn around range extremes, shows the border of the projected ordinal distribution shaded in fig. 1; and the dashed line represents the Tropic of Cancer. Dots, Sinocallipus simplipodicus; upright triangle, Paracortina (P.) leptoclada; solid diamond, P. (P.) voluta; solid square, P. (Relictus) stimula; open squares in China, P. (R.) thallina; stars, P. (Altum) viriosa; inverted triangle, P. (A.) serrata; circle, P. (A.) carinata; open squares in Vietnam, Scotopetalum warreni;?, unknown paracortinid genus and species." httpUri-0="https://zenodo.org/record/294082/files/figure.png" httpUri-1="https://zenodo.org/record/294083/files/figure.png" pageId="6" pageNumber="7">Figs. 12</figureCitation>
). Most of the material comes from montane areas around the contiguous corners of Tibet, Sichuan, and Yunnan Provinces; drawing a smooth curve around range extremes produces a general area of roughly
<quantity id="4CFE8DA39D552C2EFC34F91EFBF104A4" box="[949,1053,1706,1732]" metricMagnitude="3" metricUnit="m" metricValue="2.414016" pageId="6" pageNumber="7" unit="mi" value="1.5">1,500 mi</quantity>
(
<quantity id="4CFE8DA39D552C2EFBAAF91EFB4C04A4" box="[1067,1184,1706,1732]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="km" value="2.4">2,400 km</quantity>
), east­west, and
<quantity id="4CFE8DA39D552C2EFEB8F966FE4D048C" box="[313,417,1746,1772]" metricMagnitude="3" metricUnit="m" metricValue="1.9312128" pageId="6" pageNumber="7" unit="mi" value="1.2">1,200 mi</quantity>
(
<quantity id="4CFE8DA39D552C2EFE31F966FDC8048C" box="[432,548,1746,1772]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="6" pageNumber="7" unit="km" value="1.92">1,920 km</quantity>
), north­south, which equals some 1,800,000 sq. mi. (4,608,000 sq. km). In
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, localities exist both north and south of the Yangtze River, and the northern­ and westernmost records indicate that callipodidans may potentially be discovered in parts or all of the following 15 provinces: Tibet, Qinghai, Sichuan, Yunnan, Guizhou, Guangxi, Guangdong, Hunan, Jiangxi, Fujian, Zhejiang, Hubei, Anhui, Jiangsu, and Shanghai Shi. This includes essentially the entire area south of the Yangtze except for distant reaches near the headwaters and likely extends northward for an unknown distance. Judging from known records, the region probably extends well into
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,
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,
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, and southern
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, and possibly covers the entire Indochinese Peninsula; occurrences in the easternmost part of
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(Assam), the Malay Peninsula, and Hainan Island,
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, are also plausible. This vast, poorly investigated area is roughly equivalent in size to that in eastern North
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and surely harbors many undiscovered species and genera. Based upon its size, Shear (2000
<emphasis id="B972FC549D542C2FFDC0FDE8FDBC0015" box="[577,592,604,629]" italics="true" pageId="7" pageNumber="8">a</emphasis>
) was surely correct in suggesting that southeast Asia constitutes a third center of ordinal diversity in addition to southwestern North
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and the eastern Mediterranean (
<figureCitation id="133D3CC39D542C2FFDA1FD18FD8B00A6" box="[544,615,684,710]" captionStart="FIGURE 1" captionStartId="2.[264,368,1296,1320]" captionTargetBox="[283,1313,932,1285]" captionTargetId="figure@2.[274,1314,929,1289]" captionTargetPageId="2" captionText="FIGURE 1. Approximate global distribution of the order Callipodida." httpUri="https://zenodo.org/record/294082/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
). Indeed, this may be an understatement; its diversity may be greater, such that southeast Asia may be the primary center worldwide. Wang and Mauriès (1996) stated that probably no more than 5% of the total Chinese myriapod species had been discovered, and to some degree this statement also applies to southeast Asia in general, the
<taxonomicName id="4C065BC59D542C2FFE37FCF8FDD30106" box="[438,575,844,870]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
being a case in point. Field work in the area in which the order is projected to occur is greatly desired.
</paragraph>
</subSubSection>
</treatment>
</document>