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<document id="91D88268C221F24EBE3B4DADEEA9EDD0" ID-DOI="10.5281/zenodo.5312352" ID-ISSN="0374-1036" ID-Zenodo-Dep="5312352" ID-ZooBank="2C4DD1B7-A32F-41FF-B866-3AC74B3083EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630119354752" checkinUser="marcus" docAuthor="Kočárek, Petr" docDate="2014" docId="03AACA29C625FF868136FEF1FC6E0629" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.1-22.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Guanchia sokotrana Kočárek 2014, comb. nov." docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="FF93B251C628FF8B8069FFFBFFAF0541" masterDocTitle="Earwigs (Dermaptera) of Socotra Island: checklist, distribution, and description of a new genus and four new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="14" updateTime="1699159095649" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="63B06CF6DF77C7969459F8F64DA0E608">Earwigs (Dermaptera) of Socotra Island: checklist, distribution, and description of a new genus and four new species</mods:title>
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<mods:namePart id="A354392A6BE1737BB9263528D3A7441D">Kočárek, Petr</mods:namePart>
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<mods:date id="52F9ADFB31A0B4FB6199CBC7DCB7F8D4">2014</mods:date>
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<subSubSection id="C31928B4C625FF868136FEF1FCFC0463" box="[351,851,266,290]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BBC7B3FC625FF868136FEF1FCFC0463" blockId="13.[351,851,266,290]" box="[351,851,266,290]" pageId="13" pageNumber="14">
<heading id="D0F4CC53C625FF868136FEF1FCFC0463" bold="true" box="[351,851,266,290]" centered="true" fontSize="8" level="2" pageId="13" pageNumber="14" reason="3">
<emphasis id="B977A72DC625FF868136FEF1FCFC0463" bold="true" box="[351,851,266,290]" pageId="13" pageNumber="14">
<taxonomicName id="4C0300BCC625FF868136FEF1FD7E0463" ID-CoL="3HJ39" authority="(Burr, 1905)" authorityName="Kočárek" authorityYear="2014" baseAuthorityName="Burr" baseAuthorityYear="1905" box="[351,721,266,290]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana" status="comb. nov.">
<emphasis id="B977A72DC625FF868136FEF1FDEE0463" bold="true" box="[351,577,266,290]" italics="true" pageId="13" pageNumber="14">Guanchia sokotrana</emphasis>
(
<bibRefCitation id="EF9206CEC625FF868226FEF0FD640463" author="BURR M." box="[591,715,267,290]" pageId="13" pageNumber="14" pagination="468 - 496" refId="ref10949" refString="BURR M. 1905: Notes on Forficularia. IX. One new species with synonymic notes. Annals and Magazine of Natural History, Series 7 16: 468 - 496." type="journal article" year="1905">Burr, 1905</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2441A56C625FF8682B0FEF0FCFC0463" box="[729,851,267,290]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BBC7B3FC625FF86827BFED2FD0F047D" blockId="13.[113,672,297,351]" box="[530,672,297,316]" pageId="13" pageNumber="14">
(
<figureCitation id="133867BAC625FF868271FED2FDD5047D" box="[536,634,297,316]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="13" pageNumber="14">Figs 2023</figureCitation>
,
<figureCitation id="133867BAC625FF8682EAFED2FD36047D" box="[643,665,297,316]" captionStart="Figs 2729" captionStartId="16.[151,188,1531,1550]" captionTargetBox="[194,1079,235,1493]" captionTargetId="figure-11@16.[186,1091,219,1516]" captionTargetPageId="16" captionText="Figs 2729. Sketch maps of Socotra Island displaying the known and newly recorded distribution of Dermaptera. Published records are based on HAAS at al. (2004); the occurrence of Labidura riparia (Pallas, 1773) on Samha Island published by these authors is not displayed." figureDoi="http://doi.org/10.5281/zenodo.5312366" httpUri="https://zenodo.org/record/5312366/files/figure.png" pageId="13" pageNumber="14">28</figureCitation>
)
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<paragraph id="8BBC7B3FC625FF868018FEB7FE04041E" blockId="13.[113,672,297,351]" box="[113,427,332,351]" pageId="13" pageNumber="14">
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<treatmentCitation id="0AA25D2EC625FF868018FEB7FE04041E" author="BURR M." box="[113,427,332,351]" page="493" pageId="13" pageNumber="14" year="1905">
<taxonomicName id="4C0300BCC625FF868018FEB7FE04041E" ID-CoL="DVVJ" authority="Burr, 1905: 493" authorityName="Burr" authorityPageNumber="493" authorityYear="1905" box="[113,427,332,351]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868018FEB7FEB0041E" box="[113,287,332,351]" italics="true" pageId="13" pageNumber="14">Anechura sokotrana</emphasis>
<bibRefCitation id="EF9206CEC625FF86814DFEB7FE04041E" author="BURR M." box="[292,427,332,351]" pageId="13" pageNumber="14" pagination="468 - 496" refId="ref10949" refString="BURR M. 1905: Notes on Forficularia. IX. One new species with synonymic notes. Annals and Magazine of Natural History, Series 7 16: 468 - 496." type="journal article" year="1905">Burr, 1905: 493</bibRefCitation>
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</paragraph>
</subSubSection>
<subSubSection id="C31928B4C625FF868018FE8EFE870739" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BBC7B3FC625FF868018FE8EFE1804C9" blockId="13.[113,1090,373,632]" box="[113,439,373,392]" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FE8EFF4504C9" bold="true" box="[113,234,373,392]" pageId="13" pageNumber="14">
<typeStatus id="54B8C59DC625FF868018FE8EFF3104C9" box="[113,158,373,392]" pageId="13" pageNumber="14">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3143BAFC625FF868086FE8EFE8204C9" box="[239,301,373,392]" name="Yemen" pageId="13" pageNumber="14">Yemen</collectingCountry>
,
<collectingCountry id="F3143BAFC625FF86815CFE8EFE1C04C9" box="[309,435,373,392]" name="Socotra Island" pageId="13" pageNumber="14">Socotra Island</collectingCountry>
.
</paragraph>
<paragraph id="8BBC7B3FC625FF868018FE6BFB9C04B2" blockId="13.[113,1090,373,632]" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FE6BFF5A04E2" bold="true" box="[113,245,400,419]" pageId="13" pageNumber="14">Type material.</emphasis>
Not studied.The deposition of type material is not stated in the original description of
<bibRefCitation id="EF9206CEC625FF8683B8FE6BFB9104E2" author="BURR M." box="[977,1086,400,419]" pageId="13" pageNumber="14" pagination="468 - 496" refId="ref10949" refString="BURR M. 1905: Notes on Forficularia. IX. One new species with synonymic notes. Annals and Magazine of Natural History, Series 7 16: 468 - 496." type="journal article" year="1905">BURR (1905)</bibRefCitation>
.
<bibRefCitation id="EF9206CEC625FF868018FE50FEA804FF" author="HAAS F. &amp; POHL H. &amp; WRANIK W." box="[113,263,427,446]" pageId="13" pageNumber="14" pagination="409 - 419" refId="ref11078" refString="HAAS F., POHL H. &amp; WRANIK W. 2004: Dermaptera of the Socotra Archipelago with the description of a new species. Fauna of Arabia 20: 409 - 419." type="journal article" year="2004">HAAS et al. (2004)</bibRefCitation>
suggested that the
<typeStatus id="54B8C59DC625FF8681CAFE50FE4204FF" box="[419,493,427,446]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
could be deposited in The Natural History Museum, London.There is a specimen (male) labelled as
<taxonomicName id="4C0300BCC625FF868132FE3EFD220499" authority="Saussure, 1874" authorityName="Saussure" authorityYear="1874" box="[347,653,453,472]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fedchenkoi">
<emphasis id="B977A72DC625FF868132FE3EFDA30499" box="[347,524,453,472]" italics="true" pageId="13" pageNumber="14">Anechura fedchenkoi</emphasis>
Saussure, 1874
</taxonomicName>
(now
<taxonomicName id="4C0300BCC625FF8682A9FE3EFBB10499" authority="(Saussure, 1874))" baseAuthorityName="Saussure" baseAuthorityYear="1874" box="[704,1054,453,472]" class="Insecta" family="Forficulidae" genus="Oreasiobia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fedschenkoi">
<emphasis id="B977A72DC625FF8682A9FE3EFC2A0499" box="[704,901,453,472]" italics="true" pageId="13" pageNumber="14">Oreasiobia fedschenkoi</emphasis>
(Saussure, 1874))
</taxonomicName>
; but this specimen is marked by an additional label stating described as new /
<taxonomicName id="4C0300BCC625FF868280FE1BFCF904B2" authorityName="Burr" authorityYear="1905" box="[745,854,480,499]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868280FE1BFCF904B2" box="[745,854,480,499]" italics="true" pageId="13" pageNumber="14">A. sokotrana</emphasis>
</taxonomicName>
/ Burr inedit / Type MB.
</paragraph>
<paragraph id="8BBC7B3FC625FF868018FE00FE870739" blockId="13.[113,1090,373,632]" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FE00FE8B074F" bold="true" box="[113,292,507,526]" pageId="13" pageNumber="14">Material examined.</emphasis>
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,
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,
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,
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,
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,
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.
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lgt. (
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,
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)
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;
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,
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env.,
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N,
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,
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a.s.l.
</elevation>
,
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</date>
, 3 JJ
<specimenCount id="9D05B0B6C625FF8683C7FDEEFC700769" box="[942,991,527,552]" pageId="13" pageNumber="14" type="female">4 ♀♀</specimenCount>
,
<collectorName id="26F61EE9C625FF868382FDEEFBEF0769" box="[1003,1088,533,552]" pageId="13" pageNumber="14">J. Batelka</collectorName>
lgt. (
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)
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;
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,
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env.,
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N,
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,
<elevation id="002E9C0CC625FF8682ABFDCBFC9C0702" box="[706,819,560,579]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="13" pageNumber="14" unit="m" value="1450.0">
<quantity id="4CFBD6DAC625FF8682ABFDCBFCAB0702" box="[706,772,560,579]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="13" pageNumber="14" unit="m" value="1450.0">1450 m</quantity>
a.s.l.
</elevation>
, montane evergreen woodland,
<date id="FFBD5DFFC625FF868018FDB0FF5B071F" box="[113,244,587,606]" pageId="13" pageNumber="14" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">
<collectingDate id="EFF9A417C625FF868018FDB0FF5B071F" box="[113,244,587,606]" pageId="13" pageNumber="14" value="2012-06-16" valueMax="2012-06-18" valueMin="2012-06-16">16.18.vi.2012</collectingDate>
</date>
, 4 JJ
<specimenCount id="9D05B0B6C625FF86815DFDB0FECB071C" box="[308,356,580,606]" pageId="13" pageNumber="14" type="female">5 ♀♀</specimenCount>
,
<collectorName id="26F61EE9C625FF868119FDB0FE69071F" box="[368,454,587,606]" pageId="13" pageNumber="14">J. Bezděk</collectorName>
,
<collectorName id="26F61EE9C625FF8681A6FDB0FDB7071F" box="[463,536,587,606]" pageId="13" pageNumber="14">J. Hájek</collectorName>
,
<collectionCode id="ED12E3FAC625FF868248FDB0FD9F071F" box="[545,560,587,606]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="14" type="Museum">V</collectionCode>
.
<collectorName id="26F61EE9C625FF868250FDB0FDD4071F" box="[569,635,587,606]" pageId="13" pageNumber="14">Hula, P</collectorName>
.
<collectorName id="26F61EE9C625FF8682EDFDB0FD7D071F" box="[644,722,587,606]" pageId="13" pageNumber="14">Kment, I</collectorName>
.
<collectorName id="26F61EE9C625FF8682B5FDB0FCF4071F" box="[732,859,587,606]" pageId="13" pageNumber="14">Malenovský, J</collectorName>
. Niedobová &amp;
<collectorName id="26F61EE9C625FF868389FDB0FBEF071F" box="[992,1088,587,606]" pageId="13" pageNumber="14">L. Purchart</collectorName>
lgt. (
<collectionCode id="ED12E3FAC625FF8680F1FD9EFF770739" box="[152,216,613,632]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="14">NMPC</collectionCode>
,
<collectionCode id="ED12E3FAC625FF868088FD9EFEB10739" box="[225,286,613,632]" pageId="13" pageNumber="14">PKCO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31928B4C625FF868018FD6AFD510609" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BBC7B3FC625FF868018FD6AFD510609" blockId="13.[113,1089,657,872]" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FD6AFF5C07E9" bold="true" box="[113,243,657,680]" pageId="13" pageNumber="14">Comments.</emphasis>
The sternal plates of this species have structure typical for the subfamily
<taxonomicName id="4C0300BCC625FF868382FD6AFF0A0789" class="Insecta" family="Forficulidae" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Forficulinae">Forficulinae</taxonomicName>
with a quadratic, as wide as long mesosternum (
<figureCitation id="133867BAC625FF8682C9FD4AFD400789" box="[672,751,689,712]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
). The shape of the mesosternum is the only diagnostic characteristic that subdivides
<taxonomicName id="4C0300BCC625FF8682E4FD2AFCA107A9" box="[653,782,721,744]" class="Insecta" family="Forficulidae" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Forficulinae">Forficulinae</taxonomicName>
and
<taxonomicName id="4C0300BCC625FF868354FD2AFC6607A9" authorityName="Burr" authorityYear="1907" box="[829,969,721,744]" class="Insecta" family="Forficulidae" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Anechurinae">Anechurinae</taxonomicName>
. Therefore,
<taxonomicName id="4C0300BCC625FF868018FD0AFEE50649" authorityName="Burr" authorityYear="1905" box="[113,330,753,776]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868018FD0AFEE50649" box="[113,330,753,776]" italics="true" pageId="13" pageNumber="14">Anechura sokotrana</emphasis>
</taxonomicName>
must be moved to the subfamily
<taxonomicName id="4C0300BCC625FF8682C4FD0AFC9F0649" box="[685,816,753,776]" class="Insecta" family="Forficulidae" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Forficulinae">Forficulinae</taxonomicName>
, within which the species fits well with the generic diagnosis of the genus
<taxonomicName id="4C0300BCC625FF8682FCFCEAFC2F0669" authority="Burr, 1911" authorityName="Burr" authorityYear="1911" box="[661,896,785,808]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B977A72DC625FF8682FCFCEAFD500669" box="[661,767,785,808]" italics="true" pageId="13" pageNumber="14">Guanchia</emphasis>
Burr, 1911
</taxonomicName>
, characterised by abbreviated, obliquely posteriorly truncate tegmina (
<figureCitation id="133867BAC625FF8682CBFCCAFD5D0609" box="[674,754,817,840]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="13" pageNumber="14">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31928B4C625FF868018FCAAFC6E0629" box="[113,961,849,872]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BBC7B3FC625FF868018FCAAFC6E0629" blockId="13.[113,1089,657,872]" box="[113,961,849,872]" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FCAAFEAB0629" bold="true" box="[113,260,849,872]" pageId="13" pageNumber="14">Distribution.</emphasis>
<collectingCountry id="F3143BAFC625FF868160FCAAFEF90629" box="[265,342,849,872]" name="Yemen" pageId="13" pageNumber="14">Yemen</collectingCountry>
,
<collectingCountry id="F3143BAFC625FF868108FCAAFE550629" box="[353,506,849,872]" name="Socotra Island" pageId="13" pageNumber="14">Socotra Island</collectingCountry>
(
<figureCitation id="133867BAC625FF868260FCAAFDF70629" box="[521,600,849,872]" captionStart="Figs 2729" captionStartId="16.[151,188,1531,1550]" captionTargetBox="[194,1079,235,1493]" captionTargetId="figure-11@16.[186,1091,219,1516]" captionTargetPageId="16" captionText="Figs 2729. Sketch maps of Socotra Island displaying the known and newly recorded distribution of Dermaptera. Published records are based on HAAS at al. (2004); the occurrence of Labidura riparia (Pallas, 1773) on Samha Island published by these authors is not displayed." figureDoi="http://doi.org/10.5281/zenodo.5312366" httpUri="https://zenodo.org/record/5312366/files/figure.png" pageId="13" pageNumber="14">Fig. 28</figureCitation>
). Most likely an endemic species.
</paragraph>
</subSubSection>
</treatment>
</document>