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<document id="3CE079E3D0033BD40A5E5FB60B8FC5BB" ID-CLB-Dataset="8478" ID-DOI="10.24349/acarologia/20184227" ID-GBIF-Dataset="08352714-1e98-42d5-9f93-e638394549c3" ID-ISSN="2107-7207" ID-Zenodo-Dep="4487537" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612205300746" checkinUser="felipe" docAuthor="Moraza, Maria L. &amp; Lindquist, Evert E." docDate="2018" docId="03A6A810FF8EFFC8FE11787726671A46" docLanguage="en" docName="Acarologia.58.1.62-98.pdf.imf" docOrigin="Acarologia 58 (1)" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Lasioseius cassidini Moraza &amp; Lindquist 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="74" masterDocId="FF9FD068FF86FFC5FFFA7B1325471C25" masterDocTitle="A new species-group with new species of the genus Lasioseius (Acari: Mesostigmata: Blattisociidae) associated with Neotropical hispine beetles in furled leaves ofHeliconia" masterLastPageNumber="98" masterPageNumber="62" pageNumber="69" updateTime="1698925921708" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C2A8458016091D9852FBA98412090BB5">A new species-group with new species of the genus Lasioseius (Acari: Mesostigmata: Blattisociidae) associated with Neotropical hispine beetles in furled leaves ofHeliconia</mods:title>
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<mods:namePart id="784B09FCF8CCA3BF6B8070AF01EA0C3A">Moraza, Maria L.</mods:namePart>
<mods:affiliation id="9A203C34E4F40CC2805E5DC97DA6FABD">Universidad de Navarra, Facultad de Ciencias, Departamento de Biología Ambiental, Campus Universitario, 31080, Pamplona España.</mods:affiliation>
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<mods:namePart id="1C7FD9920D8014469E36FCE8DB877756">Lindquist, Evert E.</mods:namePart>
<mods:affiliation id="898B8FD487CC2E7BA9ED6F0585334923">Canadian National Collection of Insects, Arachnids, and Nematodes, Science &amp; Technology Branch, Agriculture and Agri-Food Canada, Ottawa, ON, K 1 A 0 C 6, Canada.</mods:affiliation>
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<mods:title id="C9A29149117CF3EDED55CA1234175DF5">Acarologia</mods:title>
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<mods:date id="FA0F9B1853BAB23DBA959AD3591FB98E">2018</mods:date>
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<treatment id="03A6A810FF8EFFC8FE11787726671A46" ID-DOI="http://doi.org/10.5281/zenodo.4501358" ID-GBIF-Taxon="177209850" ID-Zenodo-Dep="4501358" LSID="urn:lsid:plazi:treatment:03A6A810FF8EFFC8FE11787726671A46" httpUri="http://treatment.plazi.org/id/03A6A810FF8EFFC8FE11787726671A46" lastPageId="13" lastPageNumber="74" pageId="8" pageNumber="69">
<subSubSection id="C3154A8DFF8EFFCDFE11787726A11F94" pageId="8" pageNumber="69" type="nomenclature">
<paragraph id="8BB01906FF8EFFCDFE11787720191FA5" blockId="8.[491,1374,868,896]" box="[491,1374,868,896]" pageId="8" pageNumber="69">
<heading id="D0F8AE6AFF8EFFCDFE11787720191FA5" bold="true" box="[491,1374,868,896]" fontSize="12" level="2" pageId="8" pageNumber="69" reason="6">
<emphasis id="B97BC514FF8EFFCDFE11787720191FA5" bold="true" box="[491,1374,868,896]" pageId="8" pageNumber="69">
<taxonomicName id="4C0F6285FF8EFFCDFE11787726471FA5" authority="Moraza &amp; Lindquist, 2018" authorityName="Moraza &amp; Lindquist" authorityYear="2018" box="[491,768,868,896]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="cassidini" status="sp. nov.">
<emphasis id="B97BC514FF8EFFCDFE11787726471FA5" bold="true" box="[491,768,868,896]" italics="true" pageId="8" pageNumber="69">Lasioseius cassidini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248786FFF8EFFCDFCF3787726111FA5" box="[777,854,868,896]" pageId="8" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13340583FF8EFFCDFC92787726AA1FA5" box="[872,1005,868,896]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="8" pageNumber="69">Figures 1</figureCitation>
,
<figureCitation id="13340583FF8EFFCDFC04787721541FA5" box="[1022,1043,868,896]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="8" pageNumber="69">2</figureCitation>
,
<figureCitation id="13340583FF8EFFCDFBDA7877211B1FA5" box="[1056,1116,868,896]" captionStart="Figure 11" captionStartId="29.[119,186,665,686]" captionTargetBox="[310,952,245,592]" captionTargetPageId="29" captionText="Figure 11 Distal subcapitular region, adult males: A L. cassidini; B L. serripes; C. — L. fuscina" figureDoi="http://doi.org/10.5281/zenodo.4487561" httpUri="https://zenodo.org/record/4487561/files/figure.png" pageId="8" pageNumber="69">11A</figureCitation>
,
<figureCitation id="13340583FF8EFFCDFB90787721E01FA5" box="[1130,1191,868,896]" captionStart="Figure 12" captionStartId="30.[119,186,1420,1441]" captionTargetBox="[294,1295,236,1343]" captionTargetId="graphics-3@30.[294,1280,237,1343]" captionTargetPageId="30" captionText="Figure 12 A-D Leg II, dorsal view (right leg, excluding tarsus, coxa and trochanter), adult males: A—Lasioseius serripes n. sp.; B L. serripes, detail of ventral setaeon genu and tibia III; C Lasioseius cassidini n. sp.; D Lasioseius fuscina n. sp. E, F Lasioseius duobtusisetis, adult female, tarsus I right leg, dorsal view: E General aspect without detail of dorsodistal sensorial area; F Detail of dorsodistal sensorial area. G Lasioseius serripes, protonymph, tarsus IV left leg; H — Lasioseius sp. undescribed. adult female, tarsus IV right leg, ventral view. (Solid black setae in figures A-G and solid black circles in figures G and H indicate dorsal setae)." figureDoi="http://doi.org/10.5281/zenodo.4487563" httpUri="https://zenodo.org/record/4487563/files/figure.png" pageId="8" pageNumber="69">12C</figureCitation>
,
<figureCitation id="13340583FF8EFFCDFB4E787721911FA5" box="[1204,1238,868,896]" captionStart="Figure 13" captionStartId="31.[119,186,1454,1475]" captionTargetBox="[311,1289,241,1396]" captionTargetPageId="31" captionText="Figure 13 Tarsus II, right leg. A, B, H.Lasioseius cassidini n. sp.: A Adult female, dorsal view; B Adult male, dorsal view; H — Detail of distal region, lateroventral view. C, D, G.Lasioseius serripes n. sp.: C Adult female, dorsal view; D Adult male, anterolateral view; G. Detail of distal region, ventral view. E, F.Lasioseius fuscina n. sp.: E Adult male, dorsal view; F Adult female, dorsal view. I —Lasioseius duobtusisetis n. sp., adult female, dorsal view. (Solid black color indicates dorsal setae on segment)." figureDoi="http://doi.org/10.5281/zenodo.4487565" httpUri="https://zenodo.org/record/4487565/files/figure.png" pageId="8" pageNumber="69">13</figureCitation>
A-B, 14A)
</emphasis>
</heading>
</paragraph>
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Zoobank:
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</paragraph>
</subSubSection>
<subSubSection id="C3154A8DFF8EFFCDFDE978F320D5199C" pageId="8" pageNumber="69" type="diagnosis">
<paragraph id="8BB01906FF8EFFCDFDE978F320D5199C" blockId="8.[491,1520,992,1465]" pageId="8" pageNumber="69">
Diagnosis — Adult female: Sternal shield weakly lineate along lateral margins, smooth medially, its posterolateral corners not emarginated; poroids
<emphasis id="B97BC514FF8EFFCDFB837F1121DE183F" box="[1145,1177,1026,1050]" italics="true" pageId="8" pageNumber="69">iv3</emphasis>
alone, on soft cuticle, setae
<emphasis id="B97BC514FF8EFFCDFE117F36274D1818" box="[491,522,1061,1085]" italics="true" pageId="8" pageNumber="69">st4</emphasis>
on tiny metasternal platelets; epigynal shield partially reticulated, its lateral margins concave, narrowed anterior to setae
<emphasis id="B97BC514FF8EFFCDFC967F5B26CD187A" box="[876,906,1096,1119]" italics="true" pageId="8" pageNumber="69">st5</emphasis>
; ventrianal shield with posterolateral margins on either side of anus divergent anteriorly, and with four pairs of opisthogastric setae,
<emphasis id="B97BC514FF8EFFCDFAD37F79201418A7" box="[1321,1363,1130,1154]" italics="true" pageId="8" pageNumber="69">JV1</emphasis>
on shield; metapodal plates well separated on each side. Dorsal setae
<emphasis id="B97BC514FF8EFFCDFB9E7F9E213B1880" box="[1124,1148,1165,1189]" italics="true" pageId="8" pageNumber="69">r6</emphasis>
on edge of dorsal shield;
<emphasis id="B97BC514FF8EFFCDFA6A7F9E20EB1881" box="[1424,1452,1165,1188]" italics="true" pageId="8" pageNumber="69">Z5</emphasis>
,
<emphasis id="B97BC514FF8EFFCDFA407F9E20931880" box="[1466,1492,1165,1189]" italics="true" pageId="8" pageNumber="69">S5</emphasis>
attenuated like adjacent setae. Adult male: Dorsal shield setae
<emphasis id="B97BC514FF8EFFCDFB877FBC21DF18E2" box="[1149,1176,1199,1223]" italics="true" pageId="8" pageNumber="69">S5</emphasis>
dimorphically less attenuated. Sternitigenital shield incompletely reticulated, nearly smooth medially. Tibiae III-IV with setae (
<emphasis id="B97BC514FF8EFFCDFE0E7FE627471929" box="[500,512,1269,1292]" italics="true" pageId="8" pageNumber="69">v</emphasis>
) dimorphically spinelike, blunt; genu IV
<emphasis id="B97BC514FF8EFFCDFC387FE6269C1929" box="[962,987,1269,1292]" italics="true" pageId="8" pageNumber="69">av</emphasis>
similarly so. Adult attributes: Gnathotectum anterior margin truncate, irregularly denticulate. Fixed chela with file of ca 20 teeth in female, ca
<quantity id="4CF7B4E3FF8EFFCDFDF17E2A27791974" box="[523,574,1337,1361]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="8" pageNumber="69" unit="in" value="18.0">18 in</quantity>
male. Sternal setae
<emphasis id="B97BC514FF8EFFCDFCDA7E2A26791974" box="[800,830,1337,1361]" italics="true" pageId="8" pageNumber="69">st1</emphasis>
inserted in weakly sclerotized presternal region. Tarsi II-IV with ventral apical process bluntly pointed, smooth apically; tarsi II-III with seta
<emphasis id="B97BC514FF8EFFCDFAB47E4F203A1951" box="[1358,1405,1372,1396]" italics="true" pageId="8" pageNumber="69">av-1</emphasis>
slightly longer than
<emphasis id="B97BC514FF8EFFCDFD967E6D27DD19B3" box="[620,666,1406,1430]" italics="true" pageId="8" pageNumber="69">pv-1</emphasis>
; pretarsi III-IV with anterior paradactylus asymmetrically elongate, twice as long as claws; tarsus IV with distal seta
<emphasis id="B97BC514FF8EFFCDFC6C7EB22682199C" box="[918,965,1441,1465]" italics="true" pageId="8" pageNumber="69">av-1</emphasis>
clearly larger, thicker than adjacent setae.
</paragraph>
</subSubSection>
<subSubSection id="C3154A8DFF8EFFC8FE117EF226671A46" lastPageId="13" lastPageNumber="74" pageId="8" pageNumber="69" type="description">
<paragraph id="8BB01906FF8EFFCDFE117EF2273919DE" blockId="8.[491,638,1505,1531]" box="[491,638,1505,1531]" pageId="8" pageNumber="69">
<heading id="D0F8AE6AFF8EFFCDFE117EF2273919DE" box="[491,638,1505,1531]" pageId="8" pageNumber="69">
<emphasis id="B97BC514FF8EFFCDFE117EF2273919DE" bold="true" box="[491,638,1505,1531]" pageId="8" pageNumber="69">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB01906FF8EFFCDFE117D0520E91BA2" blockId="8.[491,1505,1558,2031]" pageId="8" pageNumber="69">
Idiosomatic dorsum — Adult female. Dorsal shield 703 757 long, 375 410 wide at level of
<emphasis id="B97BC514FF8EFFCDFE117D2B27441A75" box="[491,515,1592,1616]" italics="true" pageId="8" pageNumber="69">r3</emphasis>
, 428 477 wide at
<emphasis id="B97BC514FF8EFFCDFD207D2B27B51A75" box="[730,754,1592,1616]" italics="true" pageId="8" pageNumber="69">r6</emphasis>
(n=9), lightly reticulated over entire surface, without puncta behind level of setae
<emphasis id="B97BC514FF8EFFCDFD787D4827D91A56" box="[642,670,1627,1651]" italics="true" pageId="8" pageNumber="69">Z4</emphasis>
; shield with 38 pairs of setae generally similar in size and acutely pointed form,
<emphasis id="B97BC514FF8EFFCDFDD77D6E272F1AB0" box="[557,616,1661,1685]" italics="true" pageId="8" pageNumber="69">J1-J4</emphasis>
about as long or slightly longer than longitudinal intervals between their bases;
<emphasis id="B97BC514FF8EFFCDFA447D6D20911AB0" box="[1470,1494,1662,1685]" italics="true" pageId="8" pageNumber="69">J5</emphasis>
minute, barbed (8 12):
<emphasis id="B97BC514FF8EFFCDFD037DB326491A9D" box="[761,782,1696,1720]" italics="true" pageId="8" pageNumber="69">j1</emphasis>
,
<emphasis id="B97BC514FF8EFFCDFCE17DB326771A9D" box="[795,816,1696,1720]" italics="true" pageId="8" pageNumber="69">j2</emphasis>
(45 60),
<emphasis id="B97BC514FF8EFFCDFC5D7DB326FB1A9D" box="[935,956,1696,1720]" italics="true" pageId="8" pageNumber="69">j3</emphasis>
(67 76),
<emphasis id="B97BC514FF8EFFCDFBC97DB3210F1A9D" box="[1075,1096,1696,1720]" italics="true" pageId="8" pageNumber="69">j4</emphasis>
(50 57),
<emphasis id="B97BC514FF8EFFCDFB457DB221931A9D" box="[1215,1236,1697,1720]" italics="true" pageId="8" pageNumber="69">j5</emphasis>
(56 62),
<emphasis id="B97BC514FF8EFFCDFAB67DB320271A9D" box="[1356,1376,1696,1720]" italics="true" pageId="8" pageNumber="69">j6</emphasis>
(70 77),
<emphasis id="B97BC514FF8EFFCDFE117DD127441AFF" box="[491,515,1730,1754]" italics="true" pageId="8" pageNumber="69">z1</emphasis>
(37 42),
<emphasis id="B97BC514FF8EFFCDFD817DD127D41AFF" box="[635,659,1730,1754]" italics="true" pageId="8" pageNumber="69">z2</emphasis>
(56 65),
<emphasis id="B97BC514FF8EFFCDFCF07DD126651AFF" box="[778,802,1730,1754]" italics="true" pageId="8" pageNumber="69">z3</emphasis>
(60 68),
<emphasis id="B97BC514FF8EFFCDFC637DD126F61AFF" box="[921,945,1730,1754]" italics="true" pageId="8" pageNumber="69">z4</emphasis>
(72 77),
<emphasis id="B97BC514FF8EFFCDFBD27DD021071AFF" box="[1064,1088,1731,1754]" italics="true" pageId="8" pageNumber="69">z5</emphasis>
(52 62),
<emphasis id="B97BC514FF8EFFCDFB427DD121881AFF" box="[1208,1231,1730,1754]" italics="true" pageId="8" pageNumber="69">z6</emphasis>
(62 74);
<emphasis id="B97BC514FF8EFFCDFAB27DD120271AFF" box="[1352,1376,1730,1754]" italics="true" pageId="8" pageNumber="69">s1</emphasis>
(58 75),
<emphasis id="B97BC514FF8EFFCDFE117DF627441AD8" box="[491,515,1765,1789]" italics="true" pageId="8" pageNumber="69">s2</emphasis>
(65 75),
<emphasis id="B97BC514FF8EFFCDFD867DF627D41AD8" box="[636,659,1765,1789]" italics="true" pageId="8" pageNumber="69">s3</emphasis>
(70 75),
<emphasis id="B97BC514FF8EFFCDFCF67DF626631AD8" box="[780,804,1765,1789]" italics="true" pageId="8" pageNumber="69">s4</emphasis>
(70 80),
<emphasis id="B97BC514FF8EFFCDFC677DF526F21AD8" box="[925,949,1766,1789]" italics="true" pageId="8" pageNumber="69">s5</emphasis>
(82 92),
<emphasis id="B97BC514FF8EFFCDFBD77DF621021AD8" box="[1069,1093,1765,1789]" italics="true" pageId="8" pageNumber="69">s6</emphasis>
(85 96),
<emphasis id="B97BC514FF8EFFCDFB447DF621911AD8" box="[1214,1238,1765,1789]" italics="true" pageId="8" pageNumber="69">r2</emphasis>
(62 74),
<emphasis id="B97BC514FF8EFFCDFAB47DF620211AD8" box="[1358,1382,1765,1789]" italics="true" pageId="8" pageNumber="69">r3</emphasis>
erect (60 67),
<emphasis id="B97BC514FF8EFFCDFDCB7C1B270F1B05" box="[561,584,1800,1824]" italics="true" pageId="8" pageNumber="69">r4</emphasis>
(75 80),
<emphasis id="B97BC514FF8EFFCDFD3A7C1B27901B3A" box="[704,727,1800,1823]" italics="true" pageId="8" pageNumber="69">r5</emphasis>
(75 82),
<emphasis id="B97BC514FF8EFFCDFCB57C1B26201B05" box="[847,871,1800,1824]" italics="true" pageId="8" pageNumber="69">r6</emphasis>
(70 79),
<emphasis id="B97BC514FF8EFFCDFC247C1B26B01B05" box="[990,1015,1800,1824]" italics="true" pageId="8" pageNumber="69">J1</emphasis>
(65 75),
<emphasis id="B97BC514FF8EFFCDFB947C1B21C01B05" box="[1134,1159,1800,1824]" italics="true" pageId="8" pageNumber="69">J2</emphasis>
(68 77),
<emphasis id="B97BC514FF8EFFCDFB047C1B20501B05" box="[1278,1303,1800,1824]" italics="true" pageId="8" pageNumber="69">J3</emphasis>
(67 82),
<emphasis id="B97BC514FF8EFFCDFA747C1B20E01B05" box="[1422,1447,1800,1824]" italics="true" pageId="8" pageNumber="69">J4</emphasis>
(70 75),
<emphasis id="B97BC514FF8EFFCDFDCB7C39270A1B67" box="[561,589,1834,1858]" italics="true" pageId="8" pageNumber="69">Z1</emphasis>
,
<emphasis id="B97BC514FF8EFFCDFDA67C39273F1B67" box="[604,632,1834,1858]" italics="true" pageId="8" pageNumber="69">Z2</emphasis>
(92 103),
<emphasis id="B97BC514FF8EFFCDFD077C39265E1B67" box="[765,793,1834,1858]" italics="true" pageId="8" pageNumber="69">Z3</emphasis>
(100 112),
<emphasis id="B97BC514FF8EFFCDFC507C3926811B67" box="[938,966,1834,1858]" italics="true" pageId="8" pageNumber="69">Z4</emphasis>
(107 122),
<emphasis id="B97BC514FF8EFFCDFBA37C3821321B67" box="[1113,1141,1835,1858]" italics="true" pageId="8" pageNumber="69">Z5</emphasis>
(93 110),
<emphasis id="B97BC514FF8EFFCDFB037C3920531B67" box="[1273,1300,1834,1858]" italics="true" pageId="8" pageNumber="69">S1</emphasis>
(77 95),
<emphasis id="B97BC514FF8EFFCDFA767C3920E01B67" box="[1420,1447,1834,1858]" italics="true" pageId="8" pageNumber="69">S2</emphasis>
,
<emphasis id="B97BC514FF8EFFCDFA4E7C3920881B67" box="[1460,1487,1834,1858]" italics="true" pageId="8" pageNumber="69">S3</emphasis>
(88 103),
<emphasis id="B97BC514FF8EFFCDFD927C5E27C41B40" box="[616,643,1869,1893]" italics="true" pageId="8" pageNumber="69">S4</emphasis>
(97 110),
<emphasis id="B97BC514FF8EFFCDFCFF7C5E26671B40" box="[773,800,1869,1893]" italics="true" pageId="8" pageNumber="69">S5</emphasis>
(82 105). Lateral soft cuticle with seven pairs of smooth setae, including
<emphasis id="B97BC514FF8EFFCDFDAC7C7C27DC1BA2" box="[598,667,1903,1927]" italics="true" pageId="8" pageNumber="69">R1-R3</emphasis>
(48 62),
<emphasis id="B97BC514FF8EFFCDFCEA7C7C26131BA2" box="[784,852,1903,1927]" italics="true" pageId="8" pageNumber="69">R4-R6</emphasis>
(42 53) and
<emphasis id="B97BC514FF8EFFCDFC0A7C63215B1BA2" box="[1008,1052,1904,1927]" italics="true" pageId="8" pageNumber="69">ZV5</emphasis>
(47 55) posteriorly (
<figureCitation id="13340583FF8EFFCDFAFB7C7C20E61BA2" box="[1281,1441,1903,1927]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="8" pageNumber="69">Figs. 1A, B, C</figureCitation>
).
</paragraph>
<paragraph id="8BB01906FF8EFFCFFDE97C8121B11BA2" blockId="8.[491,1505,1558,2031]" lastBlockId="10.[491,1497,1730,1927]" lastPageId="10" lastPageNumber="71" pageId="8" pageNumber="69">
Adult male. Dorsal shield 493 610 long, 315 342 wide at level of setae
<emphasis id="B97BC514FF8EFFCDFAC47C8120121B8F" box="[1342,1365,1938,1962]" italics="true" pageId="8" pageNumber="69">r3</emphasis>
, 345 367 wide at
<emphasis id="B97BC514FF8EFFCDFDBA7CA7271F1BE9" box="[576,600,1972,1996]" italics="true" pageId="8" pageNumber="69">r6</emphasis>
(n = 8); shield ornamentation, complement of pore-like structures and setation as in female; posterolateral setae attenuated as in female. Setae
<emphasis id="B97BC514FF8EFFCDFB9A7CC421321BCA" box="[1120,1141,2007,2031]" italics="true" pageId="8" pageNumber="69">j1</emphasis>
(40 50),
<emphasis id="B97BC514FF8EFFCDFB177CC420451BCA" box="[1261,1282,2007,2031]" italics="true" pageId="8" pageNumber="69">j2</emphasis>
(35 47),
<emphasis id="B97BC514FF8EFFCDFA807CC420C91BCA" box="[1402,1422,2007,2031]" italics="true" pageId="8" pageNumber="69">j3</emphasis>
(50 60),
<emphasis id="B97BC514FF8CFFCFFDE67DD127771AFF" box="[540,560,1730,1754]" italics="true" pageId="10" pageNumber="71">j4</emphasis>
(38 45),
<emphasis id="B97BC514FF8CFFCFFD5C7DD027FD1AFF" box="[678,698,1731,1754]" italics="true" pageId="10" pageNumber="71">j5</emphasis>
(43 47),
<emphasis id="B97BC514FF8CFFCFFCD57DD126041AFF" box="[815,835,1730,1754]" italics="true" pageId="10" pageNumber="71">j6</emphasis>
(53 62),
<emphasis id="B97BC514FF8CFFCFFC437DD126971AFF" box="[953,976,1730,1754]" italics="true" pageId="10" pageNumber="71">z1</emphasis>
(28 38),
<emphasis id="B97BC514FF8CFFCFFBBC7DD121191AFF" box="[1094,1118,1730,1754]" italics="true" pageId="10" pageNumber="71">z2</emphasis>
,
<emphasis id="B97BC514FF8CFFCFFB907DD121C51AFF" box="[1130,1154,1730,1754]" italics="true" pageId="10" pageNumber="71">z3</emphasis>
(43 55),
<emphasis id="B97BC514FF8CFFCFFB0D7DD120491AFF" box="[1271,1294,1730,1754]" italics="true" pageId="10" pageNumber="71">z4</emphasis>
(52 62),
<emphasis id="B97BC514FF8CFFCFFA797DD020DC1AFF" box="[1411,1435,1731,1754]" italics="true" pageId="10" pageNumber="71">z5</emphasis>
(40 45),
<emphasis id="B97BC514FF8CFFCFFDE67DF627741AD8" box="[540,563,1765,1789]" italics="true" pageId="10" pageNumber="71">z6</emphasis>
(48 55),
<emphasis id="B97BC514FF8CFFCFFD527DF627F81AD8" box="[680,703,1765,1789]" italics="true" pageId="10" pageNumber="71">s1</emphasis>
(47 57),
<emphasis id="B97BC514FF8CFFCFFCC97DF6262D1AD8" box="[819,874,1765,1789]" italics="true" pageId="10" pageNumber="71">s2-s4</emphasis>
(50 65),
<emphasis id="B97BC514FF8CFFCFFC247DF526B11AD8" box="[990,1014,1766,1789]" italics="true" pageId="10" pageNumber="71">s5</emphasis>
,
<emphasis id="B97BC514FF8CFFCFFBF97DF6215C1AD8" box="[1027,1051,1765,1789]" italics="true" pageId="10" pageNumber="71">s6</emphasis>
(60 75),
<emphasis id="B97BC514FF8CFFCFFB757DF621E01AD8" box="[1167,1191,1765,1789]" italics="true" pageId="10" pageNumber="71">r2</emphasis>
(50 60),
<emphasis id="B97BC514FF8CFFCFFAE07DF620751AD8" box="[1306,1330,1765,1789]" italics="true" pageId="10" pageNumber="71">r3</emphasis>
erect (38 55),
<emphasis id="B97BC514FF8CFFCFFE117C1427441B3A" box="[491,515,1799,1823]" italics="true" pageId="10" pageNumber="71">r4</emphasis>
,
<emphasis id="B97BC514FF8CFFCFFDEB7C1B276F1B3A" box="[529,552,1800,1823]" italics="true" pageId="10" pageNumber="71">r5</emphasis>
(53 67),
<emphasis id="B97BC514FF8CFFCFFD657C1427F01B3A" box="[671,695,1799,1823]" italics="true" pageId="10" pageNumber="71">r6</emphasis>
(45 57),
<emphasis id="B97BC514FF8CFFCFFCD47C1426291B3A" box="[814,878,1799,1823]" italics="true" pageId="10" pageNumber="71">J1, J2</emphasis>
(43 55),
<emphasis id="B97BC514FF8CFFCFFC1E7C1426B91B3A" box="[996,1022,1799,1823]" italics="true" pageId="10" pageNumber="71">J3</emphasis>
(50 62),
<emphasis id="B97BC514FF8CFFCFFB8E7C1421CA1B3A" box="[1140,1165,1799,1823]" italics="true" pageId="10" pageNumber="71">J4</emphasis>
(40 47),
<emphasis id="B97BC514FF8CFFCFFAFE7C1B205A1B3A" box="[1284,1309,1800,1823]" italics="true" pageId="10" pageNumber="71">J5</emphasis>
(5 9),
<emphasis id="B97BC514FF8CFFCFFA837C1420D21B3A" box="[1401,1429,1799,1823]" italics="true" pageId="10" pageNumber="71">Z1</emphasis>
(63 80), Z2 (65 92),
<emphasis id="B97BC514FF8CFFCFFD487C3927891B67" box="[690,718,1834,1858]" italics="true" pageId="10" pageNumber="71">Z3</emphasis>
(66 98),
<emphasis id="B97BC514FF8CFFCFFCBF7C3826261B67" box="[837,865,1835,1858]" italics="true" pageId="10" pageNumber="71">Z5</emphasis>
(61 77),
<emphasis id="B97BC514FF8CFFCFFC227C3926B41B67" box="[984,1011,1834,1858]" italics="true" pageId="10" pageNumber="71">S1</emphasis>
(54 72),
<emphasis id="B97BC514FF8CFFCFFB907C3921C21B67" box="[1130,1157,1834,1858]" italics="true" pageId="10" pageNumber="71">S2</emphasis>
(55 77),
<emphasis id="B97BC514FF8CFFCFFB067C3920501B67" box="[1276,1303,1834,1858]" italics="true" pageId="10" pageNumber="71">S3</emphasis>
,
<emphasis id="B97BC514FF8CFFCFFADF7C3920781B67" box="[1317,1343,1834,1858]" italics="true" pageId="10" pageNumber="71">S4</emphasis>
(60 78),
<emphasis id="B97BC514FF8CFFCFFA4D7C3920961B67" box="[1463,1489,1834,1858]" italics="true" pageId="10" pageNumber="71">S5</emphasis>
(50 70),
<emphasis id="B97BC514FF8CFFCFFDA07C5E27311B40" box="[602,630,1869,1893]" italics="true" pageId="10" pageNumber="71">Z4</emphasis>
longest (80 102) of opisthonotal setae. Lateral soft cuticle with five or six pairs of
<emphasis id="B97BC514FF8CFFCFFDF27C6327661BA2" box="[520,545,1904,1927]" italics="true" pageId="10" pageNumber="71">R-</emphasis>
setae (33 48) (
<figureCitation id="13340583FF8CFFCFFD3C7C7C265C1BA2" box="[710,795,1903,1927]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="10" pageNumber="71">Fig. 2A</figureCitation>
),
<emphasis id="B97BC514FF8CFFCFFCC47C7C261C1BA2" box="[830,859,1903,1927]" italics="true" pageId="10" pageNumber="71">R2</emphasis>
sometimes on small platelet (
<figureCitation id="13340583FF8CFFCFFB637C7C21AC1BA2" box="[1177,1259,1903,1927]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="10" pageNumber="71">Fig. 2E</figureCitation>
).
</paragraph>
<caption id="DF70498EFF8FFFCCFF8D7DF424781B0F" ID-DOI="http://doi.org/10.5281/zenodo.4487539" ID-Zenodo-Dep="4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="9" pageNumber="70" startId="9.[119,186,1767,1788]" targetBox="[305,1283,400,1668]" targetPageId="9">
<paragraph id="8BB01906FF8FFFCCFF8D7DF424781B0F" blockId="9.[119,1513,1767,1834]" pageId="9" pageNumber="70">
<emphasis id="B97BC514FF8FFFCCFF8D7DF4258B1AD9" bold="true" box="[119,204,1767,1788]" pageId="9" pageNumber="70">Figure 1</emphasis>
<taxonomicName id="4C0F6285FF8FFFCCFF227DFB24DF1AD8" authority="Moraza &amp; Lindquist, 2018" authorityName="Moraza &amp; Lindquist" authorityYear="2018" box="[216,408,1767,1789]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="cassidini" status="sp. nov.">
<emphasis id="B97BC514FF8FFFCCFF227DFB24DF1AD8" box="[216,408,1767,1789]" italics="true" pageId="9" pageNumber="70">Lasioseius cassidini</emphasis>
</taxonomicName>
<emphasis id="B97BC514FF8FFFCCFE647DF424911AD9" bold="true" box="[414,470,1767,1788]" pageId="9" pageNumber="70">
<taxonomicNameLabel id="A248786FFF8FFFCCFE647DF424911AD9" box="[414,470,1767,1788]" pageId="9" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view.
</paragraph>
</caption>
<caption id="DF70498EFF8CFFCFFF8D7D1920AB1A69" ID-DOI="http://doi.org/10.5281/zenodo.4487541" ID-Zenodo-Dep="4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="10" pageNumber="71" startId="10.[119,186,1546,1567]" targetBox="[318,1281,249,1452]" targetPageId="10">
<paragraph id="8BB01906FF8CFFCFFF8D7D1920AB1A69" blockId="10.[119,1516,1545,1612]" pageId="10" pageNumber="71">
<emphasis id="B97BC514FF8CFFCFFF8D7D19258A1A3A" bold="true" box="[119,205,1546,1567]" pageId="10" pageNumber="71">Figure 2</emphasis>
<taxonomicName id="4C0F6285FF8CFFCFFF207D1924DC1A3A" authority="Moraza &amp; Lindquist, 2018" authorityName="Moraza &amp; Lindquist" authorityYear="2018" box="[218,411,1545,1567]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="71" phylum="Arthropoda" rank="species" species="cassidini" status="sp. nov.">
<emphasis id="B97BC514FF8CFFCFFF207D1924DC1A3A" box="[218,411,1545,1567]" italics="true" pageId="10" pageNumber="71">Lasioseius cassidini</emphasis>
</taxonomicName>
<emphasis id="B97BC514FF8CFFCFFE5B7D19249B1A3A" bold="true" box="[417,476,1546,1567]" pageId="10" pageNumber="71">
<taxonomicNameLabel id="A248786FFF8CFFCFFE5B7D19249B1A3A" box="[417,476,1546,1567]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showing
<emphasis id="B97BC514FF8CFFCFFBFA7D33215D1A13" box="[1024,1050,1568,1590]" italics="true" pageId="10" pageNumber="71">R2</emphasis>
on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view.
</paragraph>
</caption>
<paragraph id="8BB01906FF8CFFCEFE117CA72698181F" blockId="10.[491,1503,1972,2031]" lastBlockId="11.[491,1523,229,1669]" lastPageId="11" lastPageNumber="72" pageId="10" pageNumber="71">
Idiosomatic venter — Adult female. Tritosternum base sometimes biarticulate, 38 43 long and 20 22 wide at its base, with paired laciniae fused for about 22 25 of entire length (122 135). Presternal region with circumscribed pair of weakly granulated, lineated areas, flanked anteriorly by a pair of weak or distinct narrow strips on either side of tritosternal base, and with setae
<emphasis id="B97BC514FF8DFFCEFD727A3827E01D66" box="[648,679,299,323]" italics="true" pageId="11" pageNumber="72">st1</emphasis>
(33 37) inserted on this region, anterior to sternal shield. Sternal shield 127 135 long from anterior margin at transverse level with poroids
<emphasis id="B97BC514FF8DFFCEFB707A5E21ED1D40" box="[1162,1194,333,357]" italics="true" pageId="11" pageNumber="72">iv1</emphasis>
to slightly concave posterior margin, 125 136 wide, weakly lineated along lateral margins, smooth medially; sternal setae
<emphasis id="B97BC514FF8DFFCEFE117A81274C1D8F" box="[491,523,402,426]" italics="true" pageId="11" pageNumber="72">st2</emphasis>
,
<emphasis id="B97BC514FF8DFFCEFDE37A81277F1D8F" box="[537,568,402,426]" italics="true" pageId="11" pageNumber="72">st3</emphasis>
of similar length (30 37). Sternal poroids
<emphasis id="B97BC514FF8DFFCEFBE07A81217C1D8F" box="[1050,1083,402,426]" italics="true" pageId="11" pageNumber="72">iv3</emphasis>
alone on soft cuticle, setae
<emphasis id="B97BC514FF8DFFCEFA927A8120C01D8F" box="[1384,1415,402,426]" italics="true" pageId="11" pageNumber="72">st4</emphasis>
(28 35) on tiny metasternal platelets (13 15 x 8 11). Endopodal strips narrow between coxae III and IV. Epigynal shield sparsely reticulated, its broadly rounded hyaline anterior margin not overlapping posterior edge of sternal shield, its lateral margins slightly concave, narrowed anterior to setae
<emphasis id="B97BC514FF8DFFCEFD61790E27FD1E11" box="[667,698,541,564]" italics="true" pageId="11" pageNumber="72">st5</emphasis>
(38 47), and its posterolateral corners emarginated behind
<emphasis id="B97BC514FF8DFFCEFAC4790E201A1E11" box="[1342,1373,541,564]" italics="true" pageId="11" pageNumber="72">st5</emphasis>
; shield 208 225 long, 118 137 at widest level at posterior rounded corners, 80 87 at narrowest width between legs IV. Opisthosomatic venter with two pairs of well separated metapodal platelets, primary pair elongate-oval, 33 40 long, 15 19 wide, secondary pair small, 10 16 by 5 7. Ventrianal shield inversely subtriangular with angulose or rounded anterior corners, with posterolateral margins on either side of anus divergent anteriorly; surface entirely reticulate, and punctate on either side of postanal seta; shield wider (293 337) than long (212 232), width at anal level (130 155); poroids
<emphasis id="B97BC514FF8DFFCEFC61781D268E1F03" box="[923,969,782,806]" italics="true" pageId="11" pageNumber="72">ivo2</emphasis>
on shield anterior edge behind setae
<emphasis id="B97BC514FF8DFFCEFAA2781D20C21F03" box="[1368,1413,782,806]" italics="true" pageId="11" pageNumber="72">ZV1</emphasis>
; gland pore
<emphasis id="B97BC514FF8DFFCEFDDB7822270F1F6C" box="[545,584,817,841]" italics="true" pageId="11" pageNumber="72">gv3</emphasis>
on or off shield posterior to anal opening; shield with four pairs of ventral setae:
<emphasis id="B97BC514FF8DFFCEFA57782220911F6C" box="[1453,1494,817,841]" italics="true" pageId="11" pageNumber="72">JV1</emphasis>
(37 42),
<emphasis id="B97BC514FF8DFFCEFDA2784027C61F4E" box="[600,641,851,875]" italics="true" pageId="11" pageNumber="72">JV2</emphasis>
(43 50),
<emphasis id="B97BC514FF8DFFCEFD0F784026591F4E" box="[757,798,851,875]" italics="true" pageId="11" pageNumber="72">JV3</emphasis>
(44 55),
<emphasis id="B97BC514FF8DFFCEFC6B784026FA1F4E" box="[913,957,851,875]" italics="true" pageId="11" pageNumber="72">ZV2</emphasis>
(35 48); with paranal setae (21 27) shorter than postanal seta (43 65). Soft cuticle with five pairs of simple opisthogastric setae,
<emphasis id="B97BC514FF8DFFCEFABD786520331FAB" box="[1351,1396,886,910]" italics="true" pageId="11" pageNumber="72">ZV1</emphasis>
(35 44) anterior to ventrianal shield,
<emphasis id="B97BC514FF8DFFCEFCDA788B260B1F95" box="[800,844,920,944]" italics="true" pageId="11" pageNumber="72">ZV3</emphasis>
(33 43),
<emphasis id="B97BC514FF8DFFCEFC3E788B26B71F95" box="[964,1008,920,944]" italics="true" pageId="11" pageNumber="72">ZV4</emphasis>
(37 44),
<emphasis id="B97BC514FF8DFFCEFB92788A21D61F95" box="[1128,1169,921,944]" italics="true" pageId="11" pageNumber="72">JV5</emphasis>
(55 67) flanking the shield, and
<emphasis id="B97BC514FF8DFFCEFDE278A827021FF7" box="[536,581,955,978]" italics="true" pageId="11" pageNumber="72">ZV5</emphasis>
aligned behind
<emphasis id="B97BC514FF8DFFCEFD1478A8264C1FF6" box="[750,779,955,979]" italics="true" pageId="11" pageNumber="72">R6</emphasis>
; three pairs of poroids on soft cuticle. Peritrematal-exopodal shield with lines extending from stigma into rounded posterior margin (
<figureCitation id="13340583FF8DFFCEFB5578CE20441FD0" box="[1199,1283,989,1013]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Fig. 1C</figureCitation>
); peritreme extending to level between setae
<emphasis id="B97BC514FF8DFFCEFD207F1327B5183D" box="[730,754,1024,1048]" italics="true" pageId="11" pageNumber="72">s1</emphasis>
and
<emphasis id="B97BC514FF8DFFCEFCDF7F13267B183D" box="[805,828,1024,1048]" italics="true" pageId="11" pageNumber="72">z1</emphasis>
(
<figureCitation id="13340583FF8DFFCEFCB17F1326E6183D" box="[843,929,1024,1048]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Fig. 1A</figureCitation>
). Spermathecal apparatus with cup-like calyx 28 29 long to embolus, 30 31 including it (
<figureCitation id="13340583FF8DFFCEFC7A7F312695181F" box="[896,978,1058,1082]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="8BB01906FF8DFFCEFDE97F56209E1AA0" blockId="11.[491,1523,229,1669]" pageId="11" pageNumber="72">
Adult male. Tritosternum much as in female, but with base biarticulate, longer (20 32) than wide (15 19), and with laciniae fused for only about 10 15 of entire length (88 107). Presternal region faintly sclerotized, with a pair of distinct narrow strips on either side of tritosternal base, and with setae
<emphasis id="B97BC514FF8DFFCEFCA47FBF263A18E1" box="[862,893,1196,1220]" italics="true" pageId="11" pageNumber="72">st1</emphasis>
inserted on this region on either side of genital opening. Sternitigenital shield length 225 267, narrowest widths 88 113 at level between coxae II, 77 95 between coxae III, 58 87 between coxae IV; shield slightly lineated along lateral margins and posteriorly between coxae IV, unornamented medially, with endopodal apices freely acuminate beside coxae IV; setae
<emphasis id="B97BC514FF8DFFCEFC677E2426FB196A" box="[925,956,1335,1359]" italics="true" pageId="11" pageNumber="72">st1</emphasis>
(28 32) and
<emphasis id="B97BC514FF8DFFCEFBA77E24213B196B" box="[1117,1148,1335,1358]" italics="true" pageId="11" pageNumber="72">st5</emphasis>
(28 37) longer,
<emphasis id="B97BC514FF8DFFCEFABB7E2420C0196A" box="[1345,1415,1335,1359]" italics="true" pageId="11" pageNumber="72">st2-st4</emphasis>
(17 26); poroids
<emphasis id="B97BC514FF8DFFCEFD8F7E4A27D21954" box="[629,661,1369,1393]" italics="true" pageId="11" pageNumber="72">iv3</emphasis>
sometimes hardly discernible flanking faint pair of pits (
<figureCitation id="13340583FF8DFFCEFAFD7E4A201A1954" box="[1287,1373,1369,1393]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="11" pageNumber="72">Fig. 2A</figureCitation>
). Ventrianal shield well reticulated and with punctate fields in anal region as in female, its convex anterior margin sometimes excised near apices of exopodal strips behind coxae IV, and posterolateral margins slightly constricted in anal region; shield much wider (278 337) than long (170 228, including cribrum ca. 8 11 long, 58 65 wide); shield with six or usually seven pairs of opisthogastric setae,
<emphasis id="B97BC514FF8DFFCEFD337D1427B51A3B" box="[713,754,1543,1566]" italics="true" pageId="11" pageNumber="72">JV5</emphasis>
asymmetrically on or off shield (
<figureCitation id="13340583FF8DFFCEFBA97D1521941A3B" box="[1107,1235,1542,1566]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Figs. 1A, B</figureCitation>
);
<emphasis id="B97BC514FF8DFFCEFB1E7D15204A1A3B" box="[1252,1293,1542,1566]" italics="true" pageId="11" pageNumber="72">JV1</emphasis>
and
<emphasis id="B97BC514FF8DFFCEFAC57D15202B1A3B" box="[1343,1388,1542,1566]" italics="true" pageId="11" pageNumber="72">ZV1</emphasis>
(23 32),
<emphasis id="B97BC514FF8DFFCEFE117D3B27531A65" box="[491,532,1576,1600]" italics="true" pageId="11" pageNumber="72">JV2</emphasis>
(28 37),
<emphasis id="B97BC514FF8DFFCEFD7D7D3B27F41A65" box="[647,691,1576,1600]" italics="true" pageId="11" pageNumber="72">ZV2</emphasis>
(28 32),
<emphasis id="B97BC514FF8DFFCEFCDC7D3B26171A65" box="[806,848,1576,1600]" italics="true" pageId="11" pageNumber="72">JV3</emphasis>
(28 42),
<emphasis id="B97BC514FF8DFFCEFC387D3B26A91A65" box="[962,1006,1576,1600]" italics="true" pageId="11" pageNumber="72">ZV3</emphasis>
(23 30),
<emphasis id="B97BC514FF8DFFCEFB9A7D3A21CD1A65" box="[1120,1162,1577,1600]" italics="true" pageId="11" pageNumber="72">JV5</emphasis>
(39 50); postanal seta smooth (39 47), about twice as long as paranal setae (15 22). Soft opisthogastric cuticle with two pairs of setae,
<emphasis id="B97BC514FF8DFFCEFD7E7D7E27F71AA0" box="[644,688,1645,1669]" italics="true" pageId="11" pageNumber="72">ZV4</emphasis>
(26 30), and
<emphasis id="B97BC514FF8DFFCEFCAA7D7D263B1AA0" box="[848,892,1646,1669]" italics="true" pageId="11" pageNumber="72">ZV5</emphasis>
(30 42) aligned behind
<emphasis id="B97BC514FF8DFFCEFB767D7E21EE1AA0" box="[1164,1193,1645,1669]" italics="true" pageId="11" pageNumber="72">R6</emphasis>
, one pair of lyrifissures,
<emphasis id="B97BC514FF8DFFCEFA487D7D20951AA0" box="[1458,1490,1646,1669]" italics="true" pageId="11" pageNumber="72">ivp</emphasis>
.
</paragraph>
<paragraph id="8BB01906FF8DFFC9FE117DA026651D05" blockId="11.[491,1513,1715,2015]" lastBlockId="12.[491,1523,229,701]" lastPageId="12" lastPageNumber="73" pageId="11" pageNumber="72">
Gnathosoma — Adult female. Gnathotectum anterior margin broadly truncate, with two lateral smooth prongs and serrate medial edge (
<figureCitation id="13340583FF8DFFCEFBFF7DC621271AC8" box="[1029,1120,1749,1773]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Fig. 1D</figureCitation>
). Cheliceral shaft, excluding basal section, 160 173 long, with slender digits; dorsal seta blunt (13 14); fixed digit with tiny offset subapical tooth and row of 18-22 very fine teeth (
<figureCitation id="13340583FF8DFFCEFBB77C0921EF1B17" box="[1101,1192,1818,1842]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Fig. 1G</figureCitation>
), pilus dentilus minute, at level between 4
<superScript id="7C7AB44EFF8DFFCEFD697C2A27EB1B71" attach="right" box="[659,684,1849,1876]" fontSize="7" pageId="11" pageNumber="72">th-</superScript>
5
<superScript id="7C7AB44EFF8DFFCEFD3A7C2A27941B6F" attach="right" box="[704,723,1849,1866]" fontSize="7" pageId="11" pageNumber="72">th</superScript>
apical teeth; movable digit (71 75) tridentate (
<figureCitation id="13340583FF8DFFCEFB267C2E20731B70" box="[1244,1332,1853,1877]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Fig. 1H</figureCitation>
). Subcapitulum rows of deutosternal denticles progressively slightly wider posteriorly, each with numerous (ca 30-35) fine denticles, 6th and sometimes 7th widest and with ca 40-45 and 35-40 teeth, respectively (
<figureCitation id="13340583FF8DFFCEFD8C7CB7278E1B99" box="[630,713,1956,1980]" captionStart="Figure 1" captionStartId="9.[119,186,1767,1788]" captionTargetBox="[305,1283,400,1668]" captionTargetId="graphics-3@9.[305,1283,400,1667]" captionTargetPageId="9" captionText="Figure 1 Lasioseius cassidini n. sp. adult female: A Idiosoma, dorsal view; B Detail of denoted dorsal setae; C Idiosoma, ventral view; D Gnathotectum; E Subcapitulum; F Spermathecal structures; G Fixed cheliceral digit in ventral view; H movable cheliceral digit in partially lateral view." figureDoi="http://doi.org/10.5281/zenodo.4487539" httpUri="https://zenodo.org/record/4487539/files/figure.png" pageId="11" pageNumber="72">Fig. 1E</figureCitation>
); subcapitular setae
<emphasis id="B97BC514FF8DFFCEFC527CB726971B99" box="[936,976,1956,1980]" italics="true" pageId="11" pageNumber="72">hp1</emphasis>
(40 45) slightly shorter than
<emphasis id="B97BC514FF8DFFCEFAE07CB720041B99" box="[1306,1347,1956,1980]" italics="true" pageId="11" pageNumber="72">hp3</emphasis>
(48 52) and capitular seta
<emphasis id="B97BC514FF8DFFCEFD847CDB27DE1BFA" box="[638,665,1992,2015]" italics="true" pageId="11" pageNumber="72">pc</emphasis>
(43 52),
<emphasis id="B97BC514FF8DFFCEFCF07CD426741BFA" box="[778,819,1991,2015]" italics="true" pageId="11" pageNumber="72">hp3</emphasis>
much longer than
<emphasis id="B97BC514FF8DFFCEFC0C7CD421581BFA" box="[1014,1055,1991,2015]" italics="true" pageId="11" pageNumber="72">hp2</emphasis>
(23 27). Palp length (195 212); apical margin of palptrochanter with a single spine anterolaterally; palptrochanter setae dissimilar in length,
<emphasis id="B97BC514FF8AFFC9FDC17A1B27131D05" box="[571,596,264,288]" italics="true" pageId="12" pageNumber="73">v1</emphasis>
48 52,
<emphasis id="B97BC514FF8AFFC9FD427A1B27961D05" box="[696,721,264,288]" italics="true" pageId="12" pageNumber="73">v2</emphasis>
38 42.
</paragraph>
<paragraph id="8BB01906FF8AFFC9FDE97A3926781E98" blockId="12.[491,1523,229,701]" pageId="12" pageNumber="73">
Adult male. Gnathotectum as in female (
<figureCitation id="13340583FF8AFFC9FC297A39216F1D67" box="[979,1064,298,322]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="12" pageNumber="73">Fig. 2C</figureCitation>
). Cheliceral shaft, excluding short basal section 105 130 long; fixed digit with offset subapical tooth followed by minute pilus dentilis inserted on small angular protrusion and row of 11-16 small teeth (
<figureCitation id="13340583FF8AFFC9FB3B7A7C20511DA2" box="[1217,1302,367,391]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="12" pageNumber="73">Fig. 2H</figureCitation>
); movable digit (40 45) unidentate; spermatodactyl 45 54 long, 8 9 wide at medial region, extending anteriorly 28 37 beyond apex of digit, with narrowed neck and small bifid head apically (
<figureCitation id="13340583FF8AFFC9FA9B7AA7209E1DE9" box="[1377,1497,436,460]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="12" pageNumber="73">Figs. 2F, G</figureCitation>
). Corniculi more widely spaced but not more arched than in female, 33 37 long, and flanked medially by pair of short (7-10) spiculate processes (
<figureCitation id="13340583FF8AFFC9FBE37AEA21291E34" box="[1049,1134,505,529]" captionStart="Figure 2" captionStartId="10.[119,186,1546,1567]" captionTargetBox="[318,1281,249,1452]" captionTargetId="graphics-4@10.[318,1281,245,1451]" captionTargetPageId="10" captionText="Figure 2 Lasioseius cassidini n. sp., adult male: A Idiosoma, ventral view; B Variant form of ventrianal shield; C Gnathotectum; D Subcapitulum and ventral setae of palptrochanter; E Detail of denoted dorsal setae, showingR2 on variant platelet; F Cheliceral digits and spermatodactyl, dorsal view; G Cheliceral movable digit, spermatodactyl, and corniculi, dorsal view; H Detail of fixed digit, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4487541" httpUri="https://zenodo.org/record/4487541/files/figure.png" pageId="12" pageNumber="73">Figs 2D</figureCitation>
,
<figureCitation id="13340583FF8AFFC9FB827AEA21EF1E34" box="[1144,1192,505,529]" captionStart="Figure 11" captionStartId="29.[119,186,665,686]" captionTargetBox="[310,952,245,592]" captionTargetPageId="29" captionText="Figure 11 Distal subcapitular region, adult males: A L. cassidini; B L. serripes; C. — L. fuscina" figureDoi="http://doi.org/10.5281/zenodo.4487561" httpUri="https://zenodo.org/record/4487561/files/figure.png" pageId="12" pageNumber="73">11A</figureCitation>
); internal malae and labrum as in female. Subcapitulum with seven rows of deutosternal denticles much as in female, sixth and seventh rows slightly widest; hypostomatic setae
<emphasis id="B97BC514FF8AFFC9FBDA792D210E1E73" box="[1056,1097,574,598]" italics="true" pageId="12" pageNumber="73">hp1</emphasis>
(28 32) similarly as long as
<emphasis id="B97BC514FF8AFFC9FA79792C20D91E73" box="[1411,1438,575,598]" italics="true" pageId="12" pageNumber="73">pc</emphasis>
(25 37),
<emphasis id="B97BC514FF8AFFC9FDE6797327021E5D" box="[540,581,608,632]" italics="true" pageId="12" pageNumber="73">hp3</emphasis>
usually longest (30 50),
<emphasis id="B97BC514FF8AFFC9FC9A797326CD1E5D" box="[864,906,608,632]" italics="true" pageId="12" pageNumber="73">hp2</emphasis>
(12 20). Palpi (length 145 187) similar in structure, form of setae as in female; palptrochanter with inner seta longer (30 44) than outer seta (17 32) but not markedly whip-like.
</paragraph>
<paragraph id="8BB01906FF8AFFC9FE1179F8208A1858" blockId="12.[491,1523,746,1597]" pageId="12" pageNumber="73">
Legs — Adult female. Legs I (707 755) about as long as legs IV (715 750), and about as long as dorsal shield; other leg lengths (excluding pretarsi): II (565 600), III (587 615). Leg I pretarsus with biarticulate shaft (38 42 to base of claws), tarsus (187 205) about 1.6 longer than tibia; other length ratios, femur (135 147): genu (112 118): tibia (117 125) about 1.15: 0.95: 1.0. Legs II-IV with tarsus/tibia length ratios about 2.0-2.3. Chaetotactic formulae of leg segments as given for species-group. Tarsi II-IV with ventroapical process (12 17) bluntly pointed, smooth apically (
<figureCitation id="13340583FF8AFFC9FCAD78AA269D1FF4" box="[855,986,953,977]" captionStart="Figure 13" captionStartId="31.[119,186,1454,1475]" captionTargetBox="[311,1289,241,1396]" captionTargetPageId="31" captionText="Figure 13 Tarsus II, right leg. A, B, H.Lasioseius cassidini n. sp.: A Adult female, dorsal view; B Adult male, dorsal view; H — Detail of distal region, lateroventral view. C, D, G.Lasioseius serripes n. sp.: C Adult female, dorsal view; D Adult male, anterolateral view; G. Detail of distal region, ventral view. E, F.Lasioseius fuscina n. sp.: E Adult male, dorsal view; F Adult female, dorsal view. I —Lasioseius duobtusisetis n. sp., adult female, dorsal view. (Solid black color indicates dorsal setae on segment)." figureDoi="http://doi.org/10.5281/zenodo.4487565" httpUri="https://zenodo.org/record/4487565/files/figure.png" pageId="12" pageNumber="73">Figs 13A, H</figureCitation>
), and with setae (
<emphasis id="B97BC514FF8AFFC9FB6D78AA21FD1FF4" box="[1175,1210,953,977]" italics="true" pageId="12" pageNumber="73">d-1</emphasis>
) (23 30) slightly shorter than pretarsus to base of claws (28 37). Tarsi II-IV with paradactyli asymmetrical, anterior element increasingly extending beyond tip of claw from II to IV, its length 30 35, twice as long as claw (15) on leg IV (
<figureCitation id="13340583FF8AFFC9FCE37F322639181C" box="[793,894,1057,1081]" captionStart="Figure 14" captionStartId="32.[119,186,1019,1040]" captionTargetBox="[280,1301,236,936]" captionTargetId="graphics-3@32.[280,1301,237,937]" captionTargetPageId="32" captionText="Figure 14 Tarsus IV, adult females, right leg, ventral view: A Lasioseius cassidini n. sp.; B Lasioseius serripes n. sp.; C Lasioseius duobtusisetis n. sp.; D Lasioseius fuscina n. sp. (solid black circles indicate dorsal setae insertions)." figureDoi="http://doi.org/10.5281/zenodo.4487567" httpUri="https://zenodo.org/record/4487567/files/figure.png" pageId="12" pageNumber="73">Fig. 14A</figureCitation>
). Tarsi II and III with setae (
<emphasis id="B97BC514FF8AFFC9FB527F332182181D" box="[1192,1221,1056,1080]" italics="true" pageId="12" pageNumber="73">l-1</emphasis>
) and (
<emphasis id="B97BC514FF8AFFC9FAF07F33206B181D" box="[1290,1324,1056,1080]" italics="true" pageId="12" pageNumber="73">v-1</emphasis>
) symmetrical in form,
<emphasis id="B97BC514FF8AFFC9FDD77F50271E187E" box="[557,601,1091,1115]" italics="true" pageId="12" pageNumber="73">pl-1</emphasis>
(32 35) and
<emphasis id="B97BC514FF8AFFC9FD0F7F502662187E" box="[757,805,1091,1115]" italics="true" pageId="12" pageNumber="73">av-1</emphasis>
(36 47) slightly longer than
<emphasis id="B97BC514FF8AFFC9FB917F5021D1187E" box="[1131,1174,1091,1115]" italics="true" pageId="12" pageNumber="73">al-1</emphasis>
(27 30) and
<emphasis id="B97BC514FF8AFFC9FACE7F502022187E" box="[1332,1381,1091,1115]" italics="true" pageId="12" pageNumber="73">pv-1</emphasis>
(26 37), respectively. Tarsus IV with distal seta
<emphasis id="B97BC514FF8AFFC9FC6F7F7626831858" box="[917,964,1125,1149]" italics="true" pageId="12" pageNumber="73">av-1</emphasis>
(50 60) blunt, clearly larger, thicker than
<emphasis id="B97BC514FF8AFFC9FA667F76208A1858" box="[1436,1485,1125,1149]" italics="true" pageId="12" pageNumber="73">pv-1</emphasis>
</paragraph>
<paragraph id="8BB01906FF8AFFC9FE117F9B210D18C0" blockId="12.[491,1523,746,1597]" pageId="12" pageNumber="73">
(37 45); adjacent setae (
<emphasis id="B97BC514FF8AFFC9FCFE7F9B26661885" box="[772,801,1160,1184]" italics="true" pageId="12" pageNumber="73">l-1</emphasis>
) symmetrically short (20 27), slender (
<figureCitation id="13340583FF8AFFC9FB357F9B20701885" box="[1231,1335,1160,1184]" captionStart="Figure 14" captionStartId="32.[119,186,1019,1040]" captionTargetBox="[280,1301,236,936]" captionTargetId="graphics-3@32.[280,1301,237,937]" captionTargetPageId="32" captionText="Figure 14 Tarsus IV, adult females, right leg, ventral view: A Lasioseius cassidini n. sp.; B Lasioseius serripes n. sp.; C Lasioseius duobtusisetis n. sp.; D Lasioseius fuscina n. sp. (solid black circles indicate dorsal setae insertions)." figureDoi="http://doi.org/10.5281/zenodo.4487567" httpUri="https://zenodo.org/record/4487567/files/figure.png" pageId="12" pageNumber="73">Fig. 14A</figureCitation>
). Tarsi II-IV with dorsal seta
<emphasis id="B97BC514FF8AFFC9FD677FB927FA18E7" box="[669,701,1194,1218]" italics="true" pageId="12" pageNumber="73">md</emphasis>
nearly reaching base of claws, and nearly as long as smooth attenuated setae (
<emphasis id="B97BC514FF8AFFC9FDCA7FDE271B18C0" box="[560,604,1229,1253]" italics="true" pageId="12" pageNumber="73">d-2)</emphasis>
(67 80 on II, 85 100 on III, 95 110 on IV).
</paragraph>
<paragraph id="8BB01906FF8AFFC9FDE97FFC27D11A18" blockId="12.[491,1523,746,1597]" pageId="12" pageNumber="73">
Adult male. Legs I clearly longer than dorsal shield, not thicker than those of female; relative leg lengths, excluding pretarsi, similar to those of female, I 550 660, II 445 524, III 455 552, IV 575 675. Legs with segment length ratios as in female. Tarsus I with long pretarsus (25 37 to base of claws), claws 10 12 long. Tarsi II-IV with dorsoapical setal processes (
<emphasis id="B97BC514FF8AFFC9FD987E6A27C219B4" box="[610,645,1401,1425]" italics="true" pageId="12" pageNumber="73">d-1</emphasis>
), ventroapical process and pretarsal structures formed as in female, pretarsus lengths 27 35 excluding claws (10) (
<figureCitation id="13340583FF8AFFC9FC787E8F26A31991" box="[898,996,1436,1460]" captionStart="Figure 13" captionStartId="31.[119,186,1454,1475]" captionTargetBox="[311,1289,241,1396]" captionTargetPageId="31" captionText="Figure 13 Tarsus II, right leg. A, B, H.Lasioseius cassidini n. sp.: A Adult female, dorsal view; B Adult male, dorsal view; H — Detail of distal region, lateroventral view. C, D, G.Lasioseius serripes n. sp.: C Adult female, dorsal view; D Adult male, anterolateral view; G. Detail of distal region, ventral view. E, F.Lasioseius fuscina n. sp.: E Adult male, dorsal view; F Adult female, dorsal view. I —Lasioseius duobtusisetis n. sp., adult female, dorsal view. (Solid black color indicates dorsal setae on segment)." figureDoi="http://doi.org/10.5281/zenodo.4487565" httpUri="https://zenodo.org/record/4487565/files/figure.png" pageId="12" pageNumber="73">Fig. 13B</figureCitation>
). Legs III-IV with tibial setae (
<emphasis id="B97BC514FF8AFFC9FAC47E8E200D1991" box="[1342,1354,1437,1460]" italics="true" pageId="12" pageNumber="73">v</emphasis>
) and leg IV genual seta
<emphasis id="B97BC514FF8AFFC9FD907EAC27C419F3" box="[618,643,1471,1494]" italics="true" pageId="12" pageNumber="73">av</emphasis>
dimorphically thicker, spinelike, less attenuate; length and form of other setae much as in female; seta
<emphasis id="B97BC514FF8AFFC9FD0A7EF3265B19DD" box="[752,796,1504,1528]" italics="true" pageId="12" pageNumber="73">pl-3</emphasis>
of basitarsus II smooth, attenuate (36 38) (
<figureCitation id="13340583FF8AFFC9FB037EF2201C19DC" box="[1273,1371,1505,1529]" captionStart="Figure 13" captionStartId="31.[119,186,1454,1475]" captionTargetBox="[311,1289,241,1396]" captionTargetPageId="31" captionText="Figure 13 Tarsus II, right leg. A, B, H.Lasioseius cassidini n. sp.: A Adult female, dorsal view; B Adult male, dorsal view; H — Detail of distal region, lateroventral view. C, D, G.Lasioseius serripes n. sp.: C Adult female, dorsal view; D Adult male, anterolateral view; G. Detail of distal region, ventral view. E, F.Lasioseius fuscina n. sp.: E Adult male, dorsal view; F Adult female, dorsal view. I —Lasioseius duobtusisetis n. sp., adult female, dorsal view. (Solid black color indicates dorsal setae on segment)." figureDoi="http://doi.org/10.5281/zenodo.4487565" httpUri="https://zenodo.org/record/4487565/files/figure.png" pageId="12" pageNumber="73">Fig. 13B</figureCitation>
), but
<emphasis id="B97BC514FF8AFFC9FA587EF3208919DD" box="[1442,1486,1504,1528]" italics="true" pageId="12" pageNumber="73">pl-3</emphasis>
of basitarsi III, IV (21 22) similar to
<emphasis id="B97BC514FF8AFFC9FC6B7D1026FA1A3E" box="[913,957,1539,1563]" italics="true" pageId="12" pageNumber="73">
pl-
<quantity id="4CF7B4E3FF8AFFC9FC4A7D1026FA1A3E" box="[944,957,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="73" unit="in" value="2.0">2</quantity>
</emphasis>
in thicker, slightly shorter form than other three basitarsal setae.
</paragraph>
<paragraph id="8BB01906FF8AFFC8FE117D7820601D8F" blockId="12.[491,1511,1642,2045]" lastBlockId="13.[491,1498,229,426]" lastPageId="13" lastPageNumber="74" pageId="12" pageNumber="73">
Type material — Except the final one, noted below, all specimens collected at the La Selva Biological Station,
<collectingRegion id="49CBD7E4FF8AFFC9FD467D9E26381A80" box="[700,895,1677,1701]" country="Costa Rica" name="Heredia" pageId="12" pageNumber="73">Heredia Province</collectingRegion>
,
<collectingCountry id="F3185996FF8AFFC9FC717D9E21461A80" box="[907,1025,1677,1701]" name="Costa Rica" pageId="12" pageNumber="73">Costa Rica</collectingCountry>
(
<geoCoordinate id="EE3B7FC1FF8AFFC9FBEA7D9E212B1A80" box="[1040,1132,1677,1701]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="12" pageNumber="73" precision="925" value="10.433333">10°26´N</geoCoordinate>
,
<geoCoordinate id="EE3B7FC1FF8AFFC9FB837D9E21911A80" box="[1145,1238,1677,1701]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="12" pageNumber="73" precision="925" value="-84.01667">84°01´W</geoCoordinate>
, elevation
<quantity id="4CF7B4E3FF8AFFC9FAB17D9E20FF1A80" box="[1355,1464,1677,1701]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="12" pageNumber="73" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50-150 m</quantity>
):
<typeStatus id="54B4A7A4FF8AFFC9FE117DBC273B1AE2" box="[491,636,1711,1735]" pageId="12" pageNumber="73" type="holotype">HOLOTYPE</typeStatus>
: adult female,
<date id="FFB13FC6FF8AFFC9FCE77DBC26C71AE2" box="[797,896,1711,1735]" pageId="12" pageNumber="73" value="1997-07-08">8 July 1997</date>
, ex adult
<taxonomicName id="4C0F6285FF8AFFC9FBFB7DBC21B31AE2" authority="Baly" authorityName="Baly" box="[1025,1268,1711,1735]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B97BC514FF8AFFC9FBFB7DBC21871AE2" box="[1025,1216,1711,1735]" italics="true" pageId="12" pageNumber="73">Cephaloleia bella</emphasis>
Baly
</taxonomicName>
on
<taxonomicName id="4C0F6285FF8AFFC9FADB7DBC27721ACF" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="12" pageNumber="73" phylum="Tracheophyta" rank="species" species="pogonantha">
<emphasis id="B97BC514FF8AFFC9FADB7DBC27721ACF" italics="true" pageId="12" pageNumber="73">Heliconia pogonantha</emphasis>
</taxonomicName>
leaves, coll. M. Paniagua.
<typeStatus id="54B4A7A4FF8AFFC9FCA07DC126B61ACF" box="[858,1009,1746,1770]" pageId="12" pageNumber="73" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D09D28FFF8AFFC9FC017DC1210E1ACF" box="[1019,1097,1746,1770]" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
female,
<specimenCount id="9D09D28FFF8AFFC9FB5C7DC121B31ACF" box="[1190,1268,1746,1770]" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
male, with same data as
<typeStatus id="54B4A7A4FF8AFFC9FDF37DE7272C1B29" box="[521,619,1780,1804]" pageId="12" pageNumber="73" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D09D28FFF8AFFC9FD8C7DE727871B29" box="[630,704,1780,1804]" pageId="12" pageNumber="73" type="adult">2 adult</specimenCount>
females,
<specimenCount id="9D09D28FFF8AFFC9FCD27DE726311B29" box="[808,886,1780,1804]" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
male,
<date id="FFB13FC6FF8AFFC9FC457DE7217F1B29" box="[959,1080,1780,1804]" pageId="12" pageNumber="73" value="1997-06-12">12 June 1997</date>
, ex coexistent adults of
<taxonomicName id="4C0F6285FF8AFFC9FAB77DE727161B0A" authority="Baly" authorityName="Baly" authorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="belti">
<emphasis id="B97BC514FF8AFFC9FAB77DE7275D1B0A" italics="true" pageId="12" pageNumber="73">Cephaloleia belti</emphasis>
Baly
</taxonomicName>
and
<taxonomicName id="4C0F6285FF8AFFC9FD717C04262A1B0A" authority="Donckier" authorityName="Donckier" box="[651,877,1815,1839]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="ornatrix">
<emphasis id="B97BC514FF8AFFC9FD717C0426451B0A" box="[651,770,1815,1839]" italics="true" pageId="12" pageNumber="73">C. ornatrix</emphasis>
Donckier
</taxonomicName>
from
<taxonomicName id="4C0F6285FF8AFFC9FC4B7C04215D1B0A" box="[945,1050,1815,1839]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="12" pageNumber="73" phylum="Tracheophyta" rank="genus">
<emphasis id="B97BC514FF8AFFC9FC4B7C04215D1B0A" box="[945,1050,1815,1839]" italics="true" pageId="12" pageNumber="73">Heliconia</emphasis>
</taxonomicName>
leaves, coll. ALAS;
<specimenCount id="9D09D28FFF8AFFC9FAF97C0420091B0A" box="[1283,1358,1815,1839]" pageId="12" pageNumber="73" type="adult">3 adult</specimenCount>
females,
<specimenCount id="9D09D28FFF8AFFC9FA4D7C0427621B74" pageId="12" pageNumber="73" type="adult">2 adult</specimenCount>
males,
<date id="FFB13FC6FF8AFFC9FD837C2A26421B74" box="[633,773,1849,1873]" pageId="12" pageNumber="73" value="1994-04-15">15 April 1994</date>
, ex adult
<taxonomicName id="4C0F6285FF8AFFC9FC8A7C2A215C1B74" authorityName="Latreille" authorityYear="1802" box="[880,1051,1849,1873]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
, no host plant data, M. Paniagua coll.;
<specimenCount id="9D09D28FFF8AFFC9FA307C2A27621B51" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
female,
<specimenCount id="9D09D28FFF8AFFC9FD7B7C4F27891B51" box="[641,718,1884,1908]" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
male,
<date id="FFB13FC6FF8AFFC9FCEF7C4F26371B51" box="[789,880,1884,1908]" pageId="12" pageNumber="73" value="1997-06">June 1997</date>
, ex adult
<taxonomicName id="4C0F6285FF8AFFC9FC1A7C4F21941B51" authority="Baly" authorityName="Baly" box="[992,1235,1884,1908]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B97BC514FF8AFFC9FC1A7C4F21D81B51" box="[992,1183,1884,1908]" italics="true" pageId="12" pageNumber="73">Cephaloleia bella</emphasis>
Baly
</taxonomicName>
, no host plant data, coll. D. Johnson;
<specimenCount id="9D09D28FFF8AFFC9FD827C6D27811BB3" box="[632,710,1918,1942]" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
female (gravid with larva),
<specimenCount id="9D09D28FFF8AFFC9FC047C6D210B1BB3" box="[1022,1100,1918,1942]" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
male,
<date id="FFB13FC6FF8AFFC9FB6E7C6C21A81BB3" box="[1172,1263,1918,1942]" pageId="12" pageNumber="73" value="1997-06">June 1997</date>
, ex adult
<taxonomicName id="4C0F6285FF8AFFC9FAAA7C6D27161B9D" authority="Baly" authorityName="Baly" authorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="belti">
<emphasis id="B97BC514FF8AFFC9FAAA7C6D275D1B9D" italics="true" pageId="12" pageNumber="73">Cephaloleia belti</emphasis>
Baly
</taxonomicName>
, no host plant data, coll. D. Johnson;
<specimenCount id="9D09D28FFF8AFFC9FC0D7CB321061B9D" box="[1015,1089,1952,1976]" pageId="12" pageNumber="73" type="adult">2 adult</specimenCount>
males,
<date id="FFB13FC6FF8AFFC9FB697CB320651B9D" box="[1171,1314,1952,1976]" pageId="12" pageNumber="73" value="1994-02">February 1994</date>
, associated with adult
<taxonomicName id="4C0F6285FF8AFFC9FDDD7CD027DA1BFE" baseAuthorityName="Staines" baseAuthorityYear="2009" box="[551,669,1987,2011]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8AFFC9FDDD7CD027DA1BFE" box="[551,669,1987,2011]" italics="true" pageId="12" pageNumber="73">Chelobasis</emphasis>
</taxonomicName>
sp. on leaves of
<taxonomicName id="4C0F6285FF8AFFC9FCAA7CD021791BFE" box="[848,1086,1987,2011]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="12" pageNumber="73" phylum="Tracheophyta" rank="species" species="pogonantha">
<emphasis id="B97BC514FF8AFFC9FCAA7CD021791BFE" box="[848,1086,1987,2011]" italics="true" pageId="12" pageNumber="73">Heliconia pogonantha</emphasis>
</taxonomicName>
, coll. E.E. Lindquist &amp; ALAS;
<specimenCount id="9D09D28FFF8AFFC9FA6E7CD020A51BFE" box="[1428,1506,1987,2011]" pageId="12" pageNumber="73" type="adult">1 adult</specimenCount>
female,
<specimenCount id="9D09D28FFF8AFFC9FDB97CF627C91BD8" box="[579,654,2021,2045]" pageId="12" pageNumber="73" type="adult">2 adult</specimenCount>
males,
<date id="FFB13FC6FF8AFFC9FD197CF626D51BD8" box="[739,914,2021,2045]" pageId="12" pageNumber="73" value="1994-02-18">18 February 1994</date>
, ex adult
<taxonomicName id="4C0F6285FF8AFFC9FC057CF621ED1BD8" authorityName="Latreille" authorityYear="1802" box="[1023,1194,2021,2045]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
, no host plant data, coll. ALAS;
<specimenCount id="9D09D28FFF8BFFC8FDC57BF627CE1CD8" box="[575,649,229,253]" pageId="13" pageNumber="74" type="adult">3 adult</specimenCount>
females,
<date id="FFB13FC6FF8BFFC8FD0A7BF6261C1CD8" box="[752,859,229,253]" pageId="13" pageNumber="74" value="1997-06-02">2 June 1997</date>
, with adult hispine beetles on
<taxonomicName id="4C0F6285FF8BFFC8FB4A7BF620011CD8" box="[1200,1350,229,253]" class="Liliopsida" family="Heliconiaceae" kingdom="Plantae" order="Zingiberales" pageId="13" pageNumber="74" phylum="Tracheophyta" rank="family">Heliconiaceae</taxonomicName>
leaves, coll. E.E. Lindquist;
<specimenCount id="9D09D28FFF8BFFC8FD6D7A1B27A71D05" box="[663,736,264,288]" pageId="13" pageNumber="74" type="adult">2 adult</specimenCount>
females,
<date id="FFB13FC6FF8BFFC8FCBF7A1A26E71D05" box="[837,928,264,288]" pageId="13" pageNumber="74" value="1997-06">June 1997</date>
, ex adult
<taxonomicName id="4C0F6285FF8BFFC8FBEE7A1B21981D05" authorityName="Baly" authorityYear="1858" box="[1044,1247,264,288]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B97BC514FF8BFFC8FBEE7A1B21981D05" box="[1044,1247,264,288]" italics="true" pageId="13" pageNumber="74">Cephaloleia vicina</emphasis>
</taxonomicName>
on
<taxonomicName id="4C0F6285FF8BFFC8FAFE7A1B274A1D66" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="13" pageNumber="74" phylum="Tracheophyta" rank="species" species="pogonantha">
<emphasis id="B97BC514FF8BFFC8FAFE7A1B274A1D66" italics="true" pageId="13" pageNumber="74">Heliconia pogonantha</emphasis>
</taxonomicName>
, coll. not indicated;
<specimenCount id="9D09D28FFF8BFFC8FD0D7A3826021D66" box="[759,837,299,323]" pageId="13" pageNumber="74" type="adult">1 adult</specimenCount>
male,
<date id="FFB13FC6FF8BFFC8FC777A38215B1D66" box="[909,1052,299,323]" pageId="13" pageNumber="74" value="1994-02">February 1994</date>
, associated with adult
<taxonomicName id="4C0F6285FF8BFFC8FAF67A3820901D67" authorityName="Baly" authorityYear="1858" box="[1292,1495,299,323]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B97BC514FF8BFFC8FAF67A3820901D67" box="[1292,1495,299,323]" italics="true" pageId="13" pageNumber="74">Cephaloleia vicina</emphasis>
</taxonomicName>
on leaf of
<taxonomicName id="4C0F6285FF8BFFC8FDA17A5E260D1D40" box="[603,842,333,357]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="13" pageNumber="74" phylum="Tracheophyta" rank="species" species="pogonantha">
<emphasis id="B97BC514FF8BFFC8FDA17A5E260D1D40" box="[603,842,333,357]" italics="true" pageId="13" pageNumber="74">Heliconia pogonantha</emphasis>
</taxonomicName>
, coll. E.E. Lindquist &amp; ALAS.
<collectingCountry id="F3185996FF8BFFC8FB517A5E20651D40" box="[1195,1314,333,357]" name="Costa Rica" pageId="13" pageNumber="74">Costa Rica</collectingCountry>
, Osa Peninsula, Corcovado National Park (
<geoCoordinate id="EE3B7FC1FF8BFFC8FCEA7A63262C1DAD" box="[784,875,368,392]" degrees="8" direction="north" minutes="27.39" orientation="latitude" pageId="13" pageNumber="74" precision="9" value="8.4565">8°27.39´N</geoCoordinate>
,
<geoCoordinate id="EE3B7FC1FF8BFFC8FC8C7A6326A81DAD" box="[886,1007,368,392]" degrees="83" direction="west" minutes="25.45" orientation="longitude" pageId="13" pageNumber="74" precision="9" value="-83.424164">83°25.45´W</geoCoordinate>
,elevation
<quantity id="4CF7B4E3FF8BFFC8FBA07A6321811DAD" box="[1114,1222,368,392]" metricMagnitude="2" metricUnit="m" metricValue="2.55" metricValueMax="5.0" metricValueMin="0.1" pageId="13" pageNumber="74" unit="m" value="255.0" valueMax="500.0" valueMin="10.0">10-500 m</quantity>
):
<specimenCount id="9D09D28FFF8BFFC8FB277A63206B1DAD" box="[1245,1324,368,392]" pageId="13" pageNumber="74" type="adult">1 adult</specimenCount>
male, May 1997, ex adult
<taxonomicName id="4C0F6285FF8BFFC8FD737A8126CB1D8F" authority="Gray" authorityName="Gray" authorityYear="1832" box="[649,908,402,426]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B97BC514FF8BFFC8FD737A8126141D8F" box="[649,851,402,426]" italics="true" pageId="13" pageNumber="74">Chelobasis bicolor</emphasis>
Gray
</taxonomicName>
, no host plant data, coll. D. Johnson.
</paragraph>
<paragraph id="8BB01906FF8BFFC8FE117ACB207E1EB9" blockId="13.[491,1506,471,668]" pageId="13" pageNumber="74">
Host identifications — Some of the identities of the associated beetles given above are in accord with current molecular studies,
<emphasis id="B97BC514FF8BFFC8FC727AE826EF1E37" box="[904,936,507,530]" italics="true" pageId="13" pageNumber="74">i.e.</emphasis>
, those as noted below for
<taxonomicName id="4C0F6285FF8BFFC8FB3A7AE820CA1E37" authorityName="Moraza &amp; Lindquist" authorityYear="2018" box="[1216,1421,507,530]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="serripes">
<emphasis id="B97BC514FF8BFFC8FB3A7AE820CA1E37" box="[1216,1421,507,530]" italics="true" pageId="13" pageNumber="74">Lasioseius serripes</emphasis>
</taxonomicName>
. However, the following identities are discordant.
<taxonomicName id="4C0F6285FF8BFFC8FC2B790F21DA1E11" authorityName="Baly" authorityYear="1858" box="[977,1181,540,564]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B97BC514FF8BFFC8FC2B790F21DA1E11" box="[977,1181,540,564]" italics="true" pageId="13" pageNumber="74">Cephaloleia vicina</emphasis>
</taxonomicName>
does not occur at La Selva, and specimens identified as such from there are, in all cases checked by DNA barcodes,
<taxonomicName id="4C0F6285FF8BFFC8FA56792C27671E5C" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B97BC514FF8BFFC8FA56792C27671E5C" italics="true" pageId="13" pageNumber="74">C. bella</emphasis>
</taxonomicName>
. Further,
<taxonomicName id="4C0F6285FF8BFFC8FD77797127A61E5C" box="[653,737,609,633]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B97BC514FF8BFFC8FD77797127A61E5C" box="[653,737,609,633]" italics="true" pageId="13" pageNumber="74">C. bella</emphasis>
</taxonomicName>
normally does not occur on
<taxonomicName id="4C0F6285FF8BFFC8FBEE797220441E5C" box="[1044,1283,609,633]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="13" pageNumber="74" phylum="Tracheophyta" rank="species" species="pogonantha">
<emphasis id="B97BC514FF8BFFC8FBEE797220441E5C" box="[1044,1283,609,633]" italics="true" pageId="13" pageNumber="74">Heliconia pogonantha</emphasis>
</taxonomicName>
; thousands of field diet records indicate it feeds only on
<taxonomicName id="4C0F6285FF8BFFC8FC8F799620721EB9" authority="(Garcia-Robledo et al., 2017)" baseAuthorityName="Garcia-Robledo" baseAuthorityYear="2017" box="[885,1333,644,668]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="13" pageNumber="74" phylum="Tracheophyta" rank="species" species="imbricata">
<emphasis id="B97BC514FF8BFFC8FC8F799626BB1EB9" box="[885,1020,644,668]" italics="true" pageId="13" pageNumber="74">H. imbricata</emphasis>
(
<bibRefCitation id="EF9E64F7FF8BFFC8FBF0799720681EB9" author="Garcia-Robledo C. &amp; Horvitz C. C. &amp; Kress J. W. &amp; Carvajal-Acosta N. &amp; Erwin T. L. &amp; Staines C. L." box="[1034,1327,644,668]" pageId="13" pageNumber="74" pagination="803 - 810" refId="ref24363" refString="Garcia-Robledo C., Horvitz C. C., Kress J. W., Carvajal-Acosta N., Erwin T. L., Staines C. L. (2017) - Experimental assemblage of novel plant-herbivore interactions: ecological host shifts after 40 million years of isolation - Biotropica, 49 (6): 803 - 810. doi: 10.1111 / btp. 12464" type="journal article" year="2017">
Garcia-Robledo
<emphasis id="B97BC514FF8BFFC8FB46799621B51EB9" box="[1212,1266,644,668]" italics="true" pageId="13" pageNumber="74">et al.</emphasis>
, 2017
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BB01906FF8BFFC8FE1179DA27371FAB" blockId="13.[491,1520,713,910]" pageId="13" pageNumber="74">
Etymology — The specific name is based on the chrysomelid subfamilial name
<taxonomicName id="4C0F6285FF8BFFC8FA8979DA275D1F21" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="subFamily" subFamily="Cassidinae">Cassidinae</taxonomicName>
, which includes the tribes
<taxonomicName id="4C0F6285FF8BFFC8FCA179FF26F21F21" authorityName="Chapuis" authorityYear="1875" box="[859,949,748,772]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="tribe" tribe="Arescini">Arescini</taxonomicName>
and
<taxonomicName id="4C0F6285FF8BFFC8FC0E79FF21241F21" authorityName="Hope" authorityYear="1840" box="[1012,1123,748,772]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="tribe" tribe="Imatidiini">Imatidiini</taxonomicName>
(or Cephaloleini) to which beetle associates of the genera
<taxonomicName id="4C0F6285FF8BFFC8FD16781D26251F03" baseAuthorityName="Staines" baseAuthorityYear="2009" box="[748,866,782,806]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8BFFC8FD16781D26251F03" box="[748,866,782,806]" italics="true" pageId="13" pageNumber="74">Chelobasis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F6285FF8BFFC8FC6F781D215F1F03" baseAuthorityName="Staines &amp; Garcia-Robledo" baseAuthorityYear="2014" box="[917,1048,782,806]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8BFFC8FC6F781D215F1F03" box="[917,1048,782,806]" italics="true" pageId="13" pageNumber="74">Cephaloleia</emphasis>
</taxonomicName>
belong, respectively. We have chosen this name for the species with the widest array of host associates and as the species-group name, anticipating that rolled-leaf beetles of other genera in these tribes will be found to live with these mites.
</paragraph>
<paragraph id="8BB01906FF8BFFC8FE1178A82740185A" blockId="13.[491,1525,955,1635]" pageId="13" pageNumber="74">
Remarks — The male from Corcovado measured ca 12-19% smaller in dimensions than seven males measured from La Selva; however, in other descriptive aspects it fully resembles those from La Selva. Two females are asymmetrical in having humeral seta
<emphasis id="B97BC514FF8BFFC8FAED7F132068183D" box="[1303,1327,1024,1048]" italics="true" pageId="13" pageNumber="74">r3</emphasis>
inserted off the dorsal shield on one side, yet the aligned
<emphasis id="B97BC514FF8BFFC8FC517F312684181F" box="[939,963,1058,1082]" italics="true" pageId="13" pageNumber="74">r2</emphasis>
and
<emphasis id="B97BC514FF8BFFC8FC037F312176181F" box="[1017,1073,1058,1082]" italics="true" pageId="13" pageNumber="74">r4-r6</emphasis>
are on the shield. Two females show asymmetry in size and arrangement of the vertical cluster of setae
<emphasis id="B97BC514FF8BFFC8FB5D7F5621FB1878" box="[1191,1212,1093,1117]" italics="true" pageId="13" pageNumber="74">j1</emphasis>
,
<emphasis id="B97BC514FF8BFFC8FB327F56219A1878" box="[1224,1245,1093,1117]" italics="true" pageId="13" pageNumber="74">j2</emphasis>
,
<emphasis id="B97BC514FF8BFFC8FB107F5620461878" box="[1258,1281,1093,1117]" italics="true" pageId="13" pageNumber="74">z1</emphasis>
; one male lacks one
<emphasis id="B97BC514FF8BFFC8FE117F742747185A" box="[491,512,1127,1151]" italics="true" pageId="13" pageNumber="74">j2</emphasis>
.
</paragraph>
<paragraph id="8BB01906FF8BFFC8FDE97F9926671A46" blockId="13.[491,1525,955,1635]" pageId="13" pageNumber="74">
Collection records of this mite species associated with beetles of three different species of
<taxonomicName id="4C0F6285FF8BFFC8FDF07FBF27CA18E1" baseAuthorityName="Staines &amp; Garcia-Robledo" baseAuthorityYear="2014" box="[522,653,1196,1220]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8BFFC8FDF07FBF27CA18E1" box="[522,653,1196,1220]" italics="true" pageId="13" pageNumber="74">Cephaloleia</emphasis>
</taxonomicName>
and one of
<taxonomicName id="4C0F6285FF8BFFC8FCF57FBF26C218E1" baseAuthorityName="Staines" baseAuthorityYear="2009" box="[783,901,1196,1220]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8BFFC8FCF57FBF26C218E1" box="[783,901,1196,1220]" italics="true" pageId="13" pageNumber="74">Chelobasis</emphasis>
</taxonomicName>
leave uncertainty as to host preferences, particularly as beetles of these species are often coexistent (
<bibRefCitation id="EF9E64F7FF8BFFC8FC087FDC21A518C2" author="Garcia-Robledo C. &amp; Horvitz C. C. &amp; Staines C. L." box="[1010,1250,1231,1255]" pageId="13" pageNumber="74" pagination="50 - 68" refId="ref24433" refString="Garcia-Robledo C., Horvitz C. C., Staines C. L. 2010 - Larval morphology and development, host plants, adult longevity, sexual dimorphism and notes on natural history in Cephaloleia rolled-leaf beetles (Coleoptera: Chrysomelidae: Hispinae) - Zootaxa, 2610: 50 - 68." type="journal article" year="2010">Garcia-Robledo 2010</bibRefCitation>
,
<bibRefCitation id="EF9E64F7FF8BFFC8FB177FDC203318C2" author="Staines C. L." box="[1261,1396,1231,1255]" pageId="13" pageNumber="74" refId="ref26178" refString="Staines C. L. 2011 - Hispines (Chrysomelidae, Cassidinae) of La Selva Biological Station, Costa Rica" type="book" year="2011">Staines 2011</bibRefCitation>
,
<bibRefCitation id="EF9E64F7FF8BFFC8FA857FDC209F18C2" author="Schmitt M. &amp; Frank M." box="[1407,1496,1231,1255]" pageId="13" pageNumber="74" pagination="55 - 69" refId="ref26078" refString="Schmitt M., Frank M. 2013 - Notes on the ecology of rolled-leaf hispines (Chrysomelidae, Cassidinae) at La Gamba (Costa Rica) - ZooKeys, 332: 55 - 69. doi: 10.3897 / zookeys. 332.5215" type="journal article" year="2013">Schmitt</bibRefCitation>
&amp; Frank 2013,
<bibRefCitation id="EF9E64F7FF8BFFC8FD767FE126EB192F" author="Moraza M. L. &amp; Lindquist E. E." box="[652,940,1266,1290]" pageId="13" pageNumber="74" pagination="301 - 351" refId="ref25953" refString="Moraza M. L., Lindquist E. E. 2015 - Systematics and biology of mites associated with neotropical hispine beetles in unfurled leaves of Heliconia, with descriptions of two new genera of the family Melicharidae (Acari: Mesostigmata: Gamasina: Ascoides) - Zootaxa, 3931 (3): 301 - 351. doi: 10.11646 / zootaxa. 3931.3.1" type="journal article" year="2015">Moraza &amp; Lindquist 2015</bibRefCitation>
). Adults of
<taxonomicName id="4C0F6285FF8BFFC8FBC77FE02053192F" authorityName="Moraza &amp; Lindquist" authorityYear="2018" box="[1085,1300,1266,1290]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="cassidini">
<emphasis id="B97BC514FF8BFFC8FBC77FE02053192F" box="[1085,1300,1266,1290]" italics="true" pageId="13" pageNumber="74">Lasioseius cassidini</emphasis>
</taxonomicName>
are clearly the largest of the described species of this group. However, the data available are inadequate to test whether a mite/beetle size correlation exists, such as was hypothesized for larger mites being associated with larger beetles, for species of
<taxonomicName id="4C0F6285FF8BFFC8FBEF7E4A21D41954" box="[1045,1171,1369,1393]" class="Arachnida" family="Ascidae" genus="Hispiniphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8BFFC8FBEF7E4A21D41954" box="[1045,1171,1369,1393]" italics="true" pageId="13" pageNumber="74">Hispiniphis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0F6285FF8BFFC8FB3C7E4A20001954" box="[1222,1351,1369,1393]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8BFFC8FB3C7E4A20001954" box="[1222,1351,1369,1393]" italics="true" pageId="13" pageNumber="74">Makarovaia</emphasis>
</taxonomicName>
, which are more intimate subelytral associates of these same beetles (
<bibRefCitation id="EF9E64F7FF8BFFC8FB997E6F20C719B1" author="Moraza M. L. &amp; Lindquist E. E." box="[1123,1408,1404,1428]" pageId="13" pageNumber="74" pagination="301 - 351" refId="ref25953" refString="Moraza M. L., Lindquist E. E. 2015 - Systematics and biology of mites associated with neotropical hispine beetles in unfurled leaves of Heliconia, with descriptions of two new genera of the family Melicharidae (Acari: Mesostigmata: Gamasina: Ascoides) - Zootaxa, 3931 (3): 301 - 351. doi: 10.11646 / zootaxa. 3931.3.1" type="journal article" year="2015">Moraza &amp; Lindquist 2015</bibRefCitation>
, 2016). Although there are records of a few specimens of
<taxonomicName id="4C0F6285FF8BFFC8FBFF7E8C21381993" authorityName="Moraza &amp; Lindquist" authorityYear="2018" box="[1029,1151,1438,1462]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="cassidini">
<emphasis id="B97BC514FF8BFFC8FBFF7E8C21381993" box="[1029,1151,1438,1462]" italics="true" pageId="13" pageNumber="74">L. cassidini</emphasis>
</taxonomicName>
with relatively large beetles of
<taxonomicName id="4C0F6285FF8BFFC8FE117ED2272619FC" baseAuthorityName="Staines" baseAuthorityYear="2009" box="[491,609,1473,1497]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B97BC514FF8BFFC8FE117ED2272619FC" box="[491,609,1473,1497]" italics="true" pageId="13" pageNumber="74">Chelobasis</emphasis>
</taxonomicName>
, including one from the Osa Peninsula, they are based only on adult male mites, which may not present inherent preferences phoretically. Tentatively, subject to further studies, we conclude that, at La Selva,
<taxonomicName id="4C0F6285FF8BFFC8FCC67D1421551A3B" authorityName="Moraza &amp; Lindquist" authorityYear="2018" box="[828,1042,1542,1566]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="cassidini">
<emphasis id="B97BC514FF8BFFC8FCC67D1421551A3B" box="[828,1042,1542,1566]" italics="true" pageId="13" pageNumber="74">Lasioseius cassidini</emphasis>
</taxonomicName>
may be associated regularly with small beetles of the host specialist
<taxonomicName id="4C0F6285FF8BFFC8FCE57D3B26981A65" box="[799,991,1576,1600]" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="bella">
<emphasis id="B97BC514FF8BFFC8FCE57D3B26981A65" box="[799,991,1576,1600]" italics="true" pageId="13" pageNumber="74">Cephaloleia bella</emphasis>
</taxonomicName>
and to some extent with the host generalist
<taxonomicName id="4C0F6285FF8BFFC8FA4C7D3A275D1A46" authorityName="Baly" authorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="belti">
<emphasis id="B97BC514FF8BFFC8FA4C7D3A275D1A46" italics="true" pageId="13" pageNumber="74">C. belti</emphasis>
</taxonomicName>
on
<taxonomicName id="4C0F6285FF8BFFC8FDB97D58265E1A46" box="[579,793,1611,1635]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="13" pageNumber="74" phylum="Tracheophyta" rank="species" species="imbricata">
<emphasis id="B97BC514FF8BFFC8FDB97D58265E1A46" box="[579,793,1611,1635]" italics="true" pageId="13" pageNumber="74">Heliconia imbricata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>