188 lines
18 KiB
XML
188 lines
18 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4762915" ID-GBIF-Dataset="93917b8d-1ce7-425e-912a-5892214c81b4" ID-GBIF-Taxon="182073758" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762915" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1620936844672" checkinUser="felipe" docAuthor="Stark, Bill P. & Sheldon, Andrew L." docDate="2009" docId="03A621374906FF90FC18EB96FD2D34E0" docLanguage="en" docName="Illiesia.5.2.11-19.pdf" docOrigin="Illiesia 5 (2)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Phanoperla astrospinata Stark & Sheldon 2009, sp. n." docType="treatment" docVersion="5" lastPageId="7" lastPageNumber="17" masterDocId="FF9F594F4903FF97FFD4E926FFE6341B" masterDocTitle="Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species" masterLastPageNumber="19" masterPageNumber="11" pageId="5" pageNumber="16" updateTime="1632022693049" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Records Of Neoperlini (Plecoptera: Perlidae) From Brunei Darussalam And Sarawak, With Descriptions Of New Phanoperla Banks And Neoperla Needham Species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Stark, Bill P.</mods:namePart>
|
||
<mods:affiliation>and Andrew L. Sheldon & Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
|
||
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Sheldon, Andrew L.</mods:namePart>
|
||
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu & Division of Biological Sciences, University of Montana, Missoula, Montana, U. S. A. 59812 E-mail: andylsheldon @ comcast. net & and Andrew L. Sheldon & Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
|
||
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Illiesia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2009</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2009-02-05</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>11</mods:start>
|
||
<mods:end>19</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4759022</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">93917b8d-1ce7-425e-912a-5892214c81b4</mods:identifier>
|
||
<mods:identifier type="ISSN">1854-0392</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">4759022</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762915" ID-GBIF-Taxon="182073758" ID-Zenodo-Dep="4762915" LSID="urn:lsid:plazi:treatment:03A621374906FF90FC18EB96FD2D34E0" httpUri="http://treatment.plazi.org/id/03A621374906FF90FC18EB96FD2D34E0" lastPageId="7" lastPageNumber="17" pageId="5" pageNumber="16">
|
||
<subSubSection box="[972,1306,688,712]" pageId="5" pageNumber="16" type="nomenclature">
|
||
<paragraph blockId="5.[972,1306,688,712]" box="[972,1306,688,712]" pageId="5" pageNumber="16">
|
||
<heading bold="true" box="[972,1306,688,712]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="16" reason="2">
|
||
<emphasis bold="true" box="[972,1306,688,712]" pageId="5" pageNumber="16">
|
||
<taxonomicName authority="Stark & Sheldon, 2009" authorityName="Stark & Sheldon" authorityYear="2009" box="[972,1244,688,712]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="astrospinata" status="sp. n.">
|
||
<emphasis bold="true" box="[972,1244,688,712]" italics="true" pageId="5" pageNumber="16">Phanoperla astrospinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1248,1306,688,712]" pageId="5" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[1076,1204,724,748]" pageId="5" pageNumber="16" type="description">
|
||
<paragraph blockId="5.[1076,1204,724,748]" box="[1076,1204,724,748]" pageId="5" pageNumber="16">
|
||
(
|
||
<figureCitation box="[1083,1196,724,748]" captionStart="Figs" captionStartId="6.[189,240,1434,1458]" captionTargetBox="[324,1344,217,1354]" captionTargetId="figure-254@6.[324,1307,217,1354]" captionTargetPageId="6" captionText="Figs. 11‐14. Phanoperla astrospinata (11‐12) and P. belalong (13‐14). 11. Male terminalia, 12. Aedeagus, lateral aspect, 13. Male terminalia, 14. Aedeagus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4759032" httpUri="https://zenodo.org/record/4759032/files/figure.png" pageId="5" pageNumber="16">Figs. 11‐12</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="16" type="materials_examined">
|
||
<paragraph blockId="5.[835,1445,791,1885]" pageId="5" pageNumber="16">
|
||
<emphasis bold="true" box="[835,1048,792,816]" pageId="5" pageNumber="16">Material examined.</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3111531332" collectingDate="2008-07-11" collectingDateMax="2008-07-16" collectingDateMin="2008-07-11" collectionCode="ALS, USNM" collectorName="A. L. Sheldon" country="Brunei" county="Studies Centre" latitude="4.54822" location="TemburongDistrict" longLatPrecision="1" longitude="115.15823" municipality="SungaiBelalong" pageId="5" pageNumber="16" specimenCode="B43-08" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<typeStatus box="[1050,1152,792,816]" pageId="5" pageNumber="16">Holotype</typeStatus>
|
||
<specimenCount box="[1155,1175,791,816]" pageId="5" pageNumber="16" type="male">♂</specimenCount>
|
||
from
|
||
<collectingCountry box="[1235,1440,792,816]" name="Brunei" pageId="5" pageNumber="16">Brunei Darussalam</collectingCountry>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:03A621374906FF90FC18EB96FD2D34E0:8ED0C6FA4906FF92FC97EA1DFBF03748" box="[835,1046,827,851]" country="Brunei" county="Studies Centre" latitude="4.54822" longLatPrecision="1" longitude="115.15823" municipality="SungaiBelalong" name="TemburongDistrict" pageId="5" pageNumber="16">TemburongDistrict</location>
|
||
,
|
||
<collectingMunicipality box="[1052,1223,827,851]" pageId="5" pageNumber="16">SungaiBelalong</collectingMunicipality>
|
||
, KualaBelalongField
|
||
<collectingCounty box="[835,994,865,889]" pageId="5" pageNumber="16">Studies Centre</collectingCounty>
|
||
,
|
||
<geoCoordinate box="[1002,1127,860,890]" degrees="04.54822" direction="north" orientation="latitude" pageId="5" pageNumber="16" precision="1" value="4.54822">
|
||
04.54822
|
||
<emphasis bold="true" box="[1093,1105,860,890]" pageId="5" pageNumber="16">°</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1136,1269,860,890]" degrees="115.15823" direction="east" orientation="longitude" pageId="5" pageNumber="16" precision="1" value="115.15823">
|
||
115.15823
|
||
<emphasis bold="true" box="[1239,1251,860,890]" pageId="5" pageNumber="16">°</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
,
|
||
<collectionCode box="[1276,1325,865,889]" pageId="5" pageNumber="16">ALS</collectionCode>
|
||
<specimenCode box="[1329,1403,865,889]" collectionCode="ALS" pageId="5" pageNumber="16">B43‐08</specimenCode>
|
||
,
|
||
<date pageId="5" pageNumber="16" value="2008-07-11" valueMax="2008-07-16" valueMin="2008-07-11">
|
||
<collectingDate pageId="5" pageNumber="16" value="2008-07-11" valueMax="2008-07-16" valueMin="2008-07-11">11‐ 16 July 2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[973,1113,900,925]" pageId="5" pageNumber="16">A.L. Sheldon</collectorName>
|
||
(
|
||
<collectionCode box="[1126,1207,901,925]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="16" type="Museum">USNM</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="16" type="description">
|
||
<paragraph blockId="5.[835,1445,791,1885]" pageId="5" pageNumber="16">
|
||
<emphasis bold="true" box="[835,1000,936,960]" pageId="5" pageNumber="16">Adult habitus.</emphasis>
|
||
Body color pale white in alcohol. Head without distinctive pigment pattern; ocelli almost touching.Antennal bases pale but flagellum beyond basal segments, dark brown. Pronotum with pale brown rugosities and darker marginal and median sutures.Tibiae about as wide as femora and slightly darker. Wings pale with pale yellow veins.
|
||
</paragraph>
|
||
<paragraph blockId="5.[835,1445,791,1885]" pageId="5" pageNumber="16">
|
||
<emphasis bold="true" box="[835,899,1185,1209]" pageId="5" pageNumber="16">Male.</emphasis>
|
||
Forewing length
|
||
<quantity box="[1096,1182,1185,1209]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="16" unit="mm" value="7.5">7.5 mm</quantity>
|
||
. Tergum 8 unmodified, tergum 9 with a pair of lateral sensilla basiconica patches and a median patch of fine, short setae (
|
||
<figureCitation box="[1244,1313,1256,1280]" captionStart="Figs" captionStartId="6.[189,240,1434,1458]" captionTargetBox="[324,1344,217,1354]" captionTargetId="figure-254@6.[324,1307,217,1354]" captionTargetPageId="6" captionText="Figs. 11‐14. Phanoperla astrospinata (11‐12) and P. belalong (13‐14). 11. Male terminalia, 12. Aedeagus, lateral aspect, 13. Male terminalia, 14. Aedeagus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4759032" httpUri="https://zenodo.org/record/4759032/files/figure.png" pageId="5" pageNumber="16">Fig.11</figureCitation>
|
||
).Hemiterga short and relatively wide with rounded tips in dorsal aspect; apices hooked downward in lateral aspect and bearing a few enlarged sensilla basiconica; notch separating basal callus narrow and deep. Aedeagal tube shortandplump with a fewfine setalspineson bulb; apex swollen and heavily armed with spines (
|
||
<figureCitation box="[1274,1347,1469,1493]" captionStart="Figs" captionStartId="6.[189,240,1434,1458]" captionTargetBox="[324,1344,217,1354]" captionTargetId="figure-254@6.[324,1307,217,1354]" captionTargetPageId="6" captionText="Figs. 11‐14. Phanoperla astrospinata (11‐12) and P. belalong (13‐14). 11. Male terminalia, 12. Aedeagus, lateral aspect, 13. Male terminalia, 14. Aedeagus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4759032" httpUri="https://zenodo.org/record/4759032/files/figure.png" pageId="5" pageNumber="16">Fig. 12</figureCitation>
|
||
); Largest and most prominent armature consists of a subapical ring of ca. 22 black spines and a pair of star‐ like, lateral clusters. Additional small, triangular spines occur over a broad zone, proximal to star clusters and on apical zone distal to spine ring. Frontal area below spine ring, bare and produced into a membranous nose‐ like process. Dorsal sclerite slender and poorly sclerotized.
|
||
</paragraph>
|
||
<paragraph blockId="5.[835,1445,791,1885]" box="[835,1045,1754,1778]" pageId="5" pageNumber="16">
|
||
<emphasis bold="true" box="[835,923,1754,1778]" pageId="5" pageNumber="16">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph blockId="5.[835,1445,791,1885]" box="[835,1027,1789,1814]" pageId="5" pageNumber="16">
|
||
<emphasis bold="true" box="[835,904,1790,1814]" pageId="5" pageNumber="16">Larva.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="16" type="etymology">
|
||
<paragraph blockId="5.[835,1445,791,1885]" pageId="5" pageNumber="16">
|
||
<emphasis bold="true" box="[835,963,1825,1849]" pageId="5" pageNumber="16">Etymology.</emphasis>
|
||
The species name refers to the star pattern of large spines on the aedeagus sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4759032" ID-Zenodo-Dep="4759032" httpUri="https://zenodo.org/record/4759032/files/figure.png" pageId="6" pageNumber="17" startId="6.[189,240,1434,1458]" targetBox="[324,1344,217,1354]" targetPageId="6">
|
||
<paragraph blockId="6.[189,1443,1434,1494]" pageId="6" pageNumber="17">
|
||
Figs. 11‐14.
|
||
<taxonomicName authorityName="Stark & Sheldon" authorityYear="2009" box="[328,587,1434,1458]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="astrospinata">
|
||
<emphasis box="[328,587,1434,1458]" italics="true" pageId="6" pageNumber="17">Phanoperla astrospinata</emphasis>
|
||
</taxonomicName>
|
||
(11‐12) and
|
||
<taxonomicName authorityName="Stark & Sheldon" authorityYear="2009" box="[735,854,1434,1458]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="belalong">
|
||
<emphasis box="[735,854,1434,1458]" italics="true" pageId="6" pageNumber="17">P. belalong</emphasis>
|
||
</taxonomicName>
|
||
(13‐14). 11. Male terminalia, 12. Aedeagus, lateral aspect, 13. Male terminalia, 14. Aedeagus, lateral aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="7" lastPageNumber="18" pageId="6" pageNumber="17" type="diagnosis">
|
||
<paragraph blockId="6.[189,798,1576,1921]" lastBlockId="7.[189,715,227,251]" lastPageId="7" lastPageNumber="18" pageId="6" pageNumber="17">
|
||
<emphasis bold="true" box="[189,318,1576,1600]" pageId="6" pageNumber="17">Diagnosis.</emphasis>
|
||
The aedeagus and external genitalia of this species is generally similar to that of
|
||
<taxonomicName authority="(Zwick 1982)" baseAuthorityName="Zwick" baseAuthorityYear="1982" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="flaveola">
|
||
<emphasis box="[685,797,1612,1636]" italics="true" pageId="6" pageNumber="17">P. flaveola</emphasis>
|
||
(
|
||
<bibRefCitation author="Zwick, P." box="[197,334,1648,1672]" pageId="6" pageNumber="17" pagination="87 - 126" refId="ref4786" refString="Zwick, P. 1982. A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae). Systematic Entomology, 7: 87 - 126." type="journal article" year="1982">Zwick 1982</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and it appears to be a member of that group. These structures are also generally similar to, or perhaps the same as,
|
||
<taxonomicName authorityName="P.Zwick" authorityYear="1982" box="[494,763,1719,1743]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis box="[494,615,1719,1743]" italics="true" pageId="6" pageNumber="17">Phanoperla</emphasis>
|
||
cf.
|
||
<emphasis box="[665,763,1719,1743]" italics="true" pageId="6" pageNumber="17">maculata</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<bibRefCitation author="Zwick, P." box="[189,357,1754,1778]" pageId="6" pageNumber="17" pagination="151 - 158" refId="ref4845" refString="Zwick, P. 1986 b. Contribution to the knowledge of Phanoperla Banks, 1938 (Plecoptera: Perlidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 59: 151 - 158." type="journal article" year="1986">Zwick (1986b)</bibRefCitation>
|
||
. That designation was based on a pair of Bornean specimens from the 1932 Oxford University Expedition studied by
|
||
<bibRefCitation author="Zwick, P." box="[581,748,1825,1849]" pageId="6" pageNumber="17" pagination="151 - 158" refId="ref4845" refString="Zwick, P. 1986 b. Contribution to the knowledge of Phanoperla Banks, 1938 (Plecoptera: Perlidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 59: 151 - 158." type="journal article" year="1986">Zwick (1986b)</bibRefCitation>
|
||
, but whose status was not formally defined. The present specimen has the same general pattern of spines on the aedeagus, however there are fewer large lateral spines and they are arranged in a more distinct star pattern than in Zwick’s specimens. In addition, fine triangular spines are shown by
|
||
<bibRefCitation author="Zwick, P." box="[1205,1371,1683,1707]" pageId="6" pageNumber="17" pagination="151 - 158" refId="ref4845" refString="Zwick, P. 1986 b. Contribution to the knowledge of Phanoperla Banks, 1938 (Plecoptera: Perlidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 59: 151 - 158." type="journal article" year="1986">Zwick (1986b)</bibRefCitation>
|
||
along the frontal zone, and no nose shaped process is shown for his specimens. The new species is also distinguished from
|
||
<taxonomicName baseAuthorityName="Zwick" baseAuthorityYear="1982" box="[1072,1185,1790,1814]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="flaveola">
|
||
<emphasis box="[1072,1185,1790,1814]" italics="true" pageId="6" pageNumber="17">P. flaveola</emphasis>
|
||
</taxonomicName>
|
||
by the star cluster of lateral aedeagal spines. The lateral grouping of that species forms a linear group of 5‐6 large spines, and in addition, the dorsal aedeagal sclerite is much larger for
|
||
<taxonomicName baseAuthorityName="Zwick" baseAuthorityYear="1982" box="[306,417,227,251]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="flaveola">
|
||
<emphasis box="[306,417,227,251]" italics="true" pageId="7" pageNumber="18">P. flaveola</emphasis>
|
||
</taxonomicName>
|
||
than for the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |