275 lines
44 KiB
XML
275 lines
44 KiB
XML
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<mods:title id="1FC4C9FE91A82221C64A5C33289D6014">A New Find Of Discosauriscid Seymouriamorph From The Lower Permian Of Boskovice Basin In Moravia (The Czech Republic)</mods:title>
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<mods:namePart id="5C8F80B80D7A8742E7A17CFE99EDE0EE">Klembara, Jozef D.</mods:namePart>
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<paragraph id="8BB5581EFFF4FFBD2A43FC4DFDEF9EEA" blockId="1.[295,556,985,1008]" box="[295,556,985,1008]" pageId="1" pageNumber="118">
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<heading id="D0FDEF72FFF4FFBD2A43FC4DFDEF9EEA" box="[295,556,985,1008]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="118" reason="2">
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<emphasis id="B97E840CFFF4FFBD2A43FC4DFDEF9EEA" bold="true" box="[295,556,985,1008]" pageId="1" pageNumber="118">
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<taxonomicName id="4C0A239DFFF4FFBD2A43FC4DFE269EEA" authorityName="ROMER" authorityYear="1947" box="[295,485,985,1008]" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="family">Discosauriscidae</taxonomicName>
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indet.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB5581EFFF4FFBD2BF4FB98FD7B999E" blockId="1.[106,745,1036,1156]" pageId="1" pageNumber="118">
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M a t e r i a l. DE K 396, skull and anterior portion of the postcranial skeleton, is the only known specimen and is deposited in the Department of Ecology, Faculty of Natural Sciences, Comenius University in Bratislava,
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<collectingCountry id="F31D188EFFF4FFBD2937FBF9FD77999E" box="[595,692,1133,1156]" name="Slovakia" pageId="1" pageNumber="118">Slovakia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3100B95FFF4FFBF2BF4FB08FDDC9A3F" lastPageId="3" lastPageNumber="120" pageId="1" pageNumber="118" type="description">
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<paragraph id="8BB5581EFFF4FFBD2BF4FB08FE8A9829" blockId="1.[106,745,1180,1331]" pageId="1" pageNumber="118">
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L o c a l i t y a n d h o r i z o n. Boskovice Basin in
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<collectingRegion id="49CE96FCFFF4FFBD2B0EFB28FF0599C9" box="[106,198,1212,1235]" country="Czech Republic" name="Jihomoravsky" pageId="1" pageNumber="118">Moravia</collectingRegion>
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(the
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<collectingCountry id="F31D188EFFF4FFBD2A6FFB28FE0099C9" box="[267,451,1212,1235]" name="Czech Republic" pageId="1" pageNumber="118">Czech Republic</collectingCountry>
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); Kochov-Horka, western margin of the town Letovice (
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<bibRefCitation id="EF9B25EFFFF4FFBD2AFAFB48FD1E99E9" author="Klembara, J. & Meszaros, S." box="[414,733,1244,1267]" pageId="1" pageNumber="118" pagination="305 - 312" refId="ref3193" refString="Klembara, J., Meszaros, S. (1992): New finds of Discosauriscus austriacus (Makowsky 1876) from the Lower Permian of the Boskovice Furrow (Czecho-Slovakia). - Geologica Carpathica, 43: 305 - 312." type="journal article" year="1992">Klembara and Meszároš 1992</bibRefCitation>
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). Middle Letovice Formation, Autunian, Lower Permian (
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<bibRefCitation id="EF9B25EFFFF4FFBD29D5FB68FEFE9829" author="Zajic, J. & Stamberg, S." pageId="1" pageNumber="118" pagination="5 - 14" refId="ref3753" refString="Zajic, J., Stamberg, S. (2004): Selected important fossiliferous horizons of the Boskovice Basin in the light of the new zoopaleontological data. - Acta Musei Reginaehradecensis, series A, 30: 5 - 14." type="journal article" year="2004">Zajíc and Štamberg 2004</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BB5581EFFF4FFBD2BF4FAD8FF029B59" blockId="1.[106,745,1356,1603]" pageId="1" pageNumber="118">
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D i a g n o s i s. Autapomorphies relative to discosauriscids
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<taxonomicName id="4C0A239DFFF4FFBD2BFEFAF8FEFE9899" authorityName="KUHN" authorityYear="1933" box="[154,317,1388,1411]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">
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<emphasis id="B97E840CFFF4FFBD2BFEFAF8FEFE9899" box="[154,317,1388,1411]" italics="true" pageId="1" pageNumber="118">Discosauriscus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A239DFFF4FFBD2A2EFAF8FE009899" authorityName="Klembara" authorityYear="2005" box="[330,451,1388,1411]" class="Amphibia" family="Discosauriscidae" genus="Makowskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">
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<emphasis id="B97E840CFFF4FFBD2A2EFAF8FE009899" box="[330,451,1388,1411]" italics="true" pageId="1" pageNumber="118">Makowskia</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A239DFFF4FFBD2AABFAF8FDA49899" authorityName="KLEMBARA" authorityYear="2009" box="[463,615,1388,1411]" class="Amphibia" family="Discosauriscidae" genus="Spinarerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">
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<emphasis id="B97E840CFFF4FFBD2AABFAF8FDA49899" box="[463,615,1388,1411]" italics="true" pageId="1" pageNumber="118">Spinarerpeton</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0A239DFFF4FFBD29FDFAF8FF0898B9" authorityName="Ivakhnenko" authorityYear="1981" class="Amphibia" family="Discosauriscidae" genus="Ariekanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">
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<emphasis id="B97E840CFFF4FFBD29FDFAF8FF0898B9" italics="true" pageId="1" pageNumber="118">Ariekanerpeton</emphasis>
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</taxonomicName>
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include: 1) strong vermiculate ornamentation of the skull roof bones; 2) intertemporal larger than the supratemporal; and 3) a small process of parietal lying between the postfrontal and intertemporal is missing, thus the anterolateral margin of the parietal is long and almost straight.
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</paragraph>
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<paragraph id="8BB5581EFFF4FFBD2BF4F9C8FED29AAE" blockId="1.[106,745,1628,2100]" pageId="1" pageNumber="118">
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D e s c r i p t i o n. The bones of the skull roof are massively constructed and the sutures between them are mostly simple (
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<figureCitation id="1331449BFFF4FFBD2A77F908FE569BA9" box="[275,405,1692,1715]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="1" pageNumber="118">Text-fig. 1a</figureCitation>
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). The shape of any given bone may differ on the left and right sides. The ornamentation of all bones is conspicuously developed. It consists of high ridges of different lengths and course, and deep grooves and pits. Such ridges form a pronounced polygonal pattern. Due to this strong ornamentation, the ossification centres are not clearly recognizable. Deep lateral line canals are present on the left intertemporal and postfrontal and right postfrontal and postorbital.
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</paragraph>
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<paragraph id="8BB5581EFFF4FFBD2BF4F829FB4D9CB5" blockId="1.[106,745,1628,2100]" lastBlockId="1.[820,1460,152,1703]" pageId="1" pageNumber="118">
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The frontal is a mediolaterally narrow bone and is longer then the parietal (
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<figureCitation id="1331449BFFF4FFBD2A43F849FE079AEE" box="[295,452,2013,2036]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="1" pageNumber="118">Text-fig. 1a, b</figureCitation>
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). The prefrontal has a long posterior process and it joins the long anterior process of the postfrontal at a level slightly anteriorly to the mid-length of the frontal. The prefrontal, postfrontal and postorbital bear sharp orbital margins. The orbital margin is highest on the right postorbital. The right postorbital is of triradiate shape; the posterior process is the shortest and fits between the intertemporal and squamosal (
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<figureCitation id="1331449BFFF4FFBD2F09FE8CFAC09C35" box="[1133,1283,280,303]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="1" pageNumber="118">Text-fig. 1a, b</figureCitation>
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). The postorbital is subdivided; there is a small, independent bone, lying within the territory of the posteromedial portion of postorbital 1 (designated as postorbital 2 here,
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<figureCitation id="1331449BFFF4FFBD2FC2FEECFAE49C95" box="[1190,1319,376,399]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="1" pageNumber="118">Text-fig. 1a</figureCitation>
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). The lateral end of the postorbital is pointed.
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</paragraph>
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<paragraph id="8BB5581EFFF4FFBD283EFE2CFB0B9813" blockId="1.[820,1460,152,1703]" pageId="1" pageNumber="118">
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The anterior portion of the left parietal is mediolaterally wider than that of the right parietal; however, it is opposite in the posterior portion of the parietal (
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<figureCitation id="1331449BFFF4FFBD2F89FE63FAB19F14" box="[1261,1394,503,526]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="1" pageNumber="118">Text-fig. 1a</figureCitation>
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). The anterolateral margin of the parietal is long and almost straight (only very shallowly concave). It has a long suture with the intertemporal but only a very short one with the postfrontal. On the right side, the parietal-postfrontal suture is slightly longer. The pineal foramen is rounded and not closed by the parietals anteriorly. Posteriorly to the pineal foramen, the suture between the parietals is straight. The left intertemporal is a large pentagonal-shaped bone, and is larger than the left supratempotal. The right intertemporal has an approximately oval shape. As preserved, the right intertemporal is mediolaterally compressed and dorsally vaulted, giving the impression that it is smaller than the supratemporal in dorsal view. However, its reconstruction shows it is larger in size than the right supratemporal (
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<figureCitation id="1331449BFFF4FFBD2F03FC20FB3E9ED1" box="[1127,1277,948,971]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="1" pageNumber="118">Text-fig. 1a, b</figureCitation>
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). There is a small oval bone present between the right prefrontal and intertemporal bones (
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<figureCitation id="1331449BFFF4FFBD2F7DFC60FB589911" box="[1049,1179,1012,1035]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="1" pageNumber="118">Text-fig. 1b</figureCitation>
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). Such small bones, lying within the territory of the given bone or between the territories of two or three bones, are quite common within the skull roof bones of discosauriscisds (
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<bibRefCitation id="EF9B25EFFFF4FFBD2F9BFBC7FA6C9970" author="Klembara, J." box="[1279,1455,1107,1130]" pageId="1" pageNumber="118" pagination="145 - 168" refId="ref2887" refString="Klembara, J. (1993): The subdivisions and fusions of the exoskeletal skull bones of Discosauriscus austriacus (Makowsky 1876) and their possible homologues in rhipidistians. - Palaontologische Zeitschrift, 67: 145 - 168. http: // dx. doi. org / 10.1007 / BF 02985875" type="journal article" year="1993">Klembara 1993</bibRefCitation>
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,
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<bibRefCitation id="EF9B25EFFFF4FFBD2850FBE7FBE49990" author="Klembara, J & Tomasik, A. & Kathe, W." box="[820,1063,1139,1162]" pageId="1" pageNumber="118" pagination="317 - 349" refId="ref3367" refString="Klembara, J, Tomasik, A., Kathe, W. (2002): Subdivisions, fusions and extended sutural areas of dermal skull bones in Discosauriscus Kuhn (Seymouriamorpha). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 223: 317 - 349." type="journal article" year="2002">Klembara et al. 2002</bibRefCitation>
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). The posterior portion of the right intertemporal extends into a small process fitting into the anterior portion of the supratemporal. The supratemporal is quadrangular. Its suture with the parietal is slightly shorter than the intertemporal-parietal suture.
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</paragraph>
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<paragraph id="8BB5581EFFF4FFBD283EFA86FC2A98F2" blockId="1.[820,1460,152,1703]" pageId="1" pageNumber="118">The tabulars and postparietals are mediolaterally elongated. The tabular process is of quadrangular shape and extends posteriorly from the ventral surface of the tabular. The tabular occipital flange is orientated posteroventrally and has a smooth dorsal surface. It joins medially the slightly smaller occipital flange of the postparietal. The tabular has a suture with the parietal.</paragraph>
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<paragraph id="8BB5581EFFF4FFBD283EFA65FCBB9BBD" blockId="1.[820,1460,152,1703]" pageId="1" pageNumber="118">Approximately only the posterior half of the right maxilla is preserved. On its internal surface, the supradental shelf and several teeth are observable. The teeth are conical, slightly posteriorly curved and their basal portions bear dorsoventral grooves. The tooth crowns have medial and distal cutting edges.</paragraph>
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<paragraph id="8BB5581EFFF4FFBD283EF92AFC659A4F" blockId="1.[820,1459,1726,2099]" pageId="1" pageNumber="118">
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C o m p a r i s o n s. So far, three genera and four species belonging to the family
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<taxonomicName id="4C0A239DFFF4FFBD2F3FF94AFC519A0F" authority="(Seymouriamorpha)" authorityName="ROMER" authorityYear="1947" baseAuthorityName="Seymouriamorpha" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="family">Discosauriscidae (Seymouriamorpha)</taxonomicName>
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have been described from the Lower Permian localities of the Boskovice Basin in
|
||
<collectingRegion id="49CE96FCFFF4FFBD2FB0F88AFAF39A2F" box="[1236,1328,1822,1845]" country="Czech Republic" name="Jihomoravsky" pageId="1" pageNumber="118">Moravia</collectingRegion>
|
||
(the
|
||
<collectingCountry id="F31D188EFFF4FFBD2E0BF88AFC599A4F" name="Czech Republic" pageId="1" pageNumber="118">Czech Republic</collectingCountry>
|
||
):
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF4FFBD283EF8C9FAA59A6E" blockId="1.[820,1459,1726,2099]" box="[858,1382,1885,1908]" pageId="1" pageNumber="118">
|
||
1)
|
||
<taxonomicName id="4C0A239DFFF4FFBD2813F8C9FAA59A6E" authority="(MAKOWSKY, 1876)" baseAuthorityName="MAKOWSKY" baseAuthorityYear="1876" box="[887,1382,1885,1908]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="species" species="austriacus">
|
||
<emphasis id="B97E840CFFF4FFBD2813F8C9FB4C9A6E" box="[887,1167,1885,1908]" italics="true" pageId="1" pageNumber="118">Discosauriscus austriacus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9B25EFFFF4FFBD2FFAF8C9FA9D9A6E" author="Makowsky, A." box="[1182,1374,1885,1908]" pageId="1" pageNumber="118" pagination="155 - 166" refId="ref3450" refString="Makowsky, A. (1876): Uber einen neuen Labyrinthodonten: " Archegosaurus austriacus nov. spec. ". - Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe, I. Abtheilung, 73: 155 - 166." type="journal article" year="1876">MAKOWSKY, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF4FFBD283EF8E9FAA39A8E" blockId="1.[820,1459,1726,2099]" box="[858,1376,1917,1940]" pageId="1" pageNumber="118">
|
||
2)
|
||
<emphasis id="B97E840CFFF4FFBD2813F8E9FB6D9A8E" box="[887,1198,1917,1940]" italics="true" pageId="1" pageNumber="118">
|
||
<taxonomicName id="4C0A239DFFF4FFBD2813F8E9FBD99A8E" authorityName="KUHN" authorityYear="1933" box="[887,1050,1917,1940]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">Discosauriscus</taxonomicName>
|
||
pulcherrimus
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9B25EFFFF4FFBD2FD9F8E9FA9A9A8E" author="Fritsch, A." box="[1213,1369,1917,1940]" pageId="1" pageNumber="118" pagination="93 - 126" refId="ref2852" refString="Fritsch, A. (1880): Fauna der Gaskohle und der Kalksteine der Permformation Bohmens. I / 2. - F. Rivnac, Prag, pp. 93 - 126." type="book chapter" year="1880">FRITSCH, 1880</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF4FFBD283EF809FAEE9AAE" blockId="1.[820,1459,1726,2099]" box="[858,1325,1949,1972]" pageId="1" pageNumber="118">
|
||
3)
|
||
<emphasis id="B97E840CFFF4FFBD2813F809FBAC9AAE" box="[887,1135,1949,1972]" italics="true" pageId="1" pageNumber="118">
|
||
<taxonomicName id="4C0A239DFFF4FFBD2813F809FC339AAE" authorityName="Klembara" authorityYear="2005" box="[887,1008,1949,1972]" class="Amphibia" family="Discosauriscidae" genus="Makowskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">Makowskia</taxonomicName>
|
||
laticephala
|
||
</emphasis>
|
||
KLEMBARA, 2005
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF4FFBD283EF829FA479ACE" blockId="1.[820,1459,1726,2099]" box="[858,1412,1981,2004]" pageId="1" pageNumber="118">
|
||
4)
|
||
<emphasis id="B97E840CFFF4FFBD2813F829FB719ACE" box="[887,1202,1981,2004]" italics="true" pageId="1" pageNumber="118">
|
||
<taxonomicName id="4C0A239DFFF4FFBD2813F829FBCC9ACE" authorityName="KLEMBARA" authorityYear="2009" box="[887,1039,1981,2004]" class="Amphibia" family="Discosauriscidae" genus="Spinarerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">Spinarerpeton</taxonomicName>
|
||
brevicephalum
|
||
</emphasis>
|
||
<bibRefCitation id="EF9B25EFFFF4FFBD2FDDF829FA439ACE" author="Klembara, J." box="[1209,1408,1981,2004]" pageId="1" pageNumber="118" pagination="451 - 484" refId="ref3094" refString="Klembara, J. (2009 a): The skeletal anatomy and relationships of a new discosauriscid seymouriamorph from the Lower Permian of Moravia (Czech Republic). - Annals of Carnegie Museum, 77: 451 - 484. http: // dx. doi. org / 10.2992 / 0097 - 4463 - 77.4.451" type="journal article" year="2009">KLEMBARA, 2009a</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF4FFBD283EF848FA4A9529" blockId="1.[820,1459,1726,2099]" pageId="1" pageNumber="118">
|
||
The fifth species belonging to
|
||
<taxonomicName id="4C0A239DFFF4FFBD2FB5F848FA579AE9" authorityName="ROMER" authorityYear="1947" box="[1233,1428,2012,2035]" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="family">Discosauriscidae</taxonomicName>
|
||
is
|
||
<emphasis id="B97E840CFFF4FFBD2850F868FBF59509" box="[820,1078,2044,2067]" italics="true" pageId="1" pageNumber="118">
|
||
<taxonomicName id="4C0A239DFFF4FFBD2850F868FC199509" authorityName="Ivakhnenko" authorityYear="1981" box="[820,986,2044,2067]" class="Amphibia" family="Discosauriscidae" genus="Ariekanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="1" pageNumber="118" phylum="Chordata" rank="genus">Ariekanerpeton</taxonomicName>
|
||
sigalovi
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9B25EFFFF4FFBD2F21F868FB399509" author="Tatarinov, L. P." box="[1093,1274,2044,2067]" pageId="1" pageNumber="118" pagination="73 - 92" refId="ref3613" refString="Tatarinov, L. P. (1968): Verkhnepaleozoyskie i mezozoyskie zemnovodnye i presmykayutschiesya SSSR [Upper Paleozoic and Mesozoic amphibians and reptiles of USSR]. - In: Kalandadze, N. N., Otschev, V. G., Tatarinov, L. P., Tschudinov, P. K., Shishkin, M. A. (eds), Katalog permskikh i triasovykh tetrapod SSSR [Catalogue of Permian and Triassic tetrapods of USSR], Nauka, Moskva [Moscow], pp. 73 - 92. (in Russian)" type="book chapter" year="1968">TATARINOV, 1968</bibRefCitation>
|
||
) from the Lower Permian of
|
||
<collectingCountry id="F31D188EFFF4FFBD28D4F788FBFB9529" box="[944,1080,2076,2099]" name="Tajikistan" pageId="1" pageNumber="118">Tadzhikistan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9B25EFFFF4FFBD2F22F788FAA29529" author="Klembara, J. & Ruta, M." box="[1094,1377,2076,2099]" pageId="1" pageNumber="118" pagination="43 - 70" refId="ref3237" refString="Klembara, J., Ruta, M. (2005 a): The seymouriamorph tetrapod Ariekanerpeton sigalovi from the Lower Permian of Tadzhikistan. Part I. Cranial anatomy and ontogeny. - Transactions of the Royal Society of Edinburgh, Earth Sciences, 96: 43 - 70. http: // dx. doi. org / 10.1017 / S 0263593300001231" type="journal article" year="2005">Klembara and Ruta 2005a</bibRefCitation>
|
||
, b).
|
||
</paragraph>
|
||
<caption id="DF750896FFF7FFBE2BE5F870FBAC9529" ID-DOI="http://doi.org/10.5281/zenodo.5387844" ID-Zenodo-Dep="5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="2" pageNumber="119" startId="2.[129,211,2020,2041]" targetBox="[129,1482,152,1979]" targetPageId="2">
|
||
<paragraph id="8BB5581EFFF7FFBE2BE5F870FBAC9529" blockId="2.[129,1482,2020,2099]" pageId="2" pageNumber="119">
|
||
Text-fig. 1. a –
|
||
<taxonomicName id="4C0A239DFFF7FFBE2A7BF870FE089AE3" authorityName="ROMER" authorityYear="1947" box="[287,459,2020,2041]" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="2" pageNumber="119" phylum="Chordata" rank="family">Discosauriscidae</taxonomicName>
|
||
indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of
|
||
<taxonomicName id="4C0A239DFFF7FFBE2AE2F795FD4B950C" baseAuthorityName="MAKOWSKY" baseAuthorityYear="1876" box="[390,648,2049,2070]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="2" pageNumber="119" phylum="Chordata" rank="species" species="austriacus">
|
||
<emphasis id="B97E840CFFF7FFBE2AE2F795FD4B950C" bold="true" box="[390,648,2049,2070]" italics="true" pageId="2" pageNumber="119">Discosauriscus austriacus</emphasis>
|
||
</taxonomicName>
|
||
in dorsal view (on the basis of specimen
|
||
<accessionNumber id="9459C5FDFFF7FFBE2F41F795FBB4950C" box="[1061,1143,2049,2070]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/Z15529" pageId="2" pageNumber="119" type="EnaNcbi">Z 15529</accessionNumber>
|
||
from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia;
|
||
<bibRefCitation id="EF9B25EFFFF7FFBE28A0F78AFBA69529" author="Klembara, J." box="[964,1125,2078,2099]" pageId="2" pageNumber="119" pagination="257 - 302" refId="ref2997" refString="Klembara, J. (1997): The cranial anatomy of Discosauriscus Kuhn, a seymouriamorph tetrapod from the Lower Permian of the Boskovice Furrow (Czech Republic). - Philosophical Transactions of the Royal Society of London, B, 352: 257 - 302. http: // dx. doi. org / 10.1098 / rstb. 1997.0021" type="journal article" year="1997">Klembara 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB5581EFFF6FFBF2BF4FF0DFD6A9C15" blockId="3.[106,745,153,876]" pageId="3" pageNumber="120">
|
||
Among the discosauriscids, the skull of DE K 396 is most similar to that of
|
||
<taxonomicName id="4C0A239DFFF6FFBF2A4EFF2DFD8B9DCA" baseAuthorityName="MAKOWSKY" baseAuthorityYear="1876" box="[298,584,185,208]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="austriacus">
|
||
<emphasis id="B97E840CFFF6FFBF2A4EFF2DFD8B9DCA" box="[298,584,185,208]" italics="true" pageId="3" pageNumber="120">Discosauriscus austriacus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1331449BFFF6FFBF293CFF2DFD199DCA" box="[600,730,185,208]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="3" pageNumber="120">Text-fig. 1c</figureCitation>
|
||
). The specimen DE K 396 can be distinguished from all other species of
|
||
<taxonomicName id="4C0A239DFFF6FFBF2BBEFF6CFE4C9C15" authorityName="ROMER" authorityYear="1947" box="[218,399,248,271]" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="family">Discosauriscidae</taxonomicName>
|
||
by the following features:
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF6FFBF2BF4FE8CFF6C9F77" blockId="3.[106,745,153,876]" pageId="3" pageNumber="120">
|
||
Distinct ornamentation of the skull roofing bones. The ornamentation consists of high and mostly sharp ridges joined together forming a polygonal pattern. Such distinct ornamentation is developed neither in the similar-sized specimens of
|
||
<taxonomicName id="4C0A239DFFF6FFBF2A65FE03FDDC9CB4" baseAuthorityName="MAKOWSKY" baseAuthorityYear="1876" box="[257,543,407,430]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="austriacus">
|
||
<emphasis id="B97E840CFFF6FFBF2A65FE03FDDC9CB4" box="[257,543,407,430]" italics="true" pageId="3" pageNumber="120">Discosauriscus austriacus</emphasis>
|
||
</taxonomicName>
|
||
, nor in the largest known specimens of
|
||
<taxonomicName id="4C0A239DFFF6FFBF2A28FE23FE1F9CD4" baseAuthorityName="MAKOWSKY" baseAuthorityYear="1876" box="[332,476,439,462]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="austriacus">
|
||
<emphasis id="B97E840CFFF6FFBF2A28FE23FE1F9CD4" box="[332,476,439,462]" italics="true" pageId="3" pageNumber="120">D. austriacus</emphasis>
|
||
</taxonomicName>
|
||
(with the skull length up to about
|
||
<quantity id="4CF2F5FBFFF6FFBF2BA7FE43FECC9CF4" box="[195,271,471,494]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="120" unit="mm" value="62.0">62 mm</quantity>
|
||
;
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2A7DFE43FE7E9CF4" author="Klembara, J." box="[281,445,471,494]" pageId="3" pageNumber="120" pagination="265 - 281" refId="ref2941" refString="Klembara, J. (1995): The external gills and ornamentation of skull roof bones of the Lower Permian tetrapod Discosauriscus (Kuhn 1933) with remarks to its ontogeny. - Palaontologische Zeitschrift, 69: 265 - 281. http: // dx. doi. org / 10.1007 / BF 02985990" type="journal article" year="1995">Klembara 1995</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2AA2FE43FE3E9CF4" author="Klembara, J." box="[454,509,471,494]" pageId="3" pageNumber="120" pagination="257 - 302" refId="ref2997" refString="Klembara, J. (1997): The cranial anatomy of Discosauriscus Kuhn, a seymouriamorph tetrapod from the Lower Permian of the Boskovice Furrow (Czech Republic). - Philosophical Transactions of the Royal Society of London, B, 352: 257 - 302. http: // dx. doi. org / 10.1098 / rstb. 1997.0021" type="journal article" year="1997">1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2963FE43FD899CF4" author="Klembara, J." box="[519,586,471,494]" pageId="3" pageNumber="120" pagination="451 - 484" refId="ref3094" refString="Klembara, J. (2009 a): The skeletal anatomy and relationships of a new discosauriscid seymouriamorph from the Lower Permian of Moravia (Czech Republic). - Annals of Carnegie Museum, 77: 451 - 484. http: // dx. doi. org / 10.2992 / 0097 - 4463 - 77.4.451" type="journal article" year="2009">2009a</bibRefCitation>
|
||
, b). The degree of ossification and the
|
||
<typeStatus id="54B1E6BCFFF6FFBF2A3CFE63FE459F14" box="[344,390,503,526]" pageId="3" pageNumber="120">type</typeStatus>
|
||
of sutures on the skull roof bones of DE K 396 correspond to those of the similar-sized specimens of
|
||
<taxonomicName id="4C0A239DFFF6FFBF2A60FDA3FDE69F54" baseAuthorityName="MAKOWSKY" baseAuthorityYear="1876" box="[260,549,567,590]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="austriacus">
|
||
<emphasis id="B97E840CFFF6FFBF2A60FDA3FDE69F54" box="[260,549,567,590]" italics="true" pageId="3" pageNumber="120">Discosauriscus austriacus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2951FDA3FD269F54" author="Klembara, J." box="[565,741,567,590]" pageId="3" pageNumber="120" pagination="265 - 281" refId="ref2941" refString="Klembara, J. (1995): The external gills and ornamentation of skull roof bones of the Lower Permian tetrapod Discosauriscus (Kuhn 1933) with remarks to its ontogeny. - Palaontologische Zeitschrift, 69: 265 - 281. http: // dx. doi. org / 10.1007 / BF 02985990" type="journal article" year="1995">Klembara 1995</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2B0EFDC2FF609F77" author="Klembara, J." box="[106,163,598,621]" pageId="3" pageNumber="120" pagination="257 - 302" refId="ref2997" refString="Klembara, J. (1997): The cranial anatomy of Discosauriscus Kuhn, a seymouriamorph tetrapod from the Lower Permian of the Boskovice Furrow (Czech Republic). - Philosophical Transactions of the Royal Society of London, B, 352: 257 - 302. http: // dx. doi. org / 10.1098 / rstb. 1997.0021" type="journal article" year="1997">1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF6FFBF2BF4FDE2FD2A9FB7" blockId="3.[106,745,153,876]" pageId="3" pageNumber="120">
|
||
The intertemporal is larger than the supratemporal. This is in contrast to all other seymouriamorphs (
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2948FD02FD1D9FB7" author="Klembara, J." box="[556,734,662,685]" pageId="3" pageNumber="120" pagination="451 - 484" refId="ref3094" refString="Klembara, J. (2009 a): The skeletal anatomy and relationships of a new discosauriscid seymouriamorph from the Lower Permian of Moravia (Czech Republic). - Annals of Carnegie Museum, 77: 451 - 484. http: // dx. doi. org / 10.2992 / 0097 - 4463 - 77.4.451" type="journal article" year="2009">Klembara 2009a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF6FFBF2BF4FD22FF749E76" blockId="3.[106,745,153,876]" pageId="3" pageNumber="120">A small process of the parietal fitting between the postfrontal and intertemporal is absent. As a consequence, the anterolateral margin of the parietal is long and almost straight. This is clearly observable on both parietals. Such parietal morphology is not present in any known seymouriamorph.</paragraph>
|
||
<paragraph id="8BB5581EFFF6FFBF2BF4FC10FD5F98ED" blockId="3.[106,746,900,1527]" pageId="3" pageNumber="120">
|
||
R e c o n s t r u c t e d s k u l l. Although several bones of the skull roof are missing, it was possible to make a reconstruction at least in dorsal view (
|
||
<figureCitation id="1331449BFFF6FFBF294EFC57FD6F9EC0" box="[554,684,963,986]" captionStart="Text-fig" captionStartId="2.[129,211,2020,2041]" captionTargetBox="[129,1482,152,1979]" captionTargetId="figure-11@2.[152,1458,184,1956]" captionTargetPageId="2" captionText="Text-fig. 1. a – Discosauriscidae indet., DE K 396, skull in dorsal view. b – reconstruction of skull of DE K 396 in dorsal view. c – reconstruction of skull of Discosauriscus austriacus in dorsal view (on the basis of specimen Z 15529 from the locality Kochov-Horka, Boskovice Basin, deposited in the Slovak National Museum, Bratislava, Slovakia; Klembara 1997)." figureDoi="http://doi.org/10.5281/zenodo.5387844" httpUri="https://zenodo.org/record/5387844/files/figure.png" pageId="3" pageNumber="120">Text-fig. 1b</figureCitation>
|
||
).The skull is about as long as it is wide. It has a semi-elliptical shape with the long axis along the mid-line. The preorbital portion is short. The orbits are anteroposteriorly elongated and oval. They lie in the posterior portion of the anterior half of the skull length; only their posterior-most parts extend posterior to the mid-length of the skull. The prefrontal-postfrontal suture is short and lies slightly anteriorly to the mid-length of the frontals. The pineal foramen lies at the level of the anterior halves of the intertemporals and slightly posteriorly to the level of the posterior margins of the orbits. The anterior portions of both parietals are mediolaterally narrow due to the large size of the intertemporals. The intertemporal is larger than the supratemporal. The jaw joint lies at the level of the tabular. The otic notch is anteroposteriorly deep, similar to that in
|
||
<taxonomicName id="4C0A239DFFF6FFBF2B0EFA54FE1C98CD" authority="(Klembara 1997)" baseAuthorityName="Klembara" baseAuthorityYear="1997" box="[106,479,1472,1495]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97E840CFFF6FFBF2B0EFA54FED398CD" box="[106,272,1472,1495]" italics="true" pageId="3" pageNumber="120">Discosauriscus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2A40FA54FE1498CD" author="Klembara, J." box="[292,471,1472,1495]" pageId="3" pageNumber="120" pagination="257 - 302" refId="ref2997" refString="Klembara, J. (1997): The cranial anatomy of Discosauriscus Kuhn, a seymouriamorph tetrapod from the Lower Permian of the Boskovice Furrow (Czech Republic). - Philosophical Transactions of the Royal Society of London, B, 352: 257 - 302. http: // dx. doi. org / 10.1098 / rstb. 1997.0021" type="journal article" year="1997">Klembara 1997</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but not as deep as in
|
||
<taxonomicName id="4C0A239DFFF6FFBF2B0EFA74FD5A98ED" authority="KLEMBARA, 2009 a" authorityName="KLEMBARA" authorityYear="2009" box="[106,665,1504,1527]" class="Amphibia" family="Discosauriscidae" genus="Spinarerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97E840CFFF6FFBF2B0EFA74FEC198ED" box="[106,258,1504,1527]" italics="true" pageId="3" pageNumber="120">Spinarerpeton</emphasis>
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2A6CFA74FE0898ED" author="Klembara, J." box="[264,459,1504,1527]" pageId="3" pageNumber="120" pagination="451 - 484" refId="ref3094" refString="Klembara, J. (2009 a): The skeletal anatomy and relationships of a new discosauriscid seymouriamorph from the Lower Permian of Moravia (Czech Republic). - Annals of Carnegie Museum, 77: 451 - 484. http: // dx. doi. org / 10.2992 / 0097 - 4463 - 77.4.451" type="journal article" year="2009">KLEMBARA, 2009a</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EF9B25EFFFF6FFBF2ABDFA74FD5298ED" author="Klembara, J." box="[473,657,1504,1527]" pageId="3" pageNumber="120" pagination="451 - 484" refId="ref3094" refString="Klembara, J. (2009 a): The skeletal anatomy and relationships of a new discosauriscid seymouriamorph from the Lower Permian of Moravia (Czech Republic). - Annals of Carnegie Museum, 77: 451 - 484. http: // dx. doi. org / 10.2992 / 0097 - 4463 - 77.4.451" type="journal article" year="2009">Klembara 2009a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB5581EFFF6FFBF2BF4F99BFDDC9A3F" blockId="3.[106,745,1551,1829]" pageId="3" pageNumber="120">
|
||
C o n c l u s i o n s. Although the specimen DE K 396 is not yet fully prepared, its skull morphology clearly exhibits several features absent not only in
|
||
<taxonomicName id="4C0A239DFFF6FFBF2A86F9DBFD469B7C" baseAuthorityName="Klembara" baseAuthorityYear="1997" box="[482,645,1615,1638]" class="Amphibia" family="Discosauriscidae" genus="Discosauriscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97E840CFFF6FFBF2A86F9DBFD469B7C" box="[482,645,1615,1638]" italics="true" pageId="3" pageNumber="120">Discosauriscus</emphasis>
|
||
</taxonomicName>
|
||
, but also in other species of
|
||
<taxonomicName id="4C0A239DFFF6FFBF2A48F9FBFE229B9C" authorityName="ROMER" authorityYear="1947" box="[300,481,1647,1670]" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="family">Discosauriscidae</taxonomicName>
|
||
. Thus, it likely represents a new taxon. If so, this taxon will probably belong to
|
||
<taxonomicName id="4C0A239DFFF6FFBF2B0EF93AFEE59BDF" authorityName="ROMER" authorityYear="1947" box="[106,294,1710,1733]" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="family">Discosauriscidae</taxonomicName>
|
||
, because the skull of DE K 396 exhibits three of the four distinguishing features of
|
||
<taxonomicName id="4C0A239DFFF6FFBF2950F95AFD2A9BFF" authorityName="ROMER" authorityYear="1947" box="[564,745,1742,1765]" class="Amphibia" family="Discosauriscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seymouriamorpha" pageId="3" pageNumber="120" phylum="Chordata" rank="family">Discosauriscidae</taxonomicName>
|
||
(see the above Diagnosis). The taxonomic resolution of DE K 396 is pending its full preparation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |