393 lines
51 KiB
XML
393 lines
51 KiB
XML
<document id="3A96C9860BC5F78706F4FB34154703CC" ID-DOI="10.5281/zenodo.13191145" ID-ISSN="2533-4069" ID-Zenodo-Dep="13191145" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722606589039" checkinUser="felipe" docAuthor="Dedicated in memory of the late FrantiŠek Holý (1935 - 1984), an eminent Czech palaeobotanist, Holý, František, Kvaček, Zlatko & Teodoridis, Vasilis" docDate="2012" docId="03A3A81EFFB11925FF09FB9C3A63FD1A" docLanguage="en" docName="ActaMusNatPraSerB.68.3-4.53-118.pdf" docOrigin="Acta Musei Nationalis Pragae Series B 68 (3 - 4)" docStyle="DocumentStyle:2E5EC822A0C9CE74494687E2DCEF8E0C.4:ActaMusNatPraSerB.2007-2013.journal_article.0cover.type2" docStyleId="2E5EC822A0C9CE74494687E2DCEF8E0C" docStyleName="ActaMusNatPraSerB.2007-2013.journal_article.0cover.type2" docStyleVersion="4" docTitle="Leguminosites hradekensis KVACEK et TEODORIDIS 2012, comb. n." docType="treatment" docVersion="2" lastPageNumber="72" masterDocId="FF9AD066FFA31936FF98FFCA3F0FFF8B" masterDocTitle="A Review Of The Early Miocene Mastixioid Flora Of The Kristina Mine At Hrádek Nad Nisou In North Bohemia (The Czech Republic)" masterLastPageNumber="118" masterPageNumber="53" pageNumber="71" updateTime="1722673475371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="7E64698D0B360C41151B5D6B2FDEF542">A Review Of The Early Miocene Mastixioid Flora Of The Kristina Mine At Hrádek Nad Nisou In North Bohemia (The Czech Republic)</mods:title>
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<mods:namePart id="841A2148242C71C843B1D1FCA0E97F7F">Dedicated in memory of the late FrantiŠek Holý (1935 - 1984), an eminent Czech palaeobotanist</mods:namePart>
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<mods:namePart id="5026F5AF8A1C91F10C1ABBF00C807873">Holý, František</mods:namePart>
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<mods:affiliation id="FA9E5CDBAE51459A6E50340BE9AC4DE7">National Museum, Václavské nám. 68,115 79 Prague 1, the Czech Republic</mods:affiliation>
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<mods:namePart id="8BF898095A0E701C4C216842288A4EA9">Kvaček, Zlatko</mods:namePart>
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<mods:affiliation id="AD2BC103E160225BE571447B8C8D3D63">Institute of Geology and Palaeontology, Faculty of Science, Charles University in Prague, Albertov 6, 128 43 Prague 2, the Czech Republic, e-mail: kvacek @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="ABBDDB1A7DF44E07D562A8BD30E50496">Teodoridis, Vasilis</mods:namePart>
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<mods:affiliation id="4DCECDED56E2F08F9E8F8483D0C7923D">Department of Biology and Environmental Studies, Faculty of Education, Charles University in Prague, M. D. Rettigové 4, 116 39 Prague 1, the Czech Republic, e-mail: vasilis. teodoridis @ pedf. cuni. cz</mods:affiliation>
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<mods:title id="664B6CC6059483D1929E2BF42654EAE4">Acta Musei Nationalis Pragae Series B</mods:title>
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<mods:date id="F408E712853F8C4005EFF97629D15541">2012</mods:date>
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<mods:number id="AE4801BC8AE7DB427512ADD457C9621F">2012-12-31</mods:number>
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<mods:number id="BE6BF9C552895D5CDFABD4E7BA88C62A">68</mods:number>
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<mods:identifier id="4BF1A92A4941CD5908438586875B00EF" type="ISSN">2533-4069</mods:identifier>
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<treatment id="03A3A81EFFB11925FF09FB9C3A63FD1A" LSID="urn:lsid:plazi:treatment:03A3A81EFFB11925FF09FB9C3A63FD1A" httpUri="http://treatment.plazi.org/id/03A3A81EFFB11925FF09FB9C3A63FD1A" lastPageId="19" lastPageNumber="72" pageId="18" pageNumber="71">
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<subSubSection id="C3104A83FFB11924FF09FB9C3D54FB36" pageId="18" pageNumber="71" type="nomenclature">
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<paragraph id="8BB51908FFB11924FF09FB9C3D77FB06" blockId="18.[145,750,1110,1165]" pageId="18" pageNumber="71">
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<heading id="D0FDAE64FFB11924FF09FB9C3D77FB06" centered="true" fontSize="8" level="2" pageId="18" pageNumber="71" reason="2">
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<emphasis id="B97EC51AFFB11924FF09FB9C3D77FB06" bold="true" pageId="18" pageNumber="71">
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<taxonomicName id="4C0A628BFFB11924FF09FB9C3D07FB06" authority="(E. KNOBLOCH et KVACEK) KVACEK et TEODORIDIS" authorityName="KVACEK et TEODORIDIS" authorityYear="2012" baseAuthorityName="E. KNOBLOCH et KVACEK" baseAuthorityYear="2012" class="Magnoliopsida" family="Fabaceae" genus="Leguminosites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="hradekensis" status="comb. nov.">
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<emphasis id="B97EC51AFFB11924FF09FB9C3EB2FBE6" bold="true" box="[145,445,1110,1133]" italics="true" pageId="18" pageNumber="71">Leguminosites hradekensis</emphasis>
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(E.
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et
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)
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et
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<smallCapsWord id="8D538FD4FFB11924FE1DFBBC3D07FB06" baselines="1159,1160" box="[389,520,1142,1165]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Teodoridis" pageId="18" pageNumber="71">TEODORIDIS</smallCapsWord>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24D7861FFB11924FD8FFBBC3D77FB06" box="[535,632,1142,1165]" pageId="18" pageNumber="71" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB51908FFB11924FEBCFB6D3D54FB36" blockId="18.[292,603,1191,1213]" box="[292,603,1191,1213]" pageId="18" pageNumber="71">Pl. 2, figs 14-19, pl. 12, figs 4-6</paragraph>
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</subSubSection>
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<subSubSection id="C3104A83FFB11925FF1AFB1E3A63FD1A" lastPageId="19" lastPageNumber="72" pageId="18" pageNumber="71" type="description">
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<paragraph id="8BB51908FFB11924FF1AFB1E3FBDFB62" blockId="18.[130,178,1236,1257]" box="[130,178,1236,1257]" pageId="18" pageNumber="71">1976</paragraph>
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<paragraph id="8BB51908FFB11924FF5EFB1E3D38FAAF" blockId="18.[198,766,1236,1316]" pageId="18" pageNumber="71">
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<taxonomicName id="4C0A628BFFB11924FF5EFB1E3D97FB63" authority="E. KNOBLOCH et KVACEK" authorityName="E. KNOBLOCH et KVACEK" box="[198,664,1236,1257]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="hradekensis">
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<emphasis id="B97EC51AFFB11924FF5EFB1E3E91FB62" box="[198,414,1236,1257]" italics="true" pageId="18" pageNumber="71">Magnolia hradekensis</emphasis>
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E.
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et
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<smallCapsWord id="8D538FD4FFB11924FDD3FB1E3D97FB63" baselines="1252,1252" box="[587,664,1236,1257]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kvacek" pageId="18" pageNumber="71">KVAČEK</smallCapsWord>
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</taxonomicName>
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, p. 18, pl. 4, figs 15-19, pl. 5, figs 1-7, pl. 9, figs 1, 9, pl. 15, fig. 14, text-fig. 5 (basionym) (Wackersdorf).
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</paragraph>
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<paragraph id="8BB51908FFB11924FF31FA873B03FE9A" blockId="18.[130,766,1357,2096]" lastBlockId="18.[841,1476,153,851]" pageId="18" pageNumber="71">
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Detached fragmentary leaflets, lamina probably ovate to elliptic,
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<quantity id="4CF2B4EDFFB11924FF43FAA43E4DFA0E" box="[219,322,1390,1413]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="2.0" metricValueMin="0.8" pageId="18" pageNumber="71" unit="mm" value="14.0" valueMax="20.0" valueMin="8.0">8–20 mm</quantity>
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long and
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<quantity id="4CF2B4EDFFB11924FE2AFAA43D17FA0E" box="[434,536,1390,1413]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="18" pageNumber="71" unit="mm" value="9.0" valueMax="13.0" valueMin="5.0">5–13 mm</quantity>
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wide, base and apex not preserved, margin entire, venation brochidodromous, midrib strong, moderate, straight, secondary veins numerous, thinner, straight to slightly curved, looping, alternate to opposite, originating at almost acute angles, intersecondaries obvious, thinner, parallel, tertiary veins alternate percurrent, curved to sinuous, venation of higher orders regular polygonal reticulate, areolation well developed, areoles 3- or 4-sided, veinlets one branched to dichotomous branching. Adaxial epidermis medium cutinized, smooth, non-modified cells polygonal to lobate, 20–25 or up to 35 µm in diameter, anticlinal walls almost straight to strongly wavy, cells narrow elongate over veins, with cylindrical basal cells of serial trichomes 15 µm in diameter, abaxial epidermis delicately cutinized reflecting polygonal to lobate non-modified cells with curved to strongly wavy anticlinal walls, stomata amphicycloparacytic, subsidiary cells sometimes repeatedly subdivided radially, outer circle of subsidiary cells composed of larger cells not much different from the non-modified cells, pairs of slender guard cells 13–16 µm wide and 23–26 µm long, mother cells of undeveloped stomata often present. Cylindrical bases of uniserial trichomes 15–20 µm in diameter scattered among stomata, more densely in some leaf forms, complete trichomes up to 180 µm long rarely preserved with thin-walled apical cell.
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</paragraph>
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<paragraph id="8BB51908FFB11924FCF7FEE93B25FC82" blockId="18.[841,1476,153,851]" pageId="18" pageNumber="71">
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D i s c u s s i o n: The record from the Wackersdorf locality is much more completely documented than the first specimens recovered in the Kristina Mine (Kvaček 1966). Therefore the material from Wackersdorf was preferably published (
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<bibRefCitation id="EF9B64F9FFB11924FC50FE6D3A1EFE35" author="Knobloch, E. & Kvacek, Z." box="[968,1297,423,446]" pageId="18" pageNumber="71" pagination="1 - 131" refId="ref48731" refString="Knobloch, E., Kvacek, Z. (1976): Miozane Blatterfloren vom Westrand der Bohmischen Masse. - Rozpr. Ustr. Ust. Geol., 42: 1 - 131." type="journal article" year="1976">Knobloch and Kvaček 1976</bibRefCitation>
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) and the most complete specimens chosen as the
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<typeStatus id="54B1A7AAFFB11924FB7BFE023A1EFE54" box="[1251,1297,456,479]" pageId="18" pageNumber="71">type</typeStatus>
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of this foliage species, although in the manuscript (Kvaček 1966) the Kristina Mine was intended to be the
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<typeStatus id="54B1A7AAFFB11924FA98FDC03A21FDAA" box="[1280,1326,522,545]" pageId="18" pageNumber="71">type</typeStatus>
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locality. The affinity to the
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<taxonomicName id="4C0A628BFFB11924FC7AFDE63B77FDC8" authorityName="de Jussieu" authorityYear="1789" box="[994,1144,556,579]" class="Magnoliopsida" family="Magnoliaceae" kingdom="Plantae" order="Magnoliales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="family">Magnoliaceae</taxonomicName>
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is now rejected because of the lack of mesophyllous oil cells and the peculiar stomatal
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<typeStatus id="54B1A7AAFFB11924FA0FFD873ACBFDEF" box="[1431,1476,589,612]" pageId="18" pageNumber="71">type</typeStatus>
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(see
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<bibRefCitation id="EF9B64F9FFB11924FCE7FDA43B59FD0E" author="Kvacek, Z. & Teodoridis, V. & Roiron" box="[895,1110,622,645]" pageId="18" pageNumber="71" pagination="1 - 3" refId="ref50486" refString="Kvacek, Z., Teodoridis, V., Roiron (2011): A forgotten Miocene mastixioid flora of Arjuzanx (Landes, SW France). Including In memoriam of Jean Huard (1938 - 1973). - Palaeontographica, Abt. B, 285 (1 - 3): 1 - 111." type="journal article" year="2011">Kvaček et al. 2011</bibRefCitation>
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). More similar in respect of leaf morphology (short petiolule) as well as anatomy are certainly legume leaflets (for comparison see legume foliage described from the Eocene of North America by
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<bibRefCitation id="EF9B64F9FFB11924FA93FD1B3AB0FD63" author="Herendeen, P. S." box="[1291,1471,721,744]" pageId="18" pageNumber="71" pagination="85 - 167" refId="ref47367" refString="Herendeen, P. S. (1992): The fossil history of the Leguminosae from the Eocene of southeastern North America. - In: Herendeen, P. S., Dilcher, D. L. (ed.): Advances in legume systematics. Part 4. The fossil record. - The Royal Botanic Gardens, Kew, pp. 85 - 167." type="book chapter" year="1992">Herendeen 1992</bibRefCitation>
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, p. 137, figs 270-276).
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</paragraph>
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<paragraph id="8BB51908FFB11924FCF7FCD13BCBFCD8" blockId="18.[841,1476,153,851]" pageId="18" pageNumber="71">M a t e r i a l: Leaflet compressions isolated on slides, G 9172-9175 (KR 21, 43a, b, 410).</paragraph>
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<paragraph id="8BB51908FFB11924FB84FC453BFFFC2C" blockId="18.[1052,1264,911,937]" box="[1052,1264,911,937]" pageId="18" pageNumber="71">
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<taxonomicName id="4C0A628BFFB11924FB84FC453BFFFC2C" authority="GRAY" authorityName="GRAY" box="[1052,1264,911,937]" class="Magnoliopsida" family="Betulaceae" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="family">
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<emphasis id="B97EC51AFFB11924FB84FC453BFFFC2C" bold="true" box="[1052,1264,911,937]" pageId="18" pageNumber="71">
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Betulaceae
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||
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<emphasis id="B97EC51AFFB11924FB34FC453BFFFC2C" bold="true" box="[1196,1264,911,937]" pageId="18" pageNumber="71">GRAY</emphasis>
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</smallCapsWord>
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||
</emphasis>
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||
</taxonomicName>
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</paragraph>
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<paragraph id="8BB51908FFB11924FCF3FC0B3A2DFC76" blockId="18.[875,1441,961,1022]" pageId="18" pageNumber="71">
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<taxonomicName id="4C0A628BFFB11924FCF3FC0B3B09FC75" authority="(STERNBERG) KVACEK et HOLY Pl." authorityName="KVACEK et HOLY Pl." baseAuthorityName="STERNBERG" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="julianiformis">
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||
<emphasis id="B97EC51AFFB11924FCF3FC0B3AAEFC5C" bold="true" box="[875,1441,961,984]" pageId="18" pageNumber="71">
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||
<emphasis id="B97EC51AFFB11924FCF3FC0B3B4BFC53" bold="true" box="[875,1092,961,984]" italics="true" pageId="18" pageNumber="71">Alnus julianiformis</emphasis>
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(
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)
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||
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et
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<smallCapsWord id="8D538FD4FFB11924FAF9FC0B3AAEFC5C" baselines="978,978" box="[1377,1441,961,984]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Holy" pageId="18" pageNumber="71">HOLÝ</smallCapsWord>
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</emphasis>
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Pl.
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</taxonomicName>
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2, figs 20-21, pl. 12, figs 7-8
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</paragraph>
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<paragraph id="8BB51908FFB11924FCD1FBDC3C76FBA0" blockId="18.[841,889,1046,1067]" box="[841,889,1046,1067]" pageId="18" pageNumber="71">1823</paragraph>
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<paragraph id="8BB51908FFB11924FCD1FB993C76FB0D" blockId="18.[841,889,1107,1158]" box="[841,889,1107,1158]" pageId="18" pageNumber="71">1845 1974</paragraph>
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<paragraph id="8BB51908FFB11924FC0BFBDC3CD1FBC1" blockId="18.[909,1476,1046,1189]" pageId="18" pageNumber="71">
|
||
<taxonomicName id="4C0A628BFFB11924FC0BFBDC3BD2FBA1" authority="STERNBERG" authorityName="STERNBERG" box="[915,1245,1046,1067]" class="Magnoliopsida" family="Oleaceae" genus="Phyllites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="julianiformis">
|
||
<emphasis id="B97EC51AFFB11924FC0BFBDC3B65FBA0" box="[915,1130,1046,1067]" italics="true" pageId="18" pageNumber="71">Phyllites julianiformis</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB11924FBE9FBDC3BD2FBA1" baselines="1062,1062" box="[1137,1245,1046,1067]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Sternberg" pageId="18" pageNumber="71">STERNBERG</smallCapsWord>
|
||
</taxonomicName>
|
||
, p. 37, 39, pl. 36, fig. 2 (Bílina).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB11924FC15FB993A62FB2E" blockId="18.[909,1476,1046,1189]" pageId="18" pageNumber="71">
|
||
<taxonomicName id="4C0A628BFFB11924FC15FB993B68FBEC" authority="UNGER" authorityName="UNGER" box="[909,1127,1107,1128]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="feroniae">
|
||
<emphasis id="B97EC51AFFB11924FC15FB993B11FBE3" box="[909,1054,1107,1128]" italics="true" pageId="18" pageNumber="71">Fagus feroniae</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB11924FBBDFB993B68FBEC" baselines="1123,1123" box="[1061,1127,1107,1128]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Unger" pageId="18" pageNumber="71">UNGER</smallCapsWord>
|
||
</taxonomicName>
|
||
, p. 106, pl. 28, figs 3-4 (Bílina).
|
||
<taxonomicName id="4C0A628BFFB11924FC15FBBB3A7BFB0E" authority="(STERNBERG) KVACEK et HOLY" authorityName="KVACEK et HOLY" baseAuthorityName="STERNBERG" box="[909,1396,1137,1158]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="julianiformis">
|
||
<emphasis id="B97EC51AFFB11924FC15FBBB3B49FB0D" box="[909,1094,1137,1158]" italics="true" pageId="18" pageNumber="71">Alnus julianiformis</emphasis>
|
||
(
|
||
<smallCapsWord id="8D538FD4FFB11924FBCDFBBB3BCDFB0E" baselines="1153,1153" box="[1109,1218,1137,1158]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Sternberg" pageId="18" pageNumber="71">STERNBERG</smallCapsWord>
|
||
)
|
||
<smallCapsWord id="8D538FD4FFB11924FB49FBBB3A11FB0E" baselines="1153,1153" box="[1233,1310,1137,1158]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kvacek" pageId="18" pageNumber="71">KVAČEK</smallCapsWord>
|
||
et
|
||
<smallCapsWord id="8D538FD4FFB11924FAA5FBBB3A7BFB0E" baselines="1153,1153" box="[1341,1396,1137,1158]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Holy" pageId="18" pageNumber="71">HOLÝ</smallCapsWord>
|
||
</taxonomicName>
|
||
, p. 368, pls 1-3, pl. 4, fig. 1, text-fig. 1 (Bílina, Břeštany).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB11924FCF7FB1A3B34F973" blockId="18.[841,1477,1232,2097]" pageId="18" pageNumber="71">
|
||
Incomplete leaves and fragments, lamina widely elliptic to obovate,
|
||
<quantity id="4CF2B4EDFFB11924FC53FB3B3B3DFA83" box="[971,1074,1265,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="3.2" metricValueMin="0.6" pageId="18" pageNumber="71" unit="mm" value="19.0" valueMax="32.0" valueMin="6.0">6–32 mm</quantity>
|
||
long and
|
||
<quantity id="4CF2B4EDFFB11924FB3CFB3B3A16FA83" box="[1188,1305,1265,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="18" pageNumber="71" unit="mm" value="18.5" valueMax="25.0" valueMin="12.0">12–25 mm</quantity>
|
||
wide, apex not preserved, probably obtuse or acute, base cuneate with fragmentary petiole, margin irregularly simple serrate, teeth acute, sinus angular, venation craspedodromous/semicraspedodromous, midrib straight, strong, secondaries alternate, thinner, regularly spaced, originating at an angle of 30–40°, curved, looping, tertiary veins alternate percurrent, straight or curved, venation of higher orders regular polygonal reticulate, areolation well developed, 4-sided. Adaxial epidermis medium cutinized, smooth, showing outlines of non-modified cells 12–20 µm in diameter with almost straight anticlines, abaxial epidermis thinly cutinized, showing non-modified cells with shallowly wavy anticlines, stomata anomocytic, elliptic, guard cells ca. 25 µm long and 20 µm wide, scattered four-celled trichome bases ca. 28 µm in diameter.
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB11924FCF7F8C03C9EF86C" blockId="18.[841,1477,1232,2097]" pageId="18" pageNumber="71">
|
||
D i s c u s s i o n: In gross morphology and epidermal anatomy the leaf remains described above correspond to
|
||
<taxonomicName id="4C0A628BFFB11924FCD1F8863CF9F8E8" box="[841,1014,1868,1891]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="julianiformis">
|
||
<emphasis id="B97EC51AFFB11924FCD1F8863CF9F8E8" box="[841,1014,1868,1891]" italics="true" pageId="18" pageNumber="71">A. julianiformis</emphasis>
|
||
</taxonomicName>
|
||
from the Bílina area (
|
||
<bibRefCitation id="EF9B64F9FFB11924FB64F8863C8FF80F" author="Kvacek, Z. & Holy, F." pageId="18" pageNumber="71" pagination="367 - 372" refId="ref50205" refString="Kvacek, Z., Holy, F. (1974): Alnus julianaeformis (Sternberg 1823) comb. n., a noteworthy Neogene alder. - Cas. Mineral. Geol., 19: 367 - 372." type="journal article" year="1974">Kvaček and Holý 1974</bibRefCitation>
|
||
) and Wackersdorf (
|
||
<bibRefCitation id="EF9B64F9FFB11924FBF1F8A73AB8F80F" author="Knobloch, E. & Kvacek, Z." box="[1129,1463,1901,1924]" pageId="18" pageNumber="71" pagination="1 - 131" refId="ref48731" refString="Knobloch, E., Kvacek, Z. (1976): Miozane Blatterfloren vom Westrand der Bohmischen Masse. - Rozpr. Ustr. Ust. Geol., 42: 1 - 131." type="journal article" year="1976">Knobloch and Kvaček 1976</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C0A628BFFB11924FCD1F8443B44F82E" authority="HAND." authorityName="HAND." box="[841,1099,1934,1957]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="71" phylum="Tracheophyta" rank="species" species="trabeculosa">
|
||
<emphasis id="B97EC51AFFB11924FCD1F8443CF9F82E" box="[841,1014,1934,1957]" italics="true" pageId="18" pageNumber="71">A. trabeculosa</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB11924FB9DF8443B49F82E" baselines="1951,1952" box="[1029,1094,1934,1957]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Hand" pageId="18" pageNumber="71">HAND</smallCapsWord>
|
||
.
|
||
</taxonomicName>
|
||
-
|
||
<smallCapsWord id="8D538FD4FFB11924FBCBF8443B97F82E" baselines="1951,1952" box="[1107,1176,1934,1957]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Mazz" pageId="18" pageNumber="71">MAZZ</smallCapsWord>
|
||
. of
|
||
<collectingCountry id="F31D5998FFB11924FB4FF8443A45F82E" box="[1239,1354,1934,1957]" name="China" pageId="18" pageNumber="71">East Asia</collectingCountry>
|
||
has been suggested as the living relative (
|
||
<bibRefCitation id="EF9B64F9FFB11924FB71F8653C8AF86C" author="Kvacek, Z. & Holy, F." pageId="18" pageNumber="71" pagination="367 - 372" refId="ref50205" refString="Kvacek, Z., Holy, F. (1974): Alnus julianaeformis (Sternberg 1823) comb. n., a noteworthy Neogene alder. - Cas. Mineral. Geol., 19: 367 - 372." type="journal article" year="1974">Kvaček and Holý 1974</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB11924FCF7F8333A9BF7BA" blockId="18.[841,1477,1232,2097]" pageId="18" pageNumber="71">M a t e r i a l: Leaf compressions on slides, G 9176- 9183 (KR 143, 144, 147, 151, 194a, b, 224, 225, 248).</paragraph>
|
||
<paragraph id="8BB51908FFB01925FEA3FF523D2AFF39" blockId="19.[315,549,152,178]" box="[315,549,152,178]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FEA3FF523D2AFF39" authority="DUMORT." authorityName="DUMORT." box="[315,549,152,178]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="family">
|
||
<emphasis id="B97EC51AFFB01925FEA3FF523D2AFF39" bold="true" box="[315,549,152,178]" pageId="19" pageNumber="72">
|
||
Fagaceae
|
||
<smallCapsWord id="8D538FD4FFB01925FE2FFF523D11FF3B" baselines="171,171" box="[439,542,152,178]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dumort" pageId="19" pageNumber="72">DUMORT</smallCapsWord>
|
||
.
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FEE1FF023D66FE82" blockId="19.[249,617,200,265]" pageId="19" pageNumber="72">
|
||
<emphasis id="B97EC51AFFB01925FEE1FF023D66FE82" bold="true" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FEE1FF023EC9FF69" box="[377,454,200,226]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B97EC51AFFB01925FEE1FF023EC9FF69" bold="true" box="[377,454,200,226]" italics="true" pageId="19" pageNumber="72">Fagus</emphasis>
|
||
</taxonomicName>
|
||
L.
|
||
<taxonomicName id="4C0A628BFFB01925FF61FF383D17FE82" authority="UNGER" authorityName="UNGER" box="[249,536,242,265]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="deucalionis">
|
||
<emphasis id="B97EC51AFFB01925FF61FF383ECBFE82" bold="true" box="[249,452,242,265]" italics="true" pageId="19" pageNumber="72">Fagus deucalionis</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB01925FE53FF383D17FE82" baselines="259,260" box="[459,536,242,265]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Unger" pageId="19" pageNumber="72">UNGER</smallCapsWord>
|
||
</taxonomicName>
|
||
plexus
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FFE1FEE93ED3FEEB" blockId="19.[121,744,291,353]" pageId="19" pageNumber="72">Pl. 2, figs 22-27, pl. 3, figs 6-7, pl. 6, fig. 20, pl. 7, fig. 1, pl. 12, figs 9-10</paragraph>
|
||
<paragraph id="8BB51908FFB01925FFEBFEB23FACFE06" blockId="19.[115,163,376,397]" box="[115,163,376,397]" pageId="19" pageNumber="72">1847</paragraph>
|
||
<paragraph id="8BB51908FFB01925FFEBFE793FACFE43" blockId="19.[115,163,435,456]" box="[115,163,435,456]" pageId="19" pageNumber="72">1906</paragraph>
|
||
<paragraph id="8BB51908FFB01925FFEBFE273FACFD89" blockId="19.[115,163,493,514]" box="[115,163,493,514]" pageId="19" pageNumber="72">1933</paragraph>
|
||
<paragraph id="8BB51908FFB01925FFEBFDE23FACFDB6" blockId="19.[115,163,552,573]" box="[115,163,552,573]" pageId="19" pageNumber="72">1989</paragraph>
|
||
<paragraph id="8BB51908FFB01925FFEBFDA93FACFDF3" blockId="19.[115,163,611,632]" box="[115,163,611,632]" pageId="19" pageNumber="72">1989</paragraph>
|
||
<paragraph id="8BB51908FFB01925FFEBFD573FACFD39" blockId="19.[115,163,669,690]" box="[115,163,669,690]" pageId="19" pageNumber="72">1991</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF2FFEB23E81FE21" blockId="19.[183,751,376,720]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FF2FFEB23EA0FE07" authority="UNGER" authorityName="UNGER" box="[183,431,376,397]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="deucalionis">
|
||
<emphasis id="B97EC51AFFB01925FF2FFEB23E67FE06" box="[183,360,376,397]" italics="true" pageId="19" pageNumber="72">Fagus deucalionis</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB01925FEF5FEB23EA0FE07" baselines="392,392" box="[365,431,376,397]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Unger" pageId="19" pageNumber="72">UNGER</smallCapsWord>
|
||
</taxonomicName>
|
||
, p. 101, pl. 27, figs 1-6 (Počerny, in German Putschirn).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF2FFE793ED5FE6E" blockId="19.[183,751,376,720]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FF2FFE793DADFE4C" authority="MENZEL" authorityName="MENZEL" box="[183,674,435,456]" class="Magnoliopsida" family="Fagaceae" form="miocenica" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="form" species="ferruginea">
|
||
<emphasis id="B97EC51AFFB01925FF2FFE793E6CFE43" box="[183,355,435,456]" italics="true" pageId="19" pageNumber="72">Fagus ferruginea</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB01925FEF6FE793E9EFE4C" baselines="451,451" box="[366,401,435,456]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Ait" pageId="19" pageNumber="72">AIT</smallCapsWord>
|
||
. forma
|
||
<emphasis id="B97EC51AFFB01925FE7EFE793D46FE43" box="[486,585,435,456]" italics="true" pageId="19" pageNumber="72">miocenica</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB01925FDCCFE793DADFE4C" baselines="451,451" box="[596,674,435,456]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Menzel" pageId="19" pageNumber="72">MENZEL</smallCapsWord>
|
||
</taxonomicName>
|
||
, p. 48, pl. 3, figs 4-5, 10-12 (Rauno).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF2FFE273ED9FDAB" blockId="19.[183,751,376,720]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FF2FFE273EBDFD89" authority="AIT." authorityName="AIT." box="[183,434,493,514]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" isUncertain="true" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="ferruginea">
|
||
<emphasis id="B97EC51AFFB01925FF2FFE273FFCFD89" box="[183,243,493,514]" italics="true" pageId="19" pageNumber="72">Fagus</emphasis>
|
||
cf.
|
||
<emphasis id="B97EC51AFFB01925FE84FE273E8EFD89" box="[284,385,493,514]" italics="true" pageId="19" pageNumber="72">ferruginea</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB01925FE12FE273EA3FD8A" baselines="509,509" box="[394,428,493,514]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Ait" pageId="19" pageNumber="72">AIT</smallCapsWord>
|
||
.
|
||
</taxonomicName>
|
||
; Menzel in Gothan and Sapper, p. 15, pl. 3, fig. 8 (Klettwitz).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF2FFDE23D3BFDD1" blockId="19.[183,751,376,720]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FF2FFDE23EB8FDB7" authority="GOPPERT" authorityName="GOPPERT (Kvacek and Walther" authorityYear="1989" box="[183,439,552,573]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="attenuata">
|
||
<emphasis id="B97EC51AFFB01925FF2FFDE23E56FDB6" box="[183,345,552,573]" italics="true" pageId="19" pageNumber="72">Fagus attenuata</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB01925FEFCFDE23EB8FDB7" baselines="568,568" box="[356,439,552,573]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Goppert" pageId="19" pageNumber="72">GÖPPERT</smallCapsWord>
|
||
</taxonomicName>
|
||
; Kvaček and Walther, p. 214, text-fig. 1f (Hrádek/N., Kristina Mine).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF2FFDA93D9BFD1E" blockId="19.[183,751,376,720]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FF2FFDA93E52FDF3" authorityName="R.Brown ex Turner" authorityYear="1811" box="[183,349,611,632]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="decurrens">
|
||
<emphasis id="B97EC51AFFB01925FF2FFDA93E52FDF3" box="[183,349,611,632]" italics="true" pageId="19" pageNumber="72">Fagus decurrens</emphasis>
|
||
</taxonomicName>
|
||
C. et E.M.
|
||
<smallCapsWord id="8D538FD4FFB01925FE46FDA93D03FDFC" baselines="627,627" box="[478,524,611,632]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Reid" pageId="19" pageNumber="72">REID</smallCapsWord>
|
||
; Kvaček and Walther, p. 214, text-fig. 1b-c (Hrádek/N., Kristina Mine).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF2FFD573EA5FD5B" blockId="19.[183,751,376,720]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FF2FFD573D3DFD3A" authority="KVACEK et WALTHER" authorityName="KVACEK et WALTHER" box="[183,562,669,690]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="menzelii">
|
||
<emphasis id="B97EC51AFFB01925FF2FFD573E5EFD39" box="[183,337,669,690]" italics="true" pageId="19" pageNumber="72">Fagus menzelii</emphasis>
|
||
<smallCapsWord id="8D538FD4FFB01925FEC7FD573EA3FD3A" baselines="685,685" box="[351,428,669,690]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kvacek" pageId="19" pageNumber="72">KVAČEK</smallCapsWord>
|
||
et
|
||
<smallCapsWord id="8D538FD4FFB01925FE41FD573D3DFD3A" baselines="685,685" box="[473,562,669,690]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Walther" pageId="19" pageNumber="72">WALTHER</smallCapsWord>
|
||
</taxonomicName>
|
||
, p. 485, pls 7-8, text-figs 9-10 (Kausche).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF01FD333DC9FA41" blockId="19.[115,751,761,2097]" pageId="19" pageNumber="72">
|
||
Incomplete leaves and fragments, lamina elliptic to widely elliptic to ovate,
|
||
<quantity id="4CF2B4EDFFB01925FEECFCD23EE9FCA4" box="[372,486,792,815]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.8" metricValueMin="1.2" pageId="19" pageNumber="72" unit="mm" value="20.0" valueMax="28.0" valueMin="12.0">12–28 mm</quantity>
|
||
long and
|
||
<quantity id="4CF2B4EDFFB01925FDD7FCD23DA1FCA4" box="[591,686,792,815]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.5" metricValueMin="0.8" pageId="19" pageNumber="72" unit="mm" value="16.5" valueMax="25.0" valueMin="8.0">8–25mm</quantity>
|
||
wide, apex not preserved, probably acute to shortly attenuate, base cuneate with petiole up to
|
||
<quantity id="4CF2B4EDFFB01925FE4BFC923D1CFCE4" box="[467,531,856,879]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="72" unit="mm" value="8.0">8 mm</quantity>
|
||
long, margin partly entire, at places regularly widely simple serrate, teeth acute, rarely blunt, sinus angular, venation simply pseudocraspedodromous, midrib straight, strong, secondaries alternate to opposite, thinner, regularly spaced, straight, originating at an angle of 30 to 40°, curved just before entering the marginal tooth, tertiary veins alternate percurrent, straight or curved, often forked, venation of higher orders regular polygonal reticulate, areolation well developed, 4-sided. Adaxial and abaxial cuticles hardly discernible on macerated samples, more visible directly on the sub-macerated leaf compressions. Cupules broadly ovate to broadly acute,
|
||
<quantity id="4CF2B4EDFFB01925FF25FB1F3E28FB67" box="[189,295,1237,1260]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="2.1" metricValueMin="0.8" pageId="19" pageNumber="72" unit="mm" value="14.5" valueMax="21.0" valueMin="8.0">8–21 mm</quantity>
|
||
long, 6–13.3 mm wide, on apices often broken, (probable original length
|
||
<quantity id="4CF2B4EDFFB01925FE7CFB3E3D40FA80" box="[484,591,1268,1291]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="2.1" metricValueMin="0.8" pageId="19" pageNumber="72" unit="mm" value="14.5" valueMax="21.0" valueMin="8.0">8–21 mm</quantity>
|
||
), valves ovoid to broadly lanceolate, on base rounded, concentrically striate, on surface dense decurrent bases of spines present, stalk conical, short. Fruits elongate ovoid,
|
||
<quantity id="4CF2B4EDFFB01925FDDEFA9E3DA1FAE0" box="[582,686,1364,1387]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="19" pageNumber="72" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
|
||
long, 4.5–
|
||
<quantity id="4CF2B4EDFFB01925FF3BFAB93FD0FA01" box="[163,223,1395,1418]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="72" unit="mm" value="5.0">5 mm</quantity>
|
||
wide, triangular, apically narrowing into pointed apex, edges sharp to winged, bluntly rounded on base (
|
||
<bibRefCitation id="EF9B64F9FFB01925FD2FFA593FA4FA41" author="Holy, F." pageId="19" pageNumber="72" refId="ref47510" refString="Holy, F. (1975): Neogenni mastixioidni kvetena svrchniho slojoveho pasma z lomu " Kristina " (Hradek nad Nisou). (Neogene mastixioid flora of the upper seam zone from the open mine " Kristina " (Hradek upon Nisa) - MS unpublished report (in Czech); Ph. D. Thesis, Charles University, Prague." type="book" year="1975">Holý 1975</bibRefCitation>
|
||
, pp. 32-34, pl. 5, figs 15-17, as
|
||
<taxonomicName id="4C0A628BFFB01925FD9AFA793DB9FA41" authorityName="R.Brown ex Turner" authorityYear="1811" box="[514,694,1459,1482]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="decurrens">
|
||
<emphasis id="B97EC51AFFB01925FD9AFA793DB9FA41" box="[514,694,1459,1482]" italics="true" pageId="19" pageNumber="72">Fagus decurrens</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FF01FA103BA8FE1B" blockId="19.[115,751,761,2097]" lastBlockId="19.[826,1461,153,567]" pageId="19" pageNumber="72">
|
||
D i s c u s s i o n:
|
||
<taxonomicName id="4C0A628BFFB01925FEF7FA103EBDFA7A" box="[367,434,1498,1521]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B97EC51AFFB01925FEF7FA103EBDFA7A" box="[367,434,1498,1521]" italics="true" pageId="19" pageNumber="72">Fagus</emphasis>
|
||
</taxonomicName>
|
||
leaf fossils of the European Early-Middle Miocene were usually assigned to
|
||
<taxonomicName id="4C0A628BFFB01925FDE9FA303D00F9BA" authority="GOPPERT (Kvacek and Walther 1989)" authorityName="GOPPERT (Kvacek and Walther" authorityYear="1989" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="attenuata">
|
||
<emphasis id="B97EC51AFFB01925FDE9FA303DE1F99A" box="[625,750,1530,1553]" italics="true" pageId="19" pageNumber="72">F. attenuata</emphasis>
|
||
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|
||
(
|
||
<bibRefCitation id="EF9B64F9FFB01925FF7AF9D03D08F9BA" author="Kvacek, Z. & Walther, H." box="[226,519,1562,1585]" pageId="19" pageNumber="72" pagination="213 - 229" refId="ref50725" refString="Kvacek, Z., Walther, H. (1989): Paleobotanical studies in Fagaceae of the European Tertiary. - Pl. Syst. Evol., 162: 213 - 229." type="journal article" year="1989">Kvaček and Walther 1989</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but the revision by
|
||
<bibRefCitation id="EF9B64F9FFB01925FFEBF9F03ECBF9DA" author="Walther, H. & Zastawniak, E." box="[115,452,1594,1617]" pageId="19" pageNumber="72" pagination="153 - 199" refId="ref54297" refString="Walther, H., Zastawniak, E. (1991): Fagaceae from Sosnica and Malczyce (near Wroclaw, Poland). A revision of original materials by Goeppert 1852 and 1855 and a study of new collections. - Acta Palaeobot., 31: 153 - 199." type="journal article" year="1991">Walther and Zastawniak (1991)</bibRefCitation>
|
||
corrected the affinity of the
|
||
<typeStatus id="54B1A7AAFFB01925FFEBF9903FAEF9FA" box="[115,161,1626,1649]" pageId="19" pageNumber="72">type</typeStatus>
|
||
specimen to
|
||
<taxonomicName id="4C0A628BFFB01925FEB5F9903E65F9FA" box="[301,362,1626,1649]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B97EC51AFFB01925FEB5F9903E65F9FA" box="[301,362,1626,1649]" italics="true" pageId="19" pageNumber="72">Alnus</emphasis>
|
||
</taxonomicName>
|
||
(see Walther in
|
||
<bibRefCitation id="EF9B64F9FFB01925FD8EF9903FA4F91A" author="Kvacek, Z. & Walther, H." pageId="19" pageNumber="72" pagination="471 - 534" refId="ref50761" refString="Kvacek, Z., Walther, H. (1991): Revision der mitteleuropaischen tertiaren Fagaceen nach blattepidermalen Charakteristiken. IV. Teil. Fagus Linne. - Feddes Repert., 102: 471 - 534." type="journal article" year="1991">Kvaček and Walther 1991</bibRefCitation>
|
||
, p. 488). Later the leaf fossils of this
|
||
<taxonomicName id="4C0A628BFFB01925FDDDF9B03DBCF91A" box="[581,691,1658,1681]" form="have" pageId="19" pageNumber="72" rank="form">form have</taxonomicName>
|
||
been assigned either to
|
||
<taxonomicName id="4C0A628BFFB01925FEADF9503FB5F95A" authority="WALTHER et ZASTAWNIAK (1991)" authorityName="WALTHER et ZASTAWNIAK" authorityYear="1991" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="silesiaca">
|
||
<emphasis id="B97EC51AFFB01925FEADF9503ED3F93A" box="[309,476,1690,1713]" italics="true" pageId="19" pageNumber="72">Fagus silesiaca</emphasis>
|
||
<bibRefCitation id="EF9B64F9FFB01925FE7BF9503FB5F95A" author="Walther, H. & Zastawniak, E." pageId="19" pageNumber="72" pagination="153 - 199" refId="ref54297" refString="Walther, H., Zastawniak, E. (1991): Fagaceae from Sosnica and Malczyce (near Wroclaw, Poland). A revision of original materials by Goeppert 1852 and 1855 and a study of new collections. - Acta Palaeobot., 31: 153 - 199." type="journal article" year="1991">
|
||
<smallCapsWord id="8D538FD4FFB01925FE7BF9503D48F93A" baselines="1707,1708" box="[483,583,1690,1713]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Walther" pageId="19" pageNumber="72">WALTHER</smallCapsWord>
|
||
et
|
||
<smallCapsWord id="8D538FD4FFB01925FDF0F9503DE1F93A" baselines="1707,1708" box="[616,750,1690,1713]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Zastawniak" pageId="19" pageNumber="72">ZASTAWNIAK</smallCapsWord>
|
||
(1991)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
based on the Late Miocene material of Sośnica or
|
||
<taxonomicName id="4C0A628BFFB01925FD42F9703D0BF97A" authority="KVACEK et WALTHER (1991)" authorityName="KVACEK et WALTHER" authorityYear="1991" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="menzelii">
|
||
<emphasis id="B97EC51AFFB01925FD42F9703FC3F97A" italics="true" pageId="19" pageNumber="72">F. menzelii</emphasis>
|
||
<bibRefCitation id="EF9B64F9FFB01925FF4DF9103D0BF97A" author="Kvacek, Z. & Walther, H." box="[213,516,1754,1777]" pageId="19" pageNumber="72" pagination="471 - 534" refId="ref50761" refString="Kvacek, Z., Walther, H. (1991): Revision der mitteleuropaischen tertiaren Fagaceen nach blattepidermalen Charakteristiken. IV. Teil. Fagus Linne. - Feddes Repert., 102: 471 - 534." type="journal article" year="1991">
|
||
<smallCapsWord id="8D538FD4FFB01925FF4DF9103E25F97A" baselines="1771,1772" box="[213,298,1754,1777]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Kvacek" pageId="19" pageNumber="72">KVAČEK</smallCapsWord>
|
||
et
|
||
<smallCapsWord id="8D538FD4FFB01925FEC8F9103EBBF97A" baselines="1771,1772" box="[336,436,1754,1777]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Walther" pageId="19" pageNumber="72">WALTHER</smallCapsWord>
|
||
(1991)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
based on the Middle Miocene material from
|
||
<collectingCountry id="F31D5998FFB01925FEF7F9303EDDF89A" box="[367,466,1786,1809]" name="Germany" pageId="19" pageNumber="72">Germany</collectingCountry>
|
||
(Kausche and other sites). Walther (in
|
||
<bibRefCitation id="EF9B64F9FFB01925FF60F8D03D15F8BA" author="Kvacek, Z. & Walther, H." box="[248,538,1818,1841]" pageId="19" pageNumber="72" pagination="471 - 534" refId="ref50761" refString="Kvacek, Z., Walther, H. (1991): Revision der mitteleuropaischen tertiaren Fagaceen nach blattepidermalen Charakteristiken. IV. Teil. Fagus Linne. - Feddes Repert., 102: 471 - 534." type="journal article" year="1991">Kvaček and Walther 1991</bibRefCitation>
|
||
, p. 488) confirmed that differences between the two entities are negligible. Associated cupules are also very similar, usually assigned to a single fossil species (
|
||
<bibRefCitation id="EF9B64F9FFB01925FE15F8B03D0AF81A" author="Holy, F." box="[397,517,1914,1937]" pageId="19" pageNumber="72" refId="ref47510" refString="Holy, F. (1975): Neogenni mastixioidni kvetena svrchniho slojoveho pasma z lomu " Kristina " (Hradek nad Nisou). (Neogene mastixioid flora of the upper seam zone from the open mine " Kristina " (Hradek upon Nisa) - MS unpublished report (in Czech); Ph. D. Thesis, Charles University, Prague." type="book" year="1975">Holý 1975</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B64F9FFB01925FD89F8B03FA7F83A" author="Kvacek, Z. & Walther, H." pageId="19" pageNumber="72" pagination="213 - 229" refId="ref50725" refString="Kvacek, Z., Walther, H. (1989): Paleobotanical studies in Fagaceae of the European Tertiary. - Pl. Syst. Evol., 162: 213 - 229." type="journal article" year="1989">Kvaček and Walther 1989</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C0A628BFFB01925FF52F8503E72F83A" authorityName="R.Brown ex Turner" authorityYear="1811" box="[202,381,1946,1969]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="decurrens">
|
||
<emphasis id="B97EC51AFFB01925FF52F8503E72F83A" box="[202,381,1946,1969]" italics="true" pageId="19" pageNumber="72">Fagus decurrens</emphasis>
|
||
</taxonomicName>
|
||
C. et E. M.
|
||
<smallCapsWord id="8D538FD4FFB01925FE63F8503D21F83A" baselines="1963,1964" box="[507,558,1946,1969]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Reid" pageId="19" pageNumber="72">REID</smallCapsWord>
|
||
,
|
||
<bibRefCitation id="EF9B64F9FFB01925FDA3F8503FA4F85A" author="Denk, T. & Meller, B." pageId="19" pageNumber="72" pagination="869 - 897" refId="ref45825" refString="Denk, T., Meller, B. (2001): The systematic significance of the cupule / nut complex in living and fossil Fagus. - Int. J. Plant Sci., 162 (4): 869 - 897." type="journal article" year="2001">Denk and Meller 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B64F9FFB01925FF20F8703E63F85A" author="Teodoridis, V." box="[184,364,1978,2001]" pageId="19" pageNumber="72" pagination="9 - 49" refId="ref53425" refString="Teodoridis, V. (2003): Early Miocene carpological material from the Czech part of the Zittau Basin - Acta Paleobot., 43 (1): 9 - 49." type="journal article" year="2003">Teodoridis 2003</bibRefCitation>
|
||
). We include the two fossil species based on foliage, which were thriving under quite different environmental conditions into a single species plexus and, contrary to the recent treatment of fossil beach foliage (
|
||
<bibRefCitation id="EF9B64F9FFB01925FCD9FF533CB6FF3B" author="Denk, T." box="[833,953,153,176]" pageId="19" pageNumber="72" pagination="1 - 72" refId="ref45791" refString="Denk, T. (2004): Revision of Fagus from the Tertiary of Europe and southwestern Asia and its phylogenetic implications. - Documenta Naturae, 150: 1 - 72." type="journal article" year="2004">Denk 2004</bibRefCitation>
|
||
), we consider them and other fossil morphotypes of beech in Europe as infraspecific variation. As shown by
|
||
<bibRefCitation id="EF9B64F9FFB01925FCA2FF133B4CFF7B" author="Denk, T. & Meller, B." box="[826,1091,217,240]" pageId="19" pageNumber="72" pagination="869 - 897" refId="ref45825" refString="Denk, T., Meller, B. (2001): The systematic significance of the cupule / nut complex in living and fossil Fagus. - Int. J. Plant Sci., 162 (4): 869 - 897." type="journal article" year="2001">Denk and Meller (2001)</bibRefCitation>
|
||
the morphology of fossil cupules of beech conforms to a single species throughout Europe and therefore we suggest assigning the populations of the Kristina Mine and others connected with mastixioid floras (including the Late Oligocene site Počerny) into an informal species plexus
|
||
<emphasis id="B97EC51AFFB01925FC43FEB33BA8FE1B" box="[987,1191,377,400]" italics="true" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FC43FEB33BACFE1B" box="[987,1187,377,400]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="72" phylum="Tracheophyta" rank="species" species="deucalionis">Fagus deucalionis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB51908FFB01925FCC7FE6A3B20FDBC" blockId="19.[826,1461,153,567]" pageId="19" pageNumber="72">M a t e r i a l: Leaf compressions on slides, G 9184- 9194, partly missing (KR 5, 9, 10, 11, 12, 13, 14, 15, 16, 17, 18, 23, 25, 26, 105, 106, 107, 108, 109, 110, 111, 112, 113, 114, 121, 122, 123a, b), numerous cupules, several fruits, G 8821, 8822, Gs 232.</paragraph>
|
||
<paragraph id="8BB51908FFB01925FC1AFDB33A63FD1A" blockId="19.[898,1388,633,659]" box="[898,1388,633,659]" pageId="19" pageNumber="72">
|
||
<emphasis id="B97EC51AFFB01925FC1AFDB33A63FD1A" bold="true" box="[898,1388,633,659]" pageId="19" pageNumber="72">
|
||
<taxonomicName id="4C0A628BFFB01925FC1AFDB33B6CFD18" box="[898,1123,633,659]" class="Magnoliopsida" family="Fagaceae" genus="Trigonobalanopsis" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B97EC51AFFB01925FC1AFDB33B6CFD18" bold="true" box="[898,1123,633,659]" italics="true" pageId="19" pageNumber="72">Trigonobalanopsis</emphasis>
|
||
</taxonomicName>
|
||
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|
||
et
|
||
<smallCapsWord id="8D538FD4FFB01925FB6EFDB33A63FD1A" baselines="652,652" box="[1270,1388,633,659]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Walther" pageId="19" pageNumber="72">
|
||
<emphasis id="B97EC51AFFB01925FB6EFDB33A63FD1A" bold="true" box="[1270,1388,633,659]" pageId="19" pageNumber="72">WALTHER</emphasis>
|
||
</smallCapsWord>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |