treatments-xml/data/7E/65/B4/7E65B4F0822C58BABD666E44B1A0C99B.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.97.59143" ID-GBIF-Dataset="83d5b180-4e0b-44d1-ad83-67ad55ee0355" ID-GBIF-Taxon="182236446" ID-Pensoft-Pub="1860-0743-1-1" ID-Pensoft-UUID="BD006A3AD2C6516AB596CDA05E0F375D" ID-ZooBank="1D5553BA24894F2C9527B7E46B9B432B" LSID="urn:lsid:zoobank.org:act:8797FE78-84C5-4887-AF61-923C1E341734" ModsDocID="1860-0743-97-1-1" checkinTime="1621423297205" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Pall-Gergely, Barna, Sutcharit, Chirasak &amp; Tongkerd, Piyoros" docDate="2021" docId="7E65B4F0822C58BABD666E44B1A0C99B" docLanguage="en" docName="ZoosystEvol 97(1): 1-20" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-01-05" docSource="http://dx.doi.org/10.3897/zse.97.59143" docTitle="Dicharax burchi Jirapatrasilp &amp; Pall-Gergely 2021, sp. nov." docType="treatment" docUuid="8797FE78-84C5-4887-AF61-923C1E341734" docUuidSource="ZooBank" docVersion="3" id="BD006A3AD2C6516AB596CDA05E0F375D" lastPageNumber="1" masterDocId="BD006A3AD2C6516AB596CDA05E0F375D" masterDocTitle="The operculate micro land snail genus Dicharax Kobelt &amp; Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species" masterLastPageNumber="20" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1632012559486" updateUser="ExternalLinkService">
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<mods:title>The operculate micro land snail genus Dicharax Kobelt &amp; Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tongkerd, Piyoros</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9221-9293</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">piyorose@hotmail.com</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-01-05</mods:number>
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<mods:detail type="volume">
<mods:number>97</mods:number>
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<mods:number>1</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zse.97.59143</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.59143</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-1</mods:identifier>
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<treatment ID-GBIF-Taxon="182236446" LSID="urn:lsid:zoobank.org:act:8797FE78-84C5-4887-AF61-923C1E341734" httpUri="http://treatment.plazi.org/id/7E65B4F0822C58BABD666E44B1A0C99B" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/8797FE78-84C5-4887-AF61-923C1E341734" authority="Jirapatrasilp &amp; Páll-Gergely" authorityName="Jirapatrasilp &amp; Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax burchi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burchi" status="sp. nov.">
Dicharax burchi Jirapatrasilp &amp; 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">Figs 6F</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">, 11A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
CUMZ 7428/1 (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10A</figureCitation>
), 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paratypes</emphasis>
CUMZ 7428/2 (1 shell and 6 specimens in ethanol; Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">6F</figureCitation>
), HNHM (1 shell), NHMUK 20200326 (2 shells) and SMF (2 shells) from Tham Phaya Nakharat, Khon Kaen Province, Thailand, 
<geoCoordinate degrees="16" direction="north" minutes="48" orientation="latitude" precision="1" seconds="32.8" value="16.80911">16°48'32.8&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="57" orientation="longitude" precision="1" seconds="23.9" value="101.956635">101°57'23.9&quot;E</geoCoordinate>
, 21 July 2020, coll. P. Tongkerd, A. Pholyotha.
</paragraph>
<caption doi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
<taxonomicName authorityName="Jirapatrasilp &amp; Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax burchi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burchi">
<emphasis italics="true" pageId="0" pageNumber="1">Dicharax burchi</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A.</emphasis>
Holotype CUMZ 7428/1 with operculum; 
<emphasis bold="true" pageId="0" pageNumber="1">B, C.</emphasis>
Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; 
<emphasis bold="true" pageId="0" pageNumber="1">D.</emphasis>
Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="UF 347279" collectingDate="1987-06-23" collectorName="NW Ban Soppong, F. G. Thompson" country="Thailand" elevation="800" latitude="19.55305" location="Soppong" longLatPrecision="1" longitude="98.192215" specimenCount="1" stateProvince="Mae Hong Son Province">
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/UF347279">UF 347279</accessionNumber>
from limestone pass 10.0 km 
<collectorName>
NW Ban 
<location LSID="urn:lsid:plazi:treatment:7E65B4F0822C58BABD666E44B1A0C99B:755052286A1814E9B41D741532591067" country="Thailand" latitude="19.55305" longLatPrecision="1" longitude="98.192215" name="Soppong" stateProvince="Mae Hong Son Province">Soppong</location>
</collectorName>
, 
<collectingRegion country="Thailand" name="Mae Hong Son">Mae Hong Son Province</collectingRegion>
, 
<collectingCountry name="Thailand">Thailand</collectingCountry>
, 
<geoCoordinate degrees="19" direction="north" minutes="33.183" orientation="latitude" precision="1" value="19.55305">19°33.183'N</geoCoordinate>
, 
<geoCoordinate degrees="98" direction="east" minutes="11.533" orientation="longitude" precision="1" value="98.192215">98°11.533'E</geoCoordinate>
, 
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="1987-06-23">23 Jun. 1987</collectingDate>
, coll. 
<collectorName>F.G. Thompson</collectorName>
(1 shell; Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10D</figureCitation>
)
</materialsCitation>
. 
<materialsCitation accessionNumber="UF 345622" collectingDate="1988-03-20" collectingDateMax="2018-10-07" collectingDateMin="1988-03-20" collectorName="K. Auffenberg, A. Hunyadi, Hunyadi, K, Okubo, J. U, Doi Chiang Dao, S. Panha, Panha, J. B, Burch, P" country="Thailand" elevation="820" latitude="19.24075" location="Road" longLatPrecision="1" longitude="98.8805" specimenCount="1" stateProvince="Mae Hong Son Province">
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/UF345622">UF 345622</accessionNumber>
from 
<locationDeviation location="Soppong">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.02" unit="km" value="10.2">10.2 km</quantity>
WNW of Soppong
</locationDeviation>
, 
<location LSID="urn:lsid:plazi:treatment:7E65B4F0822C58BABD666E44B1A0C99B:724369B2E036B74DC723D2E4F03734B6" country="Thailand" latitude="19.24075" longLatPrecision="1" longitude="98.8805" name="Road" stateProvince="Mae Hong Son Province">Road</location>
1095, 
<collectingRegion country="Thailand" name="Mae Hong Son">Mae Hong Son Province</collectingRegion>
, 
<collectingCountry name="Thailand">Thailand</collectingCountry>
, 
<geoCoordinate degrees="19" direction="north" minutes="33" orientation="latitude" precision="925" value="19.55">19°33'N</geoCoordinate>
, 
<geoCoordinate degrees="98" direction="east" minutes="03" orientation="longitude" precision="925" value="98.05">98°03'E</geoCoordinate>
, 
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.2" unit="m" value="820.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.2" unit="m" value="820.0">820 m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="1988-03-20">20 Mar. 1988</collectingDate>
, coll. 
<collectorName>K. Auffenberg</collectorName>
(3 shells). HA (locality code: 2018/41) from Hsihseng centre E ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
, right side of rd. + 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
</quantity>
on unpaved rd., limestone hill, Shan State, 
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
, 
<geoCoordinate degrees="20" direction="north" minutes="8.002" orientation="latitude" precision="1" value="20.133368">20°8.002'N</geoCoordinate>
, 
<geoCoordinate degrees="97" direction="east" minutes="18.024" orientation="longitude" precision="1" value="97.3004">97°18.024'E</geoCoordinate>
, 
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="2018-10-07">7 Oct. 2018</collectingDate>
, coll. 
<collectorName>A. 
<collectorName>Hunyadi, K</collectorName>
</collectorName>
. 
<collectorName>Okubo, J.U</collectorName>
. Otani (1 shell; Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11A</figureCitation>
). CUMZ 7431 from 
<collectorName>Doi Chiang Dao</collectorName>
, 
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
, 
<collectingCountry name="Thailand">Thailand</collectingCountry>
, 
<geoCoordinate degrees="19" direction="north" minutes="14" orientation="latitude" precision="1" seconds="26.7" value="19.24075">19°14'26.7&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="49.8" value="98.8805">98°52'49.8&quot;E</geoCoordinate>
, 
<collectingDate value="1997-05-24" valueMax="1997-05-28" valueMin="1997-05-24">24-28 May 1997</collectingDate>
, coll. 
<collectorName>S. 
<collectorName>Panha, J.B</collectorName>
</collectorName>
. 
<collectorName>Burch, P</collectorName>
. Dumrongrojwattana (4 shells; Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10B, C</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shell small (SH up to 3.6 mm, SW up to 3.5 mm), conical. Spire ca. 
<normalizedToken originalValue="½">1/2</normalizedToken>
of shell height. R1 with regular, strong ribs; R2 as long as R3; R2 with 36-43 ribs. R3 with either sharp and narrow or blunt and more elongated swelling in the middle of R3 or near aperture. Aperture round. Inner peristome blunt, slightly protruding. Outer peristome expanded (most conspicuously at the upper palatal position and near the umbilicus), reflected. Umbilicus partly or entirely covered by expanded outer peristome. Operculum multispiral.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shell small (SH up to 3.6 mm, SW up to 3.5 mm), conical, solid, semitransparent, yellowish to pale orange. Shell outline round in apical view, spire ca. 
<normalizedToken originalValue="½">1/2</normalizedToken>
of shell height. Whorls approaching 5, last whorl large. Protoconch low, ca. two whorls, glossy and smooth. R1 ca. 2 whorls, with regular, strong ribs; with ca. 21 ribs in 
<normalizedToken originalValue="¼">1/4</normalizedToken>
whorl adjoining R2, ribs sharper near suture and flatter near umbilicus, most prominent when approaching R2. Boundary between R1 and R2 distinct as R2 contains more close-set ribs than R1; R2 with 36-43 ribs that are curved towards aperture; R2 ca. 
<normalizedToken originalValue="¼">1/4</normalizedToken>
whorl, as long as R3. Boundary between R2 and R3 distinct due to shallow constriction; R3 smooth with indistinct growth lines; with either sharp and narrow or blunt and more elongated swelling in the middle of R3 or near aperture. Aperture round; slightly oblique to shell axis. Peristome double with prominent outer peristome. Inner peristome blunt, slightly protruding. Outer peristome strongly thickened, expanded, reflected, most conspicuously at the upper palatal position and near the umbilicus, multilayered (visible in lateral view). Umbilicus is partly or entirely closed by the reflected outer peristome. Operculum thin; multispiral ridges not significantly elevated.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">In honor of Prof. John B. Burch, a prominent American malacologist, who, together with Prof. S. Panha, extensively studied the taxonomy and systematics of micro land snails of Thailand.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">The new species is known from the type locality, Tham Phaya Nakharat, Khon Kaen Province, Northeastern Thailand. The other localities are Doi Chiang Dao, Chiang Mai Province and Ban Soppong, Mae Hong Son Province, Northern Thailand, and also from Shan State, Myanmar.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Differs from 
<taxonomicName genus="D." lsidName="D. vestitus" pageId="0" pageNumber="1" rank="species" species="vestitus">
<emphasis italics="true" pageId="0" pageNumber="1">D. vestitus</emphasis>
</taxonomicName>
(Blanford, 1862) from Moditoung [Modi Taung, Arakan Hills] (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11B</figureCitation>
), and its subspecies 
<taxonomicName genus="D." lsidName="D. v. subsp. akyabensis" pageId="0" pageNumber="1" rank="subSpecies" species="v." subSpecies="akyabensis">
<emphasis italics="true" pageId="0" pageNumber="1">D. v. akyabensis</emphasis>
</taxonomicName>
(Godwin-Austen, 1914) from Baumi, Akyab [Sittwe] (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11C</figureCitation>
), both from Rhakhine State, Myanmar, by having a smaller shell with a higher spire, a longer last whorl and a less distinct and a narrower constriction between R2 and R3. Differs from 
<taxonomicName genus="D." lsidName="D. conicus" pageId="0" pageNumber="1" rank="species" species="conicus">
<emphasis italics="true" pageId="0" pageNumber="1">D. conicus</emphasis>
</taxonomicName>
(Godwin-Austen, 1871) from Samiamri, E of the Kopili R., Assam State, India (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Dicharax burchi sp. nov., specimen HA (locality code: 2018 / 41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; B. Dicharax vestitus, holotype NHMUK 1906.4.4.53; C. Dicharax vestitus akyabensis, syntype NHMUK 1888.12.4.251 - 252; D. Dicharax conicus, syntype NHMUK 1903.7.1.2674. Photo: NHM (B-D)." figureDoi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1">11D</figureCitation>
) in having a longer last whorl, more close-set ribs on R2, and the inner and outer peristomes are narrower and less reflected. Differs from 
<taxonomicName genus="D." lsidName="D. imitator" pageId="0" pageNumber="1" rank="species" species="imitator">
<emphasis italics="true" pageId="0" pageNumber="1">D. imitator</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi in 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Do, DS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331: 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al., 2017
</bibRefCitation>
from Guangxi, China in having a less smooth R1, a much thicker peristome, a longer R2, a shallower constriction between R2 and R3, the last whorl less oblique compared to the columellar axis and a less triangular aperture (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B45" refString="Pall-Gergely, B, Hunyadi, A, Do, DS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331: 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zse.97.59143.figure11" httpUri="https://binary.pensoft.net/fig/494375" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">A.</emphasis>
<taxonomicName authorityName="Jirapatrasilp &amp; Pall-Gergely" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax burchi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burchi">
<emphasis italics="true" pageId="0" pageNumber="1">Dicharax burchi</emphasis>
</taxonomicName>
sp. nov., specimen HA (locality code: 2018/41) from Hsihseng centre E ca. 6 km, limestone hill, Shan State, Myanmar; 
<emphasis bold="true" pageId="0" pageNumber="1">B.</emphasis>
<taxonomicName baseAuthorityName="W.T.Blanford" baseAuthorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax vestitus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vestitus">
<emphasis italics="true" pageId="0" pageNumber="1">Dicharax vestitus</emphasis>
</taxonomicName>
, holotype NHMUK 1906.4.4.53; 
<emphasis bold="true" pageId="0" pageNumber="1">C.</emphasis>
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax vestitus subsp. akyabensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="vestitus" subSpecies="akyabensis">
<emphasis italics="true" pageId="0" pageNumber="1">Dicharax vestitus akyabensis</emphasis>
</taxonomicName>
, syntype NHMUK 1888.12.4.251-252; 
<emphasis bold="true" pageId="0" pageNumber="1">D.</emphasis>
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1871" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" kingdom="Animalia" lsidName="Dicharax conicus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="conicus">
<emphasis italics="true" pageId="0" pageNumber="1">Dicharax conicus</emphasis>
</taxonomicName>
, syntype NHMUK 1903.7.1.2674. Photo: NHM (
<emphasis bold="true" pageId="0" pageNumber="1">B-D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
This species exhibits the variation of R1 sculpture from nearly smooth (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10A</figureCitation>
) to distinctly ribbed (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10C, D</figureCitation>
) and R3 swelling from sharp (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10B, C</figureCitation>
) to blunter and longer by having the anterior slope of swelling steeper than posterior slope (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dicharax burchi sp. nov. A. Holotype CUMZ 7428 / 1 with operculum; B, C. Specimen CUMZ 7431 from Doi Chiang Dao, Chiang Mai Province; D. Specimen UF 345622 from 10.2 km WNW of Soppong, Road 1095, Mae Hong Son Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure10" httpUri="https://binary.pensoft.net/fig/494374" pageId="0" pageNumber="1">10D</figureCitation>
). In addition, the umbilicus is open, or covered by the reflected outer peristome. The latter trait even shows variation within the same population (maybe also depends on the age of the examined specimens).
</paragraph>
</subSubSection>
</treatment>
</document>