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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.781.21106" ID-GBIF-Dataset="52fd4469-c493-40e0-adef-67c8f04619f6" ID-PMC="PMC6160833" ID-Pensoft-Pub="1313-2970-781-81" ID-PubMed="30271236" ID-ZBK="50D89B9AA54F4FF1BDEFE847458EF03D" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-781-81" ModsDocOrigin="ZooKeys 781" ModsDocTitle="Taxonomic review of genus Microrhagus Dejean, 1833 from Korea, with description of a new species (Coleoptera, Eucnemidae, Melasinae, Dirhagini)" checkinTime="1534245671565" checkinUser="pensoft" docAuthor="Seung, Jinbae &amp; Lee, Seunghwan" docDate="2018" docId="3E2CD522D066D6AACBFAC7E71B8207E9" docLanguage="en" docName="ZooKeys 781: 81-95" docOrigin="ZooKeys 781" docSource="http://dx.doi.org/10.3897/zookeys.781.21106" docTitle="Microrhagus mystagogus Fleutiaux 1923" docType="treatment" docVersion="4" lastPageNumber="81" masterDocId="3B22FFE6FFFE143EFFC14322FFF37608" masterDocTitle="Taxonomic review of genus Microrhagus Dejean, 1833 from Korea, with description of a new species (Coleoptera, Eucnemidae, Melasinae, Dirhagini)" masterLastPageNumber="95" masterPageNumber="81" pageNumber="81" updateTime="1668166161126" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic review of genus Microrhagus Dejean, 1833 from Korea, with description of a new species (Coleoptera, Eucnemidae, Melasinae, Dirhagini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seung, Jinbae</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Seunghwan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>781</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>95</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.781.21106</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.781.21106</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-781-81</mods:identifier>
<mods:identifier type="ZBK">50D89B9AA54F4FF1BDEFE847458EF03D</mods:identifier>
<mods:identifier type="ZooBank">50D89B9AA54F4FF1BDEFE847458EF03D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="147279698" LSID="urn:lsid:plazi:treatment:3E2CD522D066D6AACBFAC7E71B8207E9" httpUri="http://treatment.plazi.org/id/3E2CD522D066D6AACBFAC7E71B8207E9" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName authority="Fleutiaux, 1923" authorityName="Fleutiaux" authorityYear="1923" class="Insecta" family="Eucnemidae" genus="Microrhagus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microrhagus mystagogus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="mystagogus">Microrhagus mystagogus (Fleutiaux, 1923)</taxonomicName>
Fig. 3
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="reference_group">
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Eucnemidae" genus="Dirhagus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dirhagus mystagogus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="mystagogus">Dirhagus mystagogus</taxonomicName>
Fleutiaux, 1923: 309.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="diagnosis">
<paragraph pageId="0" pageNumber="81">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="81">
Body: mostly dull black. Head: antennae pectinate from antennomere IV in male. Prothorax: pronotum convex, as long as wide; notosternal antennal grooves subparallel-sided. Pterothorax: elytra 2.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than combined width; metepisternum narrow, slightly widened posteriorly, its greatest width narrower than outer edge of metacoxal plate; metacoxal plate expanded inward; abdominal ventrite V simply rounded at apex.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="redescription">
<paragraph pageId="0" pageNumber="81">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="81">
Male (Fig. 3A,
<normalizedToken originalValue="CD">C-D</normalizedToken>
) 3.2-3.9 mm long and 1.0-1.2 mm wide. Body mostly black; antennae and femur dark red-brown; tibiae and tarsi yellow-brown; surface weakly glossy, with yellow pubescence. Head with circular punctures, becoming denser and irregularly sized near frontoclypeal region; frons without carina or groove at midline; frontoclypeal region slightly depressed at base, rounded and weakly sinuate at anterior edge, anterior edge 3.9
<normalizedToken originalValue="×">x</normalizedToken>
wider than distance between antennal sockets (Fig. 3G). Antennae (Fig. 3E) almost exceeding metacoxal plate, with yellow-brown pubescence, and pectinate from antennomere IV; processes of antennomeres IV, V, and VI 1.4, 2.3 and 2.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as corresponding antennomeres; antennomere I stout; antennomere II shortest; antennomere III gradually expanded toward apex, 1.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, three
<normalizedToken originalValue="×">x</normalizedToken>
longer than II, and 1.8
<normalizedToken originalValue="×">x</normalizedToken>
longer than IV; antennomeres
<normalizedToken originalValue="IVX">IV-X</normalizedToken>
with processes near apex, gradually lengthened toward apex; apical antennomere strongly elongate, curved, 11.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, and 2.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than previous one. Pronotum as long as wide, parallel-sided; surface with more regularly sized and spaced punctures than on head, slightly larger and sparser posteriorly; disc with a short carina at base of midline; anterolateral carina short, almost reaching one-third length of pronotum; posterolateral carina exceeding two-thirds length of pronotum; antescutellar area almost straight; pronotal posterior angles sharply projecting, exceeding posterior edge of antescutellar area. Scutellum subtriangular, 1.1
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, gradually narrowed posteriorly, and rounded at apex; surface rough, barely pubescent. Elytra 2.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than combined width, parallel-sided, gradually narrowing near apices; disc barely striate, with shallow and irregularly sized and spaced punctures; several large and deep punctures present near apices; apices simply rounded. Prosternum subparallel-sided, anterior margin shallowly bisinuate; surface mostly with punctures as on pronotum, slightly larger laterally; prosternal process robust basally, abruptly tapered and curved dorsally at posterior end; hypomeron mostly punctate as prosternum; surface wrinkled at coxal cavities; notosternal antennal grooves (Fig. 3I) subparallel-sided, with outer marginal carina, rarely punctate, glabrous, and with pits. Mesoventrite with coarse surface; mesopleuron with irregularly sized and spaced punctures, especially anteriorly. Metaventrite mostly with finer punctures than on prosternum, slightly larger laterally; disc with a groove at midline, not reaching anterior margin; metepisternum (Fig. 3J) widened posteriorly, its greatest width four-fifths of outer edge of metacoxal plate; metacoxal plate (Fig. 3K) expanded inward, medially 1.8
<normalizedToken originalValue="×">x</normalizedToken>
wider than laterally. Legs (Fig. 3O) slender; metatarsomere I 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than
<normalizedToken originalValue="IIIV">II-IV</normalizedToken>
combined; metatarsomere II 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than III; metatarsomere V 1.2
<normalizedToken originalValue="×">x</normalizedToken>
longer than II; claws simple. Abdomen with finer and denser punctures than on metaventrite; ventrite V simply rounded at apex (Fig. 3L). Aedeagus (Fig. 3
<normalizedToken originalValue="MN">M-N</normalizedToken>
) 3.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, compressed dorsoventrally; median lobe slightly bent ventrally, fused with lateral lobes; lateral lobes strongly curved ventrally, enlarged near apex, densely setose; ventral lobe bifurcate apically, with dense short pubescence; phallobase strongly emarginate basally, 1.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, approximately one-fifth as long as aedeagus. Female (Fig. 3B) is distinguished from male by following characters: body stouter, 3.4-3.9 mm long and 1.1-1.3 mm wide; frontoclypeal region (Fig. 3H) with anterior edge, 3.7
<normalizedToken originalValue="×">x</normalizedToken>
wider than distance between antennal sockets; antennae (Fig. 3F) serrate, almost reaching metacoxal plate; antennomere III subrectangular, 3.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, 2.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than II, and 1.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than IV; antennomeres
<normalizedToken originalValue="IVX">IV-X</normalizedToken>
gradually more strongly serrate; apical antennomere 3.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, and approximately twice longer than X.
</paragraph>
<caption pageId="0" pageNumber="81">
<paragraph pageId="0" pageNumber="81">
Figure 3.
<taxonomicName class="Insecta" family="Eucnemidae" genus="Microrhagus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microrhagus mystagogus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="mystagogus">Microrhagus mystagogus</taxonomicName>
(Fleutiaux, 1923). A,
<normalizedToken originalValue="CE">C-E</normalizedToken>
, G,
<normalizedToken originalValue="IO">I-O</normalizedToken>
male B, F, H, female A, B dorsal habitus C ventral habitus D lateral habitus
<normalizedToken originalValue="EF">E-F</normalizedToken>
antenna
<normalizedToken originalValue="GH">G-H</normalizedToken>
frons I antennal groove J metepisternum K metacoxal plate L abdominal ventrite V
<normalizedToken originalValue="MN">M-N</normalizedToken>
aedeagus O metatarsus. Scale bar: 1 mm (
<normalizedToken originalValue="AF">A-F</normalizedToken>
); 0.5 mm (
<normalizedToken originalValue="GO">G-O</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="materials_examined">
<paragraph pageId="0" pageNumber="81">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="81">
&lt;Gyeonggi-do&gt; 1♀, Deoksu-ri, Danwol-myeon, Yangpyeong-gun,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="37.55638">N37°33'22.97&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="127.687126">E127°41'13.65&quot;</geoCoordinate>
, 166m alt., flight intercept trap, 29
<normalizedToken originalValue="June">June-</normalizedToken>
16 July, 2016, Seung and Jung leg. (SNU). &lt;Gangwon-do&gt; 1♀, Yeongheung-ri, Yeongwol-eup, Yeongwol-gun,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="37.205513">N37°12'19.84&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="128.45493">E128°27'17.77&quot;</geoCoordinate>
, 308m alt., flight intercept trap, 19
<normalizedToken originalValue="June">June-</normalizedToken>
02 July, 2015, Seung and Lee leg. (SNU); 1♂, Beopheung-ri, Suju-myeon, Yeongwol-gun,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="37.378105">N37°22'41.19&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="128.2543">E128°15'15.50&quot;</geoCoordinate>
, 550m alt., flight intercept trap, 03-16 July. 2015, Seung and Lee leg. (SNU); 1♂2♀, Beopheung-ri, Suju-myeon, Yeongwol-gun,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="37.378105">N37°22'41.19&quot;</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="128.2543">E128°15'15.50&quot;</geoCoordinate>
, 550m alt., flight intercept trap, 05-29 June, 2016, Seung and Jung leg. (SNU).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">Korea (New record), Japan, Russia (Far East).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="remarks">
<paragraph pageId="0" pageNumber="81">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Eucnemidae" genus="Microrhagus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microrhagus mystagogus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="mystagogus">Microrhagus mystagogus</taxonomicName>
is easily distinguished from other Korean
<taxonomicName class="Insecta" family="Eucnemidae" genus="Microrhagus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microrhagus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Microrhagus</taxonomicName>
species by its antennae: male antennae pectinate from antennomere IV. Also, convex pronotum and structure of aedeagus are characteristic. Its aedeagal structure is well-illustrated in
<bibRefCitation author="Kovalev, AV" journalOrPublisher="Proceedings of the Zoological Institute RAS" pageId="0" pageNumber="81" pagination="268 - 274" title="Two new species of the tribe Dirhagini (Coleoptera: Eucnemidae) from Palaearctic region." volume="317" year="2013">
<normalizedToken originalValue="Kovalevs">Kovalev's</normalizedToken>
(2013)
</bibRefCitation>
work.
</paragraph>
</subSubSection>
</treatment>
</document>