424 lines
60 KiB
XML
424 lines
60 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.46.104548" ID-Pensoft-Pub="1314-2615-46-21" ID-Pensoft-UUID="7C8C1294D4B05D749CC9DF10F2D7458D" ID-ZooBank="17340FB714C84903B1E48B209FE73C93" ModsDocID="1314-2615-46-21" checkinTime="1692896423125" checkinUser="pensoft" docAuthor="Curcic, Srecko, Vesovic, Nikola, Vrbica, Maja, Popovic, Slađana, Radovanovic, Zeljko, Curcic, Nina B., Yamashkin, Anatoliy A., Radovic, Dejan, Yamashkin, Stanislav A., Vranic, Sofija & Rađa, Tonci" docDate="2023" docId="D9C1EB2C360F50F28CB780B4DCCB189D" docLanguage="en" docName="SubterBiol 46: 21-46" docOrigin="Subterranean Biology 46" docPubDate="2023-08-24" docSource="http://dx.doi.org/10.3897/subtbiol.46.104548" docTitle="Spelaeobates (Spelaeobates) novaki Mueller 1901" docType="treatment" docVersion="2" id="7C8C1294D4B05D749CC9DF10F2D7458D" lastPageNumber="21" masterDocId="7C8C1294D4B05D749CC9DF10F2D7458D" masterDocTitle="The surprising discovery of two new subterranean Leptodirini of the genus Spelaeobates Mueller, 1901 (Coleoptera, Leiodidae, Cholevinae) from Croatia after more than a century" masterLastPageNumber="46" masterPageNumber="21" pageNumber="21" updateTime="1692896725838" updateUser="ExternalLinkService">
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<mods:title id="8E690BC1C7F3463BD5B0798AF7BC502C">The surprising discovery of two new subterranean Leptodirini of the genus Spelaeobates Mueller, 1901 (Coleoptera, Leiodidae, Cholevinae) from Croatia after more than a century</mods:title>
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<mods:namePart id="CC296ECAAC4DBC88EECD29800A564C8B">Curcic, Srecko</mods:namePart>
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<mods:affiliation id="3C8BD8F08C52C6A687E756D5465285A3">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:nameIdentifier id="2D268C1AAADCCC27B79DBCE07AD6F215" type="email">srecko@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="473530BC96EEC0E5C7BC69E7954C75FE">Vesovic, Nikola</mods:namePart>
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<mods:affiliation id="E4A99E300A4CAF5D63941808DEF6879C">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="D0194348BB3862602694467D373BFBB1">Vrbica, Maja</mods:namePart>
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<mods:affiliation id="44AC61AFB0A22C7CBE5BDFFAE3396B7D">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="9C8F0A3170D7B2455215C1E0C746BF3A">Popovic, Slađana</mods:namePart>
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<mods:affiliation id="A5924B44FF31BEB34950E7D4139E720F">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="8ED59EE6C9DC7E5975559113386F455C">Radovanovic, Zeljko</mods:namePart>
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<mods:affiliation id="AF6CBB93F213D92F7E536EF5F457F27B">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="55AD829A2E4EC4ECD6858722E7DD4004">Curcic, Nina B.</mods:namePart>
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<mods:affiliation id="CD5726B2DC1E85847D75B51725F2BB13">University of Belgrade, Innovation Center of the Faculty of Technology and Metallurgy, Karnegijeva 4, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="884F0A5E692EFDA01465D0699082A558">Yamashkin, Anatoliy A.</mods:namePart>
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<mods:affiliation id="E14CDD8019E36173D77C539F9B2904B7">Geographical Institute " Jovan Cvijic ", Serbian Academy of Sciences and Arts, Đure Jaksica 9, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="42DC753F2C817D7DE64047CB03612EF6">Radovic, Dejan</mods:namePart>
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<mods:affiliation id="91A70D20C15F4DCC8377A5C4C73D193D">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="66813309FA47B6F885813E39E43253EC">Yamashkin, Stanislav A.</mods:namePart>
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<mods:affiliation id="BE1C081E63DD3E9DDB45624DE9770E03">Geographical Institute " Jovan Cvijic ", Serbian Academy of Sciences and Arts, Đure Jaksica 9, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="9116F8729C7B424B889A359A06D79390">Vranic, Sofija</mods:namePart>
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<mods:affiliation id="6977EE3FD32F06E91D3FFCC88D7ED0F9">University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia</mods:affiliation>
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<mods:namePart id="F6C1D52D81887A2E4A8BBC64CC1A26CC">Rađa, Tonci</mods:namePart>
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<mods:affiliation id="F80250A9602C9CB87BFFDD5B561F5DF6">National Research Ogarev Mordovia State University, Faculty of Geography, Sovetskaya Str. 24, 430005 Saransk, Russia</mods:affiliation>
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<mods:title id="F96314C251B84909125742A8A7AE8A48">Subterranean Biology</mods:title>
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<mods:part id="AE4CB61493F86653923420A6A7BA677E">
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<mods:date id="F14CF9A0A8754263594376ABB07BA7FC">2023</mods:date>
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<mods:number id="9E3F35DC32397F9781DB9EA6775B0D7B">2023-08-24</mods:number>
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<mods:start id="79D61C775747F6497CBA23B768878E90">21</mods:start>
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<treatment id="D9C1EB2C360F50F28CB780B4DCCB189D" LSID="urn:lsid:plazi:treatment:D9C1EB2C360F50F28CB780B4DCCB189D" httpUri="http://treatment.plazi.org/id/D9C1EB2C360F50F28CB780B4DCCB189D" lastPageNumber="21" pageId="0" pageNumber="21">
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<subSubSection id="586CCA7B14216AF20497C3735EE0039A" pageId="0" pageNumber="21" type="nomenclature">
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<paragraph id="9144A77103066B5B699E53F3344A667A" pageId="0" pageNumber="21">
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<taxonomicName id="EB18FC837B1CD5ADE9CF0310274B8097" ID-CoL="4YH9P" LSID="D9C1EB2C-360F-50F2-8CB7-80B4DCCB189D" authority="Mueller, 1901" authorityName="Mueller" authorityYear="1901" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) novaki" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="novaki" subGenus="Spelaeobates">
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Spelaeobates (Spelaeobates) novaki
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<normalizedToken id="D3DF0664AA7688317FA7E2B2C3A9F5F7" originalValue="Müller">Mueller</normalizedToken>
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, 1901
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</taxonomicName>
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<figureCitation id="554BAB2380BD31886F99FE119ED0C5E8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">Figs 7</figureCitation>
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<figureCitation id="731C5D786A3F3B47A8C536240F57CF53" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">, 8</figureCitation>
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</paragraph>
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<subSubSection id="D46E4DF9F4451051B771EDB4873E3BA9" pageId="0" pageNumber="21" type="type material">
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<paragraph id="38715C5471998DC440DEAA8A068B4697" pageId="0" pageNumber="21">Type material.</paragraph>
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<paragraph id="723A44D398B1A5F30C5FD0C1427129B8" pageId="0" pageNumber="21">
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<taxonomicName id="7FC17105CCE7593B907B945577087DDC" genus="Topotypes" lsidName="Topotypes" pageId="0" pageNumber="21" rank="genus">
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<emphasis id="7A00FE27238269FE5AF28519ED3982E6" bold="true" italics="true" pageId="0" pageNumber="21">Topotypes</emphasis>
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</taxonomicName>
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: one male and four females (IZFB) labeled as follows: "CROATIA, NORTHERN DALMATIA: island of Dugi Otok, village of Savar,
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<normalizedToken id="2DE0BEA11256A68E33A46A6A77522638" originalValue="Strašna">Strasna</normalizedToken>
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<normalizedToken id="F5A92FEFE8C0C330EB8555D12A2BF048" originalValue="Peć">Pec</normalizedToken>
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Cave, 70 m a.s.l.,
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<geoCoordinate id="EDCC9BB259818E1B45136EE1724E8EA1" degrees="44" direction="north" minutes="00" orientation="latitude" precision="1" seconds="16.6" value="44.004612">44°00'16.6"N</geoCoordinate>
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,
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<geoCoordinate id="49C96DF10C0EAB4206FCBEE50809B67D" degrees="15" direction="east" minutes="02" orientation="longitude" precision="1" seconds="19.1" value="15.038639">15°02'19.1"E</geoCoordinate>
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, 1.VII.1997, TR". All topotypes are labeled with white, printed locality labels (Fig.
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<figureCitation id="27AB923915EA17BFD3A81C9AD1DBF8C9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7</figureCitation>
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).
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</paragraph>
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<caption id="446113DD747BC0958BCA39A77A0BE039" doi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21" start="Figure 7" startId="F7">
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<paragraph id="57AB89E4F94D25BC8A87CD8417AF62D5" pageId="0" pageNumber="21">
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<emphasis id="40B33365981EA118071774C2397FD8AD" bold="true" pageId="0" pageNumber="21">Figure 7.</emphasis>
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Bright-field images of the morphological structures of topotype male (
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<emphasis id="02D1C8276769058980BC6DBEE2331BC7" bold="true" pageId="0" pageNumber="21">A-G</emphasis>
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) and topotype female (
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<emphasis id="FD7C575325A9AC4DC9EA51DA9CB1C546" bold="true" pageId="0" pageNumber="21">H</emphasis>
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) of
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<taxonomicName id="6EE25E9B53773DEC9FEA329FA5E858A2" authorityName="Mueller" authorityYear="1901" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) novaki" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="novaki" subGenus="Spelaeobates">Spelaeobates (Spelaeobates) novaki</taxonomicName>
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from the
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<normalizedToken id="400102E171309317CFC75DA9944335E7" originalValue="Strašna">Strasna</normalizedToken>
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<normalizedToken id="2A2DB5C242409093086087C69AAD70BB" originalValue="Peć">Pec</normalizedToken>
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Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia
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<emphasis id="CC56438DBE9BAC3368621C63F1CAA83D" bold="true" pageId="0" pageNumber="21">A</emphasis>
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habitus, dorsal view
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<emphasis id="B32EA3AEF74B2F864AF04449FECEDCB4" bold="true" pageId="0" pageNumber="21">B</emphasis>
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habitus, lateral view
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<emphasis id="C0135A8B518EE44C4DCA663D73DEF26B" bold="true" pageId="0" pageNumber="21">C</emphasis>
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head, dorsal view
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<emphasis id="94BA6A2E5A2C3D4C7F05DA0D780ABE24" bold="true" pageId="0" pageNumber="21">D</emphasis>
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left antenna, dorsal view
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<emphasis id="06FB73C0459972B8BBA215583482F444" bold="true" pageId="0" pageNumber="21">E</emphasis>
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pronotum and scutellum, dorsal view
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<emphasis id="EED7AD131948375279B23B4934D882EE" bold="true" pageId="0" pageNumber="21">F</emphasis>
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elytra, dorsal view
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<emphasis id="DBFC0F5E3C9867CD347ECE8FB1A1F893" bold="true" pageId="0" pageNumber="21">G</emphasis>
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mesoventral carina, lateral view
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<emphasis id="9874DEAC41F2AE96BAA5C4E5CD2029BA" bold="true" pageId="0" pageNumber="21">H</emphasis>
|
|
mesoventrite, ventral view. Scale bars: 1.0 mm (
|
|
<emphasis id="948F42F778E5717AF84628984DFB17FF" bold="true" pageId="0" pageNumber="21">A, B</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis id="16482CE96258E1BC3997C9C7301E93A5" bold="true" pageId="0" pageNumber="21">D, F</emphasis>
|
|
); 0.25 mm (
|
|
<emphasis id="4C9F29B37021B2A04C963E557849303F" bold="true" pageId="0" pageNumber="21">C, G</emphasis>
|
|
); 0.2 mm (
|
|
<emphasis id="35B11A919191CD7D5AAF94CD12F0EC2E" bold="true" pageId="0" pageNumber="21">E, H</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="9C94FDCBEECB55D4CD75ABAD2B6B5A10" pageId="0" pageNumber="21" type="remarks">
|
|
<paragraph id="522C1854DCF6730713D913D454A0325E" pageId="0" pageNumber="21">Remarks.</paragraph>
|
|
<paragraph id="2A86D8658597A09EA778C2081096F2CA" pageId="0" pageNumber="21">
|
|
For purpose of comparisons, we have examined the topotype material of
|
|
<taxonomicName id="367BBAA6F3B5B95143F098D8C0B59821" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
collected by the last author of this study. This species is described on the basis of the type series of specimens collected in September 1900 by Josef
|
|
<normalizedToken id="C47B0B2F407BC9FE8F75273A15F1D0CB" originalValue="Müller">Mueller</normalizedToken>
|
|
and Petar Novak in two caves on two northern Dalmatian islands - the
|
|
<normalizedToken id="4A3B1E7989EAC38F45CF5A3E2793E0DE" originalValue="Strašna">Strasna</normalizedToken>
|
|
<normalizedToken id="F9C838FFC61F1A12E9923D61FDDC8D3E" originalValue="Peć">Pec</normalizedToken>
|
|
Cave, village od Savar, island of Dugi Otok, and a small cave in the village of Mali
|
|
<normalizedToken id="A4F5A778E4386A3DE24964BC3C6644A4" originalValue="Iž">Iz</normalizedToken>
|
|
, island of
|
|
<normalizedToken id="D6A1760B8756D29803F3DB53AF1C0615" originalValue="Iž">Iz</normalizedToken>
|
|
(
|
|
<bibRefCitation id="FE3AA6BD2AA92DB0C195BA9691B5881A" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="F5BAC34E78954E88B3720065BDE39AFD" originalValue="Müller">Mueller</normalizedToken>
|
|
1901
|
|
</bibRefCitation>
|
|
). Later
|
|
<bibRefCitation id="17515C614E3D4E80B2254110C9B52741" author="Pretner, E" journalOrPublisher="Krs Jugoslavije" pageId="0" pageNumber="21" pagination="101 - 239" refId="B28" refString="Pretner, E, 1973. Koleopteroloska fauna pecina i jama Hrvatske s historijskim pregledom istrazivanja. Krs Jugoslavije 8: 101 - 239" title="Koleopteroloska fauna pecina i jama Hrvatske s historijskim pregledom istrazivanja." volume="8" year="1973">Pretner (1973)</bibRefCitation>
|
|
determined the correct name of the second cave site (its exact name is the Jezero Cave). In his original description of
|
|
<taxonomicName id="A4AB48CBF689B004F8E63301228069A3" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
,
|
|
<bibRefCitation id="6BD7EB75872CE3BA39998C513D60E399" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="D41D838085D794760100179BEB511C10" originalValue="Müller">Mueller</normalizedToken>
|
|
(1901)
|
|
</bibRefCitation>
|
|
did not indicate how many specimens are included in the type series, nor is there any information about their sex or where they were deposited. Based on the data and illustrations in the paper of
|
|
<bibRefCitation id="6338341D1A2490F13D082DCDFDF693E6" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="A6329E9E0CFFB4377F82465AC52F66B0" originalValue="Müller">Mueller</normalizedToken>
|
|
(1901)
|
|
</bibRefCitation>
|
|
, it can be concluded that the type series of this species consisted of both male and female specimens.
|
|
</paragraph>
|
|
<paragraph id="EA16A57B8634DB80466E53C2615DB0DE" pageId="0" pageNumber="21">
|
|
After reading both the original description of
|
|
<taxonomicName id="3B44D7981AD686EFD3889738E16B2FFE" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
by
|
|
<bibRefCitation id="29C68F5279A48F8882E5D0CEEDED12E6" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="762C4D3013576E23D9E905AA11BD57FB" originalValue="Müller">Mueller</normalizedToken>
|
|
(1901)
|
|
</bibRefCitation>
|
|
and the subsequent morphological data on the species by
|
|
<bibRefCitation id="32E4B7F49831748F33CC009134BBA05B" author="Jeannel, R" journalOrPublisher="Archives de Zoologie Experimentale et Generale" pageId="0" pageNumber="21" pagination="1 - 436" refId="B15" refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie Experimentale et Generale 63: 1 - 436" title="Monographie des Bathysciinae. Biospeologica L." volume="63" year="1924">Jeannel (1924)</bibRefCitation>
|
|
, as well as a careful examination of the topotype specimens of
|
|
<taxonomicName id="F245879AD88440C11DC600D2A65492AA" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
, we have found that some of the data given in the earlier literature on the morphology of the species do not agree with the characteristics of the topotype specimens we have observed. Namely, both
|
|
<bibRefCitation id="F9993242D3A3D4B4D05F1C6C3ED66547" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="C43B78D831501E7C69FFC570B0D179AF" originalValue="Müller">Mueller</normalizedToken>
|
|
(1901)
|
|
</bibRefCitation>
|
|
and
|
|
<bibRefCitation id="C333D69058BA7F594FF9001083CFBBB5" author="Jeannel, R" journalOrPublisher="Archives de Zoologie Experimentale et Generale" pageId="0" pageNumber="21" pagination="1 - 436" refId="B15" refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie Experimentale et Generale 63: 1 - 436" title="Monographie des Bathysciinae. Biospeologica L." volume="63" year="1924">Jeannel (1924)</bibRefCitation>
|
|
reported that the parameres of the aedeagus of
|
|
<taxonomicName id="088B4C7D8F767E59BADF4FE394E54F35" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
lack setae. Furthermore, in the drawing of the aedeagus by
|
|
<bibRefCitation id="6EBCC1FA1B350E1E5E92878CA9BAAEAF" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="2B38AF18CAD024195E339AF89C664C7F" originalValue="Müller">Mueller</normalizedToken>
|
|
(1901)
|
|
</bibRefCitation>
|
|
, no parameral setae are present. However, in the topotype male of
|
|
<taxonomicName id="F32231871011F974935A7D2FD8C22DED" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
, we observed that each paramere has four apical setae, as in
|
|
<taxonomicName id="213027265A421B8BA6646E4F1F583732" lsidName="S. (S.) coriniensis" pageId="0" pageNumber="21" rank="species" species="coriniensis" subGenus="S.">S. (S.) coriniensis</taxonomicName>
|
|
sp. nov. In the work of
|
|
<bibRefCitation id="345426EA4F676C214DABCD609EA9C60F" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="8F7BE5CB04C0DC7A4E6559FEB03282A3" originalValue="Müller">Mueller</normalizedToken>
|
|
(1901)
|
|
</bibRefCitation>
|
|
, it was noted that the head of
|
|
<taxonomicName id="C3500E58763D20BF91B588A669FFF9EF" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
is nearly twice as long as wide, the pronotum is 1.5 times longer than wide, antennomeres I and II are of similar width, the last antennomere is wider than the preceding ones, and the first protarsomere of the males is about twice as long as wide. However, in the specimens of the same species that we have examined, the head is about 1.5 times as long as wide, the pronotum is nearly one third longer than wide, antennomere I is wider than antennomere II, the last antennomere is narrower than the preceding ones, and the first protarsomere of the male is about 1⅔ times longer than wide (Table
|
|
<tableCitation id="16A3233EAD886AA5E748AD063A877340" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Linear measurements and morphometric ratios in Spelaeobates (Spelaeobates) coriniensis coriniensis ssp. nov., S. (S.) coriniensis nonveilleri ssp. nov., and S. (S.) novaki. Values outside parentheses are mean values, while those inside parentheses are ranges." httpUri="http://table.plazi.org/id/A94F7FBCE48202F29A888BF18D2FCAD6" pageId="0" pageNumber="21" tableUuid="A94F7FBCE48202F29A888BF18D2FCAD6">1</tableCitation>
|
|
).
|
|
<bibRefCitation id="8A4E72D59DD970BBB50422795BF28040" author="Mueller, J" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="0" pageNumber="21" pagination="16 - 33" refId="B17" refString="Mueller, J, 1901. Beitrag zur Kenntniss der Hoehlensilphiden. Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 16 - 33" title="Beitrag zur Kenntniss der Hoehlensilphiden." volume="51" year="1901">
|
|
<normalizedToken id="93ECEA41A88BAAB179AE64C0643B4CA3" originalValue="Müller">Mueller</normalizedToken>
|
|
(1901)
|
|
</bibRefCitation>
|
|
noted that the pronotum of
|
|
<taxonomicName id="29A981FFF70BFF7A74295B4C5DEF40D6" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
is finely punctate, but
|
|
<bibRefCitation id="EF0F69A716130D885B53F47822808D14" author="Jeannel, R" journalOrPublisher="Archives de Zoologie Experimentale et Generale" pageId="0" pageNumber="21" pagination="1 - 436" refId="B15" refString="Jeannel, R, 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie Experimentale et Generale 63: 1 - 436" title="Monographie des Bathysciinae. Biospeologica L." volume="63" year="1924">Jeannel (1924)</bibRefCitation>
|
|
reported that it is strongly punctate, which we also observed in our specimens. For these reasons, we have decided to redescribe the species
|
|
<taxonomicName id="9345E6A50964B0A886FEE6EFC5AFEC4D" lsidName="S. (S.) novaki" pageId="0" pageNumber="21" rank="species" species="novaki" subGenus="S.">S. (S.) novaki</taxonomicName>
|
|
and add additional data on its morphology here.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E5E7733631822AC89292CF3FB39BF6A8" pageId="0" pageNumber="21" type="redescription">
|
|
<paragraph id="50656B379FEB20531667E42C5E9C470D" pageId="0" pageNumber="21">Redescription.</paragraph>
|
|
<paragraph id="2FF3B05C9D24A0471076A7D0F6ACA6A6" pageId="0" pageNumber="21">Small-sized leptodirine. TL M 2.63 mm (2.57 mm in males, 2.64 mm in females), R 2.57-2.70 mm (2.57 mm in males, 2.60-2.70 mm in females).</paragraph>
|
|
<paragraph id="9D397C2D5533D791D20D0932FEEF0CDB" pageId="0" pageNumber="21">
|
|
<emphasis id="8C225F6B62196BA65F8C4A6BFDA9AB14" bold="true" italics="true" pageId="0" pageNumber="21">Habitus</emphasis>
|
|
: Body shape leptodiroid (Fig.
|
|
<figureCitation id="9F8A26F8547B1D17888C47338ADD472B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7A, B</figureCitation>
|
|
), colour yellowish-brown.
|
|
</paragraph>
|
|
<paragraph id="359A33C8C0FEEE4BFB05257097680F9E" pageId="0" pageNumber="21">
|
|
<emphasis id="9F65425425773CAE0610CBC3CC6D11B6" bold="true" italics="true" pageId="0" pageNumber="21">Integument</emphasis>
|
|
: Lustrous, microsculptured both dorsally and ventrally (Fig.
|
|
<figureCitation id="B6B321F36763DA316848F6CF7F736AEE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7C, E, F, H</figureCitation>
|
|
). Densely distributed deep punctures present on head, pronotum (often merged) and elytra (particularly strong) (Fig.
|
|
<figureCitation id="4A51FCA2366CF9A2E28322E4750328B3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7C, E, F</figureCitation>
|
|
). Entire body dorsally covered with yellow pubescence of short length (erect on head, while recumbent on both pronotum and elytra) (Fig.
|
|
<figureCitation id="587F259D091CFCE58B918ED53A8C5D8C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7A, B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="A6D64D0F6009F610DA8AADCB3A2225E8" pageId="0" pageNumber="21">
|
|
<emphasis id="EFCC8D77A1A7D84DE2F008E3C8F82005" bold="true" italics="true" pageId="0" pageNumber="21">Head</emphasis>
|
|
: About one and a half times as long as wide (HL/HW M 1.47, R 1.38-1.53), slightly more elongate in males (HL/HW M 1.49 in males, M 1.46 in females), with no eyes, occipital carina in the shape of a curved concave line (Fig.
|
|
<figureCitation id="9E1EFEE52513EAD497720C6EBF31AB03" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7A, C</figureCitation>
|
|
). Head widest between first third and half. Frons roundly impressed between antennal insertions. Labrum transverse, with a few long setae. First maxillary palpomere of similar length and width, shorter than second maxillary palpomere. Maxillary palpomeres II and III of similar length (M3L/M2L M 1.09, R 1.00-1.18). Penultimate maxillary palpomere widened apically. Ultimate maxillary palpomere short, slender, gradually narrowing apically. Antennae inserted in basal quarter of head, thin, narrow proximally (except for first two antennomeres, which are thickened), slightly widened distally, longer in males, AL M 1.81 mm, R 1.75-1.93 mm (1.93 mm in males, 1.75-1.79 mm in females), not reaching end of elytra in both sexes (Fig.
|
|
<figureCitation id="E6C7403D1715AF8A98F44477D375AB21" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7A, B, D</figureCitation>
|
|
). Antennomeres I and II short and wide, of similar length, second of which slightly narrower. Following four antennomeres thinner and slightly longer than antennomere II. Antennomere III longer than adjacent antennomeres (A3L/A2L M 1.37, R 1.23-1.42; A3L/A4L M 1.24, R 1.21-1.31). Antennomeres VII, IX, and X quite expanded distally. Antennomere VIII relatively short and narrow, shorter and narrower than anatennomeres VII, IX, X, and XI. Ultimate antennomere slender, widened sub-distally, then narrowing apically, narrower than penultimate one (A11W/A10W M 0.75, R 0.67-0.83). Antennomere VIII shortest, while antennomeres IX and XI longest. Other ratios of length of certain antennomeres: A6L/A3L M 0.79, R 0.76-0.82; A8L/A3L M 0.65, R 0.59-0.71; A11L/A8L M 2.27, R 2.09-2.40.
|
|
</paragraph>
|
|
<paragraph id="F9B33A32DBA46B7A7D6EE144B2131A11" pageId="0" pageNumber="21">
|
|
<emphasis id="049D0007DC42C7684509B9649264A15F" bold="true" italics="true" pageId="0" pageNumber="21">Prothorax</emphasis>
|
|
: Pronotum bell-shaped, elongate, longer than wide (PL/PW M 1.29, R 1.26-1.32; M 1.32, R 1.32 in males; M 1.28, R 1.26-1.30 in females), widest slightly before anterior third, broader (HW/PW M 0.93, R 0.88-0.98) and shorter than head (PL/HL M 0.96, R 0.90-0.98) (Fig.
|
|
<figureCitation id="17A359AB91B56B75B80EFBFA806C2D59" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7A, E</figureCitation>
|
|
). Lateral margins rounded anteriorly, after which they constrict towards posterior end, slightly concave posteriorly. Pronotal base straight, somewhat shorter than elytral base. PB/AM M 0.86, R 0.81-0.90. Anterior margin straight. Lateral margins and pronotal base rimmed. Fore pronotal angles weakly expressed, rounded, obtuse. Hind pronotal angles well-expressed, obtuse, not protruding backwards. Pronotal disc moderately convex (Fig.
|
|
<figureCitation id="93C7562E4B19508A6870E1DB277A744D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="3B04B289DF3995124304154277422AA4" pageId="0" pageNumber="21">
|
|
<emphasis id="850950E13DAF8DF4946CAD5E9CF69DC2" bold="true" italics="true" pageId="0" pageNumber="21">Mesothorax</emphasis>
|
|
: Mesoventral carina very low, barely noticeable, with a few setae (Fig.
|
|
<figureCitation id="92EF5AAAA4FBBF76938D1C1AE4E54096" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7G</figureCitation>
|
|
). No tooth, anterior and posterior margins observed. Mesoventrite with a long, sub-parallel process between mesocoxae (Fig.
|
|
<figureCitation id="8CEE6F6D34143C21572AC06E1CD6FD29" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7H</figureCitation>
|
|
). Scutellum large, sub-triangular (Fig.
|
|
<figureCitation id="199F7ACCCEF20CF6F5B8573B3F0173C6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7E, F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="C941A36A596080E250844ACF9F321C45" pageId="0" pageNumber="21">
|
|
<emphasis id="8F95EEF99045FACFF962477B02FF6373" bold="true" italics="true" pageId="0" pageNumber="21">Metathorax</emphasis>
|
|
: Metaventrite without carina.
|
|
</paragraph>
|
|
<paragraph id="9D5262DAC7650C23760F559CC54D04F4" pageId="0" pageNumber="21">
|
|
<emphasis id="B04964DB02A85AB411FF0C58E4138423" bold="true" italics="true" pageId="0" pageNumber="21">Elytra</emphasis>
|
|
: Wide, ovoid, of similar width in males and females (EL/EW M 1.53, R 1.53 in males; M 1.55, R 1.49-1.60 in females), markedly wider than pronotum (EW/PW M 2.44, R 2.37-2.50) (Fig.
|
|
<figureCitation id="2EE79B362857DFDD37DD2EE3A87B8123" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7A, F</figureCitation>
|
|
). Maximum width a little before middle. Lateral margins arcuate. Marginal furrows not visible from above. Shoulders barely visible, obtuse, covered by hind pronotal angles. Elytral disc markedly convex, steeply declining both basally and apically in lateral view (Fig.
|
|
<figureCitation id="90CBF5BEB4875341070B5D19B5486DDA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7B</figureCitation>
|
|
). Parasutural stria absent. Elytral apex slightly attenuated, rounded. Pygidium not entirely covered by elytra.
|
|
</paragraph>
|
|
<paragraph id="5F4E8102A022B6ED732DC49739237FE2" pageId="0" pageNumber="21">
|
|
<emphasis id="67B54EC2590DDD67EEA237808F569FA5" bold="true" italics="true" pageId="0" pageNumber="21">Legs</emphasis>
|
|
: Elongate and slender (Fig.
|
|
<figureCitation id="969C3D3DBF4DC6126505065FA48FBCA9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bright-field images of the morphological structures of topotype male (A-G) and topotype female (H) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A habitus, dorsal view B habitus, lateral view C head, dorsal view D left antenna, dorsal view E pronotum and scutellum, dorsal view F elytra, dorsal view G mesoventral carina, lateral view H mesoventrite, ventral view. Scale bars: 1.0 mm (A, B); 0.5 mm (D, F); 0.25 mm (C, G); 0.2 mm (E, H)." figureDoi="10.3897/subtbiol.46.104548.figure7" httpUri="https://binary.pensoft.net/fig/896586" pageId="0" pageNumber="21">7A, B</figureCitation>
|
|
). Femora widened basally, constricted in distal half. Tibiae thin, gently curved, gradually widening distally. Each protibia with a very fine comb over entire apical third of outer margin. Fore tarsi four-segmented in both sexes, only first protarsomere in males dilated (P1W/P2W M 1.67, R 1.67). Tarsal claws thin, elongate, curved, pointed apically.
|
|
</paragraph>
|
|
<paragraph id="78913D345C606C22791DB83E8A2DA68B" pageId="0" pageNumber="21">
|
|
<emphasis id="A1B7AEDAB6081F6775A6197E4B2B91C0" bold="true" italics="true" pageId="0" pageNumber="21">Male genitalia</emphasis>
|
|
: Aedeagus elongate, slender, small, well chitinised (Fig.
|
|
<figureCitation id="60C94C862A70D9411AB4E62C0A628AC3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8A, B</figureCitation>
|
|
). Median lobe in dorsal view straight, sub-parallel, gradually narrowing distally, with a rounded apex, longer than parameres (Fig.
|
|
<figureCitation id="00884A8F44F8EB93DE0EC1D3751FEDBC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8A</figureCitation>
|
|
). Median lobe in lateral view quite flattened, curved basally, straight proximally and slightly bent downward distally, narrowing apically (Fig.
|
|
<figureCitation id="914C40B7C007ED294DD8F745C5C35F11" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8B</figureCitation>
|
|
). Basal bulb small, narrow, sub-parallel and slightly widened distally in dorsal view (Fig.
|
|
<figureCitation id="F493A0441484675D03BCC8D57AFACAB1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8A</figureCitation>
|
|
), while elongate and widened basally in lateral view (Fig.
|
|
<figureCitation id="8740618E61C3C44BFDD22A4AB2162342" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8B</figureCitation>
|
|
). Tegmen wide from above (Fig.
|
|
<figureCitation id="C2A2F89632160652C050D3F0D8C4577A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8A</figureCitation>
|
|
), in the shape of a ring around basal bulb (Fig.
|
|
<figureCitation id="ED9D5BC3A5763C59E9FFA6917CB77512" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8B</figureCitation>
|
|
). Parameres elongate, slender, arcuate, sub-apically curved exteriorly, each with a moderately widened rounded apex in dorsal view (Fig.
|
|
<figureCitation id="75DBA1936F04B44A11A418CC612B674E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8A</figureCitation>
|
|
), while almost straight, sub-parallel in lateral view (Fig.
|
|
<figureCitation id="C765BB6B1A80F067A4BD3B874E84B681" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8B</figureCitation>
|
|
). Each paramera bearing four apical close-set setae, two of which longer, while two shorter (Fig.
|
|
<figureCitation id="3F09BCCBDF0A8F726A2A207AA4704CF6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8A, B</figureCitation>
|
|
). No copulatory piece observed within inner sac (Fig.
|
|
<figureCitation id="2265EF93DCB6AAB63DB7130293408D24" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8A, B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="2C8DA5AB049F24755516E4B0AE811AAB" doi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21" start="Figure 8" startId="F8">
|
|
<paragraph id="5E8CAF2D02C4A87A938F20BE056FF420" pageId="0" pageNumber="21">
|
|
<emphasis id="A1618F46B3DE5B2DB2B764DA68B22659" bold="true" pageId="0" pageNumber="21">Figure 8.</emphasis>
|
|
Bright-field images of certain morphological traits of topotype male (
|
|
<emphasis id="9CF519C6CDEA7878989217900726A39C" bold="true" pageId="0" pageNumber="21">A, B</emphasis>
|
|
) and topotype female (
|
|
<emphasis id="8260CEC2CC12447D8E27D2FC068650C7" bold="true" pageId="0" pageNumber="21">C, D</emphasis>
|
|
) of
|
|
<taxonomicName id="A18338B18D27EDE1EB697E8A12489942" authorityName="Mueller" authorityYear="1901" class="Insecta" family="Leiodidae" genus="Spelaeobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spelaeobates (Spelaeobates) novaki" order="Coleoptera" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="novaki" subGenus="Spelaeobates">Spelaeobates (Spelaeobates) novaki</taxonomicName>
|
|
from the
|
|
<normalizedToken id="CF92D50CAEB901C249ABD4C172A951AD" originalValue="Strašna">Strasna</normalizedToken>
|
|
<normalizedToken id="841545E479EF167721CD5B4975360C06" originalValue="Peć">Pec</normalizedToken>
|
|
Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia
|
|
<emphasis id="6B094F0D317EC6923BC9A0CE21F449A5" bold="true" pageId="0" pageNumber="21">A</emphasis>
|
|
aedeagus, dorsal view
|
|
<emphasis id="276ECB086553971DB0D350B9EE9BFA06" bold="true" pageId="0" pageNumber="21">B</emphasis>
|
|
aedeagus, lateral view
|
|
<emphasis id="0D9E4406E98728ED9F987D96A7B57105" bold="true" pageId="0" pageNumber="21">C</emphasis>
|
|
spermatheca, lateral view
|
|
<emphasis id="E821440AFF5D4D3BD89E40918A2F237D" bold="true" pageId="0" pageNumber="21">D</emphasis>
|
|
abdominal ventrite VIII. Scale bars: 0.2 mm (
|
|
<emphasis id="0B5A2FD5E3BA77D46B77D82204EB78A3" bold="true" pageId="0" pageNumber="21">A, B, D</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="26AA98E335C4C40A9AB9E0AF9BD58B90" bold="true" pageId="0" pageNumber="21">C</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="D94E101703755C3288EBAD173F4A54B0" pageId="0" pageNumber="21">
|
|
<emphasis id="700646DEFDFD361E118950A0A4D4AA15" bold="true" italics="true" pageId="0" pageNumber="21">Female genitalia</emphasis>
|
|
: Spermatheca small, chitinised, straight basally, curved sub-apically, spherical both basally and apically (Fig.
|
|
<figureCitation id="EF67CA637FD836527CC6D4E081307A16" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8C</figureCitation>
|
|
). Gonostyli short, straight, moderately widened, gradually narrowing distally, pointed apically. Each gonostylus carrying one long apical seta.
|
|
</paragraph>
|
|
<paragraph id="4EFA6949C1036FD0B8F526BD9AF9194E" pageId="0" pageNumber="21">
|
|
<emphasis id="1485B724969445763EFE14E595D63927" bold="true" italics="true" pageId="0" pageNumber="21">Male abdominal sternite IX (urite)</emphasis>
|
|
: Small, narrowing apically, sub-triangular.
|
|
</paragraph>
|
|
<paragraph id="B716709D0DC2727086812AB632DBE46B" pageId="0" pageNumber="21">
|
|
<emphasis id="01F41A20C97F139C6D181F72DA8E8A80" bold="true" italics="true" pageId="0" pageNumber="21">Female abdominal ventrite VIII</emphasis>
|
|
: Small, transverse, with no anterior process, hairy, especially posteriorly, slightly bilobed distally (Fig.
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<figureCitation id="CB3178F6AFEBEBF402A54E52299608CE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bright-field images of certain morphological traits of topotype male (A, B) and topotype female (C, D) of Spelaeobates (Spelaeobates) novaki from the Strasna Pec Cave, village of Savar, island of Dugi Otok, northern Dalmatia, Croatia A aedeagus, dorsal view B aedeagus, lateral view C spermatheca, lateral view D abdominal ventrite VIII. Scale bars: 0.2 mm (A, B, D); 0.1 mm (C)." figureDoi="10.3897/subtbiol.46.104548.figure8" httpUri="https://binary.pensoft.net/fig/896587" pageId="0" pageNumber="21">8D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="E42BC3DE94D899FF46953D7B1F6DB817" pageId="0" pageNumber="21" type="distribution">
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<paragraph id="B2323742D5E339618E77FD59AB17521A" pageId="0" pageNumber="21">Geographic distribution.</paragraph>
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<paragraph id="5145F89989FED073BA2F71F86705C1C0" pageId="0" pageNumber="21">
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This species inhabits two caves located on two northern Dalmatian islands - the
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<normalizedToken id="63FD7C2AE3E1AFBAF3AB5DCFE2FE0952" originalValue="Strašna">Strasna</normalizedToken>
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<normalizedToken id="4DB599F71E1487F28B645FC87B8DFDCB" originalValue="Peć">Pec</normalizedToken>
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Cave (island of Dugi Otok) and the Jezero Cave (island of
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<normalizedToken id="BE8B80C11073D5030979C0A3187AB0A7" originalValue="Iž">Iz</normalizedToken>
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) (
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<bibRefCitation id="BD48A7419BCDEE2E34937408427C5E6C" author="Pretner, E" journalOrPublisher="Krs Jugoslavije" pageId="0" pageNumber="21" pagination="101 - 239" refId="B28" refString="Pretner, E, 1973. Koleopteroloska fauna pecina i jama Hrvatske s historijskim pregledom istrazivanja. Krs Jugoslavije 8: 101 - 239" title="Koleopteroloska fauna pecina i jama Hrvatske s historijskim pregledom istrazivanja." volume="8" year="1973">Pretner 1973</bibRefCitation>
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). It is possible that it also inhabits other subterranean sites on the same and neighbouring islands.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |