544 lines
67 KiB
XML
544 lines
67 KiB
XML
<document id="2497ABF2A9FCBCDA3A64AC42AED15DE6" ID="10.11646/zootaxa.4121.5.2" ID-CLB-Dataset="38883" ID-DOI="10.11646/zootaxa.4121.5.2" ID-GBIF-Dataset="e50114c8-81ec-4aff-b985-104934aa648e" ID-ISSN="1175-5326" ID-Zenodo-Dep="257253" ID-ZooBank="5BFF6D6F-A325-4200-A99A-ED6D01BBE381" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465798714731" checkinUser="plazi" docAuthor="Köhler, Gunther, Rodríguez Bobadilla, Marcos J. & Hedges, S. Blair" docDate="2016" docId="943987CBCE37F23FA0F2F9EEFC7FF952" docLanguage="en" docName="zootaxa.4121.5.2.pdf" docOrigin="Zootaxa 4121 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leiocephalus sixtoi Köhler, Bobadilla & Hedges, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="527" masterDocId="6800FFB3CE35F235A065FF96FFF7FFE6" masterDocTitle="A new dune-dwelling lizard of the genus Leiocephalus (Iguania, Leiocephalidae) from the Dominican Republic" masterLastPageNumber="532" masterPageNumber="517" pageNumber="519" updateTime="1698662822167" updateUser="ExternalLinkService">
|
||
<mods:mods id="2C6DA6C40C1AF444F5F960001DC2A964" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="E80F26256D1E274A50E363953EEEFC99">
|
||
<mods:title id="82CFF86D53D102B2A709F9FF036575CC">A new dune-dwelling lizard of the genus Leiocephalus (Iguania, Leiocephalidae) from the Dominican Republic</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="05C612BCB13CFE61FB94148250D348EE" type="personal">
|
||
<mods:role id="B55827C5EC74CA949C60F5053AD50EDC">
|
||
<mods:roleTerm id="54C615A5512F7ED238F10C2094B43CD0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E0E3C34CD3AAE76282564FA6157F50AC">Köhler, Gunther</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0D19A2EC286B230E8A3C113E0E2FCD6D" type="personal">
|
||
<mods:role id="9CE6537B174D6B5ECFDB2073AB1C25BA">
|
||
<mods:roleTerm id="5C2F7C9BCEB2C5E3B06F11545320A3A2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="49A50A4D51C9675FAD3E354E2B769727">Rodríguez Bobadilla, Marcos J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D33FFB582CA91F876F21830CC1DC9F37" type="personal">
|
||
<mods:role id="456FC5423A30ED121574B5BFF58CB47B">
|
||
<mods:roleTerm id="B09257BAE4114081FFA19C5F9B229747">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="EA38FE8BF710FF2708E0379B3DBFA186">Hedges, S. Blair</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="ABE32048BFD443BCE7319EB8A831C936">text</mods:typeOfResource>
|
||
<mods:relatedItem id="0C3C90424224D6E8A568210AF23F0DFC" type="host">
|
||
<mods:titleInfo id="ABB1EE9EA7E60FC67458F5C47141FAD4">
|
||
<mods:title id="CE1FCCB32756DBEF8C2D02DBBAFFC1C3">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="3355E08267D2E9B311DBADFF365E8CA6">
|
||
<mods:date id="15797D92B0FDAFB506AEEAD1FEC48EB8">2016</mods:date>
|
||
<mods:detail id="EDF4C0FD14228A0EC77944E0BA2945D3" type="volume">
|
||
<mods:number id="7215C5EA9B647CD02DB0F16F2B0C11C4">4121</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="FA3BC059376D0AEAD6EA04AAA8EECF8B" type="issue">
|
||
<mods:number id="4D8F76349A26CF9C3528CEA3319DDA21">5</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="80AD763A33BADE58B9FB52ACD5618286" unit="page">
|
||
<mods:start id="813FBBA9E2D24299893C5C798999896A">517</mods:start>
|
||
<mods:end id="74823A73D74FEDA22A1F2777404C38A1">532</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="12BC0E52541B069D29953723F0143A31">journal article</mods:classification>
|
||
<mods:identifier id="FB3B8D7F276DC6AC956CAFDA35B09C33" type="CLB-Dataset">38883</mods:identifier>
|
||
<mods:identifier id="4AB3D95AF0D3AB1BB1B7AB96B599E987" type="DOI">10.11646/zootaxa.4121.5.2</mods:identifier>
|
||
<mods:identifier id="33A1BF7BB2C3114BC544D9C73A53713F" type="GBIF-Dataset">e50114c8-81ec-4aff-b985-104934aa648e</mods:identifier>
|
||
<mods:identifier id="BBD7394023EBBA44B527C6D6748CC173" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="060734C8F9B0A3D343782A31EFFDF257" type="Zenodo-Dep">257253</mods:identifier>
|
||
<mods:identifier id="E648EB963E5DDCEF718FEDB99F60AF83" type="ZooBank">5BFF6D6F-A325-4200-A99A-ED6D01BBE381</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="943987CBCE37F23FA0F2F9EEFC7FF952" ID-DOI="http://doi.org/10.5281/zenodo.6059205" ID-GBIF-Taxon="120827680" ID-Zenodo-Dep="6059205" LSID="urn:lsid:plazi:treatment:943987CBCE37F23FA0F2F9EEFC7FF952" httpUri="http://treatment.plazi.org/id/943987CBCE37F23FA0F2F9EEFC7FF952" lastPageId="10" lastPageNumber="527" pageId="2" pageNumber="519">
|
||
<subSubSection id="548A6556CE37F237A0F2F9EEFE4BF952" pageId="2" pageNumber="519" type="nomenclature">
|
||
<paragraph id="1C2F36DDCE37F237A0F2F9EEFE11F974" blockId="2.[151,486,1656,1716]" box="[151,486,1656,1682]" pageId="2" pageNumber="519">
|
||
<heading id="476781B1CE37F237A0F2F9EEFE11F974" bold="true" box="[151,486,1656,1682]" fontSize="11" level="1" pageId="2" pageNumber="519" reason="1">
|
||
<emphasis id="2EE4EACFCE37F237A0F2F9EEFE11F974" bold="true" box="[151,486,1656,1682]" pageId="2" pageNumber="519">
|
||
<taxonomicName id="DB904D5ECE37F237A0F2F9EEFE77F974" box="[151,384,1656,1682]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="519" phylum="Chordata" rank="species" species="sixtoi" status="sp. nov.">
|
||
<emphasis id="2EE4EACFCE37F237A0F2F9EEFE77F974" bold="true" box="[151,384,1656,1682]" italics="true" pageId="2" pageNumber="519">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="35D757B4CE37F237A1EDF9EEFE11F974" box="[392,486,1656,1682]" pageId="2" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE37F237A0F2F90DFE4BF952" blockId="2.[151,486,1656,1716]" box="[151,444,1691,1716]" pageId="2" pageNumber="519">
|
||
<figureCitation id="84AB2A58CE37F237A0F2F90DFF14F952" box="[151,227,1691,1716]" captionStart="FIGURE 2" captionStartId="3.[151,250,1753,1775]" captionTargetBox="[150,1436,193,1730]" captionTargetId="figure@3.[150,1436,193,1732]" captionTargetPageId="3" captionText="FIGURE 2. Cranial osteology of Leiocephalus sixtoi (SMF 99151): (a) dorsal view; (b) lateral view; (c) ventral view of upper cranium; (d) right mandible in lateral view. Cranial osteology of Leiocephalus schreibersii (SMF 26278): (e) dorsal view; (f) lateral view; (g) ventral view of upper cranium; (h) right mandible in lateral view. Scale bars equal 1.0 mm. Photographs by G. K." httpUri="https://zenodo.org/record/257255/files/figure.png" pageId="2" pageNumber="519">Figs. 2</figureCitation>
|
||
a–d, 3a,c, 4a,c, 5–8
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="548A6556CE37F237A0F2F975FB66F856" pageId="2" pageNumber="519" type="materials_examined">
|
||
<paragraph id="1C2F36DDCE37F237A0F2F975FE39F8A2" blockId="2.[151,1437,1763,1968]" pageId="2" pageNumber="519">
|
||
<emphasis id="2EE4EACFCE37F237A0F2F975FEF3F91A" bold="true" box="[151,260,1763,1788]" pageId="2" pageNumber="519">
|
||
<typeStatus id="C32B887FCE37F237A0F2F975FEF3F91A" box="[151,260,1763,1788]" pageId="2" pageNumber="519" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="7A81AE18CE37F237A177F972FEBDF91A" LSID="urn:lsid:biocol.org:col:34838" box="[274,330,1764,1788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="519">SMF</collectionCode>
|
||
99143, an adult male from Dunas de Baní, near the village of Las Salinas, 18.21285°, -70.53131°,
|
||
<quantity id="DB689B38CE37F237A0F2F891FF32F8F9" box="[151,197,1799,1824]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="519" unit="m" value="5.0">5 m</quantity>
|
||
asl, Provincia de Peravia,
|
||
<collectingCountry id="6487764DCE37F237A19BF891FD1AF8C6" box="[510,749,1799,1824]" name="Dominican Republic" pageId="2" pageNumber="519">Dominican Republic</collectingCountry>
|
||
, collected
|
||
<date id="682E101DCE37F237A30FF891FBECF8F9" box="[874,1051,1799,1824]" pageId="2" pageNumber="519" value="2014-03-22">22 March 2014</date>
|
||
by Marcos Rodriguez Bobadilla. Field tag number GK-5061.
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE37F237A0A2F8D9FB66F856" blockId="2.[151,1437,1763,1968]" pageId="2" pageNumber="519">
|
||
<emphasis id="2EE4EACFCE37F237A0A2F8D9FEC8F88E" bold="true" box="[199,319,1871,1896]" pageId="2" pageNumber="519">
|
||
<typeStatus id="C32B887FCE37F237A0A2F8D9FEC8F88E" box="[199,319,1871,1896]" pageId="2" pageNumber="519" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="7A81AE18CE37F237A12BF8D9FE71F881" LSID="urn:lsid:biocol.org:col:34838" box="[334,390,1871,1895]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="519">SMF</collectionCode>
|
||
99144–51, same collecting data as the
|
||
<typeStatus id="C32B887FCE37F237A328F8D9FC58F88E" box="[845,943,1871,1896]" pageId="2" pageNumber="519" type="holotype">holotype</typeStatus>
|
||
;
|
||
<collectionCode id="7A81AE18CE37F237A3A5F8D9FBCEF881" box="[960,1081,1871,1895]" httpUri="http://grbio.org/cool/736g-snne" name="Museo Nacional de Historia Natural, Santo Domingo" pageId="2" pageNumber="519">MNHNSD</collectionCode>
|
||
23.2193–202, same locality as the
|
||
<typeStatus id="C32B887FCE37F237A0A4F8E5FED4F86A" box="[193,291,1907,1932]" pageId="2" pageNumber="519" type="holotype">holotype</typeStatus>
|
||
but collected
|
||
<date id="682E101DCE37F237A1A6F8E5FDB0F86A" box="[451,583,1907,1932]" pageId="2" pageNumber="519" value="2012-07-06">6 July 2012</date>
|
||
by Marcos Rodríguez, Luis M. Díaz, and Nils Navarro.
|
||
<collectionCode id="7A81AE18CE37F237A4A2F8E2FAB6F86A" box="[1223,1345,1908,1932]" httpUri="http://grbio.org/cool/736g-snne" name="Museo Nacional de Historia Natural, Santo Domingo" pageId="2" pageNumber="519">MNHNSD</collectionCode>
|
||
23.202,
|
||
<collectionCode id="7A81AE18CE37F237A0F2F801FF38F849" LSID="urn:lsid:biocol.org:col:34838" box="[151,207,1943,1967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="519">SMF</collectionCode>
|
||
99145, 99147, 99150–51 are adult males, the remaining specimens are adult females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="48EF6655CE36F236A0F2F94FFF34F8AD" httpUri="https://zenodo.org/record/257255/files/figure.png" pageId="3" pageNumber="520" targetBox="[150,1436,193,1730]" targetPageId="3">
|
||
<paragraph id="1C2F36DDCE36F236A0F2F94FFF34F8AD" blockId="3.[151,1436,1753,1867]" pageId="3" pageNumber="520">
|
||
<emphasis id="2EE4EACFCE36F236A0F2F94FFEE4F909" bold="true" box="[151,275,1753,1775]" pageId="3" pageNumber="520">FIGURE 2.</emphasis>
|
||
Cranial osteology of
|
||
<taxonomicName id="DB904D5ECE36F236A18AF94FFD44F909" box="[495,691,1753,1775]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE36F236A18AF94FFD44F909" box="[495,691,1753,1775]" italics="true" pageId="3" pageNumber="520">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99151): (a) dorsal view; (b) lateral view; (c) ventral view of upper cranium; (d) right mandible in lateral view. Cranial osteology of
|
||
<taxonomicName id="DB904D5ECE36F236A351F96EFBCFF8E8" box="[820,1080,1784,1806]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE36F236A351F96EFBCFF8E8" box="[820,1080,1784,1806]" italics="true" pageId="3" pageNumber="520">Leiocephalus schreibersii</emphasis>
|
||
</taxonomicName>
|
||
(SMF 26278): (e) dorsal view; (f) lateral view; (g) ventral view of upper cranium; (h) right mandible in lateral view. Scale bars equal 1.0 mm. Photographs by G.K.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="548A6556CE36F231A0A2F8E4FBB1FD16" lastPageId="4" lastPageNumber="521" pageId="3" pageNumber="520" type="diagnosis">
|
||
<paragraph id="1C2F36DDCE36F236A0A2F8E4FA6BF834" blockId="3.[151,1436,1906,2003]" pageId="3" pageNumber="520">
|
||
<emphasis id="2EE4EACFCE36F236A0A2F8E4FEB6F86D" bold="true" box="[199,321,1906,1931]" pageId="3" pageNumber="520">Diagnosis.</emphasis>
|
||
<taxonomicName id="DB904D5ECE36F236A12EF8E5FDD0F86C" box="[331,551,1907,1931]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE36F236A12EF8E5FDD0F86C" box="[331,551,1907,1931]" italics="true" pageId="3" pageNumber="520">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
differs from all other congeners except
|
||
<emphasis id="2EE4EACFCE36F236A39FF8E5FE23F849" italics="true" pageId="3" pageNumber="520">
|
||
<taxonomicName id="DB904D5ECE36F236A39FF8E5FB57F86D" box="[1018,1184,1907,1931]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="schreibersii">L. schreibersii</taxonomicName>
|
||
,
|
||
<taxonomicName id="DB904D5ECE36F236A4CBF8E5FA80F86D" box="[1198,1399,1907,1931]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="melanochlorus">L. melanochlorus</taxonomicName>
|
||
,
|
||
<taxonomicName id="DB904D5ECE36F236A5E3F8E5FEBDF849" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="psammodromus">L. psammodromus</taxonomicName>
|
||
,
|
||
<taxonomicName id="DB904D5ECE36F236A130F80EFE27F849" box="[341,464,1944,1967]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="inaguae">L. inaguae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="DB904D5ECE36F236A26EF80EFD61F849" box="[523,662,1944,1967]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="macropus">
|
||
<emphasis id="2EE4EACFCE36F236A26EF80EFD61F849" box="[523,662,1944,1967]" italics="true" pageId="3" pageNumber="520">L. macropus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a lateral fold. It differs from
|
||
<emphasis id="2EE4EACFCE36F236A4D6F80EFEB9F835" italics="true" pageId="3" pageNumber="520">
|
||
<taxonomicName id="DB904D5ECE36F236A4D6F80EFA8DF849" box="[1203,1402,1943,1967]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="melanochlorus">L. melanochlorus</taxonomicName>
|
||
,
|
||
<taxonomicName id="DB904D5ECE36F236A5E0F80EFEBDF835" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="psammodromus">L. psammodromus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="DB904D5ECE36F236A1E2F82DFDE5F834" box="[391,530,1979,2002]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="macropus">
|
||
<emphasis id="2EE4EACFCE36F236A1E2F82DFDE5F834" box="[391,530,1979,2002]" italics="true" pageId="3" pageNumber="520">L. macropus</emphasis>
|
||
</taxonomicName>
|
||
in having 3 internasals (vs.
|
||
<quantity id="DB689B38CE36F236A331F82DFC77F835" box="[852,896,1978,2003]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="520" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="DB904D5ECE36F236A3EDF82DFBBAF835" box="[904,1101,1979,2003]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="melanochlorus">
|
||
<emphasis id="2EE4EACFCE36F236A3EDF82DFBBAF835" box="[904,1101,1979,2003]" italics="true" pageId="3" pageNumber="520">L. melanochlorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="2EE4EACFCE36F236A4E3F82DFAEEF834" box="[1158,1305,1979,2002]" italics="true" pageId="3" pageNumber="520">
|
||
<taxonomicName id="DB904D5ECE36F236A4E3F82DFAE3F834" box="[1158,1300,1979,2002]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="520" phylum="Chordata" rank="species" species="macropus">L. macropus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<quantity id="DB689B38CE36F236A534F82DFA8AF835" box="[1361,1405,1978,2003]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="520" unit="in" value="4.0">4 in</quantity>
|
||
<emphasis id="2EE4EACFCE36F236A5E0F82DFA6BF834" box="[1413,1436,1979,2002]" italics="true" pageId="3" pageNumber="520">L.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE31F231A0F2FF0EFBB1FD16" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="521">
|
||
<taxonomicName id="DB904D5ECE31F231A0F2FF0EFEB0FF56" box="[151,327,152,176]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="psammodromus">
|
||
<emphasis id="2EE4EACFCE31F231A0F2FF0EFEB0FF56" box="[151,327,152,176]" italics="true" pageId="4" pageNumber="521">psammodromus</emphasis>
|
||
</taxonomicName>
|
||
, respectively). It further differs from
|
||
<taxonomicName id="DB904D5ECE31F231A28DFF0FFC5CFF56" box="[744,939,152,176]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="melanochlorus">
|
||
<emphasis id="2EE4EACFCE31F231A28DFF0FFC5CFF56" box="[744,939,152,176]" italics="true" pageId="4" pageNumber="521">L. melanochlorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DB904D5ECE31F231A385FF0FFB5AFF56" box="[992,1197,152,176]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="psammodromus">
|
||
<emphasis id="2EE4EACFCE31F231A385FF0FFB5AFF56" box="[992,1197,152,176]" italics="true" pageId="4" pageNumber="521">L. psammodromus</emphasis>
|
||
</taxonomicName>
|
||
in having 4 lorilabial scales anterior to the enlarged subocular (vs. 5–6).
|
||
<taxonomicName id="DB904D5ECE31F231A28BFF2AFC3CFF32" box="[750,971,188,212]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE31F231A28BFF2AFC3CFF32" box="[750,971,188,212]" italics="true" pageId="4" pageNumber="521">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="DB904D5ECE31F231A40FFF2BFB12FF32" box="[1130,1253,189,212]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="inaguae">
|
||
<emphasis id="2EE4EACFCE31F231A40FFF2BFB12FF32" box="[1130,1253,189,212]" italics="true" pageId="4" pageNumber="521">L. inaguae</emphasis>
|
||
</taxonomicName>
|
||
in having a Ushaped bony parietal table (vs. V-shaped in
|
||
<taxonomicName id="DB904D5ECE31F231A2F4FF77FCFCFF11" box="[657,779,224,248]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="inaguae">
|
||
<emphasis id="2EE4EACFCE31F231A2F4FF77FCFCFF11" box="[657,779,224,248]" italics="true" pageId="4" pageNumber="521">L. inaguae</emphasis>
|
||
</taxonomicName>
|
||
), 3 or 4 enlarged postcloacal scales in males (vs.
|
||
<quantity id="DB689B38CE31F231A52AFF76FA8BFF1E" box="[1359,1404,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="521" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="DB904D5ECE31F231A5E0FF77FF06FEFA" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="inaguae">
|
||
<emphasis id="2EE4EACFCE31F231A5E0FF77FF06FEFA" italics="true" pageId="4" pageNumber="521">L. inaguae</emphasis>
|
||
</taxonomicName>
|
||
), most scales on the snout posterior to the internasal scales rugose to keeled (vs. smooth in
|
||
<taxonomicName id="DB904D5ECE31F231A577FE93FA7CFEFA" box="[1298,1419,261,284]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="inaguae">
|
||
<emphasis id="2EE4EACFCE31F231A577FE93FA7CFEFA" box="[1298,1419,261,284]" italics="true" pageId="4" pageNumber="521">L. inaguae</emphasis>
|
||
</taxonomicName>
|
||
), and a patternless throat in males, spots on the throat in females (vs. throat with dark streaks and bars in males and females of
|
||
<taxonomicName id="DB904D5ECE31F231A171FEDBFE7BFE82" box="[276,396,333,356]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="inaguae">
|
||
<emphasis id="2EE4EACFCE31F231A171FEDBFE7BFE82" box="[276,396,333,356]" italics="true" pageId="4" pageNumber="521">L. inaguae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="DB904D5ECE31F231A1C1FEDAFD8AFE82" box="[420,637,332,356]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE31F231A1C1FEDAFD8AFE82" box="[420,637,332,356]" italics="true" pageId="4" pageNumber="521">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="DB904D5ECE31F231A376FEDBFC43FE82" box="[787,948,332,356]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A376FEDBFC43FE82" box="[787,948,332,356]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
in having the scales of the lateral fold only slightly smaller than adjacent scales (vs. scales of lateral fold distinctly smaller than adjacent scales;
|
||
<figureCitation id="84AB2A58CE31F231A543FEF9FA9BFE6E" box="[1318,1388,367,392]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1965]" captionTargetBox="[150,1436,1057,1921]" captionTargetId="figure@5.[150,1436,1057,1921]" captionTargetPageId="5" captionText="FIGURE 4. Region of the lateral fold in (a) Leiocephalus sixtoi (SMF 99143) and (b) L. schreibersii (SMF 26132). Lateral tail surface in (c) Leiocephalus sixtoi (SMF 99143) and (d) L. schreibersii (SMF 26133). Scale bars equal 1.0 mm. Photographs by G. K." httpUri="https://zenodo.org/record/257257/files/figure.png" pageId="4" pageNumber="521">Fig. 4</figureCitation>
|
||
a,b), having prominent caudal crest scales in adult males (vs. caudal crest scales of moderate size, even in very large males in
|
||
<taxonomicName id="DB904D5ECE31F231A099FE2FFE6AFE36" box="[252,413,440,464]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A099FE2FFE6AFE36" box="[252,413,440,464]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="84AB2A58CE31F231A1C8FE21FE06FE36" box="[429,497,439,464]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1965]" captionTargetBox="[150,1436,1057,1921]" captionTargetId="figure@5.[150,1436,1057,1921]" captionTargetPageId="5" captionText="FIGURE 4. Region of the lateral fold in (a) Leiocephalus sixtoi (SMF 99143) and (b) L. schreibersii (SMF 26132). Lateral tail surface in (c) Leiocephalus sixtoi (SMF 99143) and (d) L. schreibersii (SMF 26133). Scale bars equal 1.0 mm. Photographs by G. K." httpUri="https://zenodo.org/record/257257/files/figure.png" pageId="4" pageNumber="521">Fig. 4</figureCitation>
|
||
c,d), a pattern of dark gray bars on a grayish brown background in the region above the lateral body fold (vs. dense turquoise blue mottling with heavy suffusion of red pigment in
|
||
<taxonomicName id="DB904D5ECE31F231A4B1FE4BFA80FE12" box="[1236,1399,476,500]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A4B1FE4BFA80FE12" box="[1236,1399,476,500]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
), a darker dorsal ground color (vs. paler in
|
||
<taxonomicName id="DB904D5ECE31F231A233FD97FD00FDFE" box="[598,759,512,536]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A233FD97FD00FDFE" box="[598,759,512,536]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
), and a red iris in adult males (vs. pale grayish blue in adult male
|
||
<taxonomicName id="DB904D5ECE31F231A0B3FDB3FE8FFDDA" box="[214,376,548,572]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A0B3FDB3FE8FFDDA" box="[214,376,548,572]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="DB904D5ECE31F231A1F4FDB2FD9BFDDA" box="[401,620,548,572]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE31F231A1F4FDB2FD9BFDDA" box="[401,620,548,572]" italics="true" pageId="4" pageNumber="521">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
differs further from
|
||
<taxonomicName id="DB904D5ECE31F231A33CFDB3FC0BFDDA" box="[857,1020,548,572]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A33CFDB3FC0BFDDA" box="[857,1020,548,572]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
in several osteological characters as follows: in
|
||
<taxonomicName id="DB904D5ECE31F231A173FDDFFE87FDB9" box="[278,368,584,608]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE31F231A173FDDFFE87FDB9" box="[278,368,584,608]" italics="true" pageId="4" pageNumber="521">L. sixtoi</emphasis>
|
||
</taxonomicName>
|
||
the nasal process of the premaxilla reaches to mid-level of the bony external nares (vs. to level of posterior margin of the bony external nares in
|
||
<taxonomicName id="DB904D5ECE31F231A2B1FDFBFC80FD62" box="[724,887,620,644]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A2B1FDFBFC80FD62" box="[724,887,620,644]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
), lacking a constriction at the base of the nasal process of the premaxilla (vs. such a constriction is present in
|
||
<taxonomicName id="DB904D5ECE31F231A3EEFD07FBC6FD4E" box="[907,1073,656,680]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A3EEFD07FBC6FD4E" box="[907,1073,656,680]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
), and having a reduced nasalprefrontal contact that leaves the nasal processes of the frontal bone exposed (vs. nasal and prefrontal bones in contact, thereby obscuring the nasal processes of the frontal bone in
|
||
<taxonomicName id="DB904D5ECE31F231A3F0FD4FFBC1FD16" box="[917,1078,728,752]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A3F0FD4FFBC1FD16" box="[917,1078,728,752]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="48EF6655CE31F231A0F2F90DFAACF908" httpUri="https://zenodo.org/record/257256/files/figure.png" pageId="4" pageNumber="521" targetBox="[150,1436,805,1669]" targetPageId="4">
|
||
<paragraph id="1C2F36DDCE31F231A0F2F90DFAACF908" blockId="4.[151,1437,1691,1774]" pageId="4" pageNumber="521">
|
||
<emphasis id="2EE4EACFCE31F231A0F2F90DFEE2F957" bold="true" box="[151,277,1691,1713]" pageId="4" pageNumber="521">FIGURE 3.</emphasis>
|
||
Shape and extension of the nasal process of the premaxilla in (a)
|
||
<taxonomicName id="DB904D5ECE31F231A3B5F90DFB61F957" box="[976,1174,1691,1713]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE31F231A3B5F90DFB61F957" box="[976,1174,1691,1713]" italics="true" pageId="4" pageNumber="521">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99151) and (b)
|
||
<taxonomicName id="DB904D5ECE31F231A5E2F90AFEF9F929" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A5E2F90AFEF9F929" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
(SMF 26278). Shape and extension of the nasal processes of the frontal bone in (c)
|
||
<taxonomicName id="DB904D5ECE31F231A40CF92CFB4BF929" box="[1129,1212,1722,1743]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE31F231A40CF92CFB4BF929" box="[1129,1212,1722,1743]" italics="true" pageId="4" pageNumber="521">L. sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99151) and (d)
|
||
<taxonomicName id="DB904D5ECE31F231A0F2F94FFEDFF908" box="[151,296,1752,1774]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="521" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE31F231A0F2F94FFEDFF908" box="[151,296,1752,1774]" italics="true" pageId="4" pageNumber="521">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
(SMF 26278). Relevant sutures highlighted with red lines. Scale bars equal 1.0 mm. Photographs by G.K.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="548A6556CE31F23CA0A2F883FB9AFE13" lastPageId="9" lastPageNumber="526" pageId="4" pageNumber="521" type="description">
|
||
<paragraph id="1C2F36DDCE31F230A0A2F883FADBFC0A" blockId="4.[151,1436,1813,2018]" lastBlockId="5.[151,1437,151,1005]" lastPageId="5" lastPageNumber="522" pageId="4" pageNumber="521">
|
||
<emphasis id="2EE4EACFCE31F231A0A2F883FD4BF8C8" bold="true" box="[199,700,1813,1838]" pageId="4" pageNumber="521">
|
||
Description of the
|
||
<typeStatus id="C32B887FCE31F231A1C1F883FDFCF8C8" box="[420,523,1813,1838]" pageId="4" pageNumber="521" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="84AB2A58CE31F231A27CF883FD9EF8C8" box="[537,617,1813,1838]" captionStart="FIGURE 5" captionStartId="6.[151,250,1521,1543]" captionTargetBox="[150,1435,193,1499]" captionTargetId="figure@6.[150,1436,193,1500]" captionTargetPageId="6" captionText="FIGURE 5. Holotype of Leiocephalus sixtoi (SMF 99143): (a) lateral view; (b) dorsal view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (a – c) and 1.0 mm in (d – f), respectively. Photographs by G. K." httpUri="https://zenodo.org/record/257258/files/figure.png" pageId="4" pageNumber="521">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="84AB2A58CE31F231A2C1F883FD44F8C8" box="[676,691,1813,1838]" captionStart="FIGURE 6" captionStartId="7.[151,250,1976,1998]" captionTargetBox="[371,1221,193,1951]" captionTargetId="figure@7.[371,1222,193,1951]" captionTargetPageId="7" captionText="FIGURE 6. Holotype of Leiocephalus sixtoi (SMF 99143): (a) dorsal region (b) flank; (c) midventer. Scale bars equal 1.0 mm. Photographs by G. K." httpUri="https://zenodo.org/record/257259/files/figure.png" pageId="4" pageNumber="521">6</figureCitation>
|
||
)
|
||
</emphasis>
|
||
. Adult male, as indicated by partly everted hemipenes; SVL 87.0 mm; tail length 125.0 mm (incomplete); tail strongly compressed in cross section, tail height
|
||
<quantity id="DB689B38CE31F231A4D9F8ACFAD4F8B7" box="[1212,1315,1850,1874]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="4" pageNumber="521" unit="mm" value="12.3">12.3 mm</quantity>
|
||
, tail width
|
||
<quantity id="DB689B38CE31F231A0F2F8CBFF07F893" box="[151,240,1885,1910]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="521" unit="mm" value="5.6">5.6 mm</quantity>
|
||
; axilla to groin distance
|
||
<quantity id="DB689B38CE31F231A267F8CBFD9EF893" box="[514,617,1885,1910]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="4" pageNumber="521" unit="mm" value="38.5">38.5 mm</quantity>
|
||
; head length 20.0 mm, head length/SVL ratio 0.23; snout length
|
||
<quantity id="DB689B38CE31F231A55AF8C8FA6FF893" box="[1343,1432,1886,1910]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="4" pageNumber="521" unit="mm" value="8.2">8.2 mm</quantity>
|
||
; head width 14.0 mm; head height
|
||
<quantity id="DB689B38CE31F231A27FF814FD76F87F" box="[538,641,1922,1946]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="4" pageNumber="521" unit="mm" value="11.1">11.1 mm</quantity>
|
||
; shank length
|
||
<quantity id="DB689B38CE31F231A341F814FC7CF87F" box="[804,907,1922,1946]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="4" pageNumber="521" unit="mm" value="23.1">23.1 mm</quantity>
|
||
, shank length/head length ratio 1.16; scales on parietal and occipital regions large, rugose to wrinkled, juxtaposed; parietal eye visible; supraoculars in a single row, multicarinate, total number 6; circumorbital row complete, separating supraoculars from supraorbital semicircles; 3 overlapping superciliaries, posteriormost longest; canthals two; anteriormost canthal separated from nasal by a small scale; internasals 3, 2 in contact with rostral; postrostrals, much longer than wide; supralabials 4 to point level of mideye, supralabial series slightly sloping down after this point; 8 (right) / 6 (left) lorilabials in a single row; 4 (right) / 3 (left) lorilabials anterior to enlarged subocular; infralabials 5 to point below center of eye; 4 loreals in two rows; 1 preocular; single, large, elongate subocular scale, in contact with one supralabial; lateral temporals keeled, subimbricate; transverse count of gular scales 29 between tympanic openings; all gulars cycloid, smooth, imbricate, some slightly pointed; first pair of sublabials in contact with anteriormost infralabials; first pair of postmentals in contact medially; interparietal scale very elongate, subtriangular; parietal scales large, almost rectangular, inner pair almost equal to size of outer one; 2–3 scales between parietals and dorsal scales; dorsal scales of neck and body large, strongly keeled, imbricate, mucronate, keels mostly orientated longitudanally, with the vertebral scales forming a serrated crest that continues along body onto tail; lateral scales of neck somewhat heterogeneous in size and shape, mostly small, weakly keeled and subimbricate, granular at base lateral neck folds; lateral scales of body mostly strongly keeled, imbricate, mucronate, keels oriented obliquely upwards, becoming smaller and less keeled toward axilla and toward groin, respectively; scales of longitudinal fold mostly large, keeled, imbricate, micronate, except at lower base of fold where they are slightly smaller and less distinctly keeled; scales around midbody 68; vertebral crest scales from occiput to level of posterior margin of thigh 77; ventrals smooth, cycloid, imbricate, distinctly larger than dorsals; preauricular fringe present with 4–5 projecting scales; antegular, antehumeral, gular, longitudinal, postauricular, supra-auricular, and transverse antegular neck folds present; dorsal scales of forelimbs and hindlimbs keeled, mucronate, imbricate; scales on ventral surfaces of limbs smooth to slightly keeled, imbricate, cycloid to pointed; lamellae on Finger
|
||
<date id="682E101DCE30F230A39FFCD3FB94FCBB" box="[1018,1123,836,861]" pageId="5" pageNumber="522" value="1916-04-15">IV 15/16</date>
|
||
; lamellae on Toe
|
||
<date id="682E101DCE30F230A54BFCD3FA6FFCBB" box="[1326,1432,836,861]" pageId="5" pageNumber="522" value="1925-04-25">IV 25/25</date>
|
||
; lateral caudal scales strongly keeled, mucronate, imbricate, dorsal caudal scales greatly enlarged, forming a high, serrated crest; ventral caudal scales in a double series of pointed scales, smooth on base of tail, becoming strongly keeled on middle and distal portions; posthumeral mite pocket present as 2–3 vertical folds [
|
||
<typeStatus id="C32B887FCE30F230A4A8FC26FAFFFC2E" box="[1229,1288,944,968]" pageId="5" pageNumber="522">Type</typeStatus>
|
||
1 of Torres-
|
||
<bibRefCitation id="78014B2CCE30F230A0F2FC42FEBBFC0A" author="Carvajal" box="[151,332,980,1005]" pageId="5" pageNumber="522" refString="Torres-Carvajal, O. (2007) A taxonomic revision of South American Stenocercus (Squamata: Iguania) lizards. Herpetological Monographs, 21, 76 - 178. http: // dx. doi. org / 10.1655 / 06 - 001.1" type="journal article" year="2007">Carvajal (2007)</bibRefCitation>
|
||
]; postfemoral mite pocket absent; a pair of enlarged, elongate postcloacal scales present.
|
||
</paragraph>
|
||
<caption id="48EF6655CE30F230A0F2F801FF34F80C" httpUri="https://zenodo.org/record/257257/files/figure.png" pageId="5" pageNumber="522" targetBox="[150,1436,1057,1921]" targetPageId="5">
|
||
<paragraph id="1C2F36DDCE30F230A0F2F801FF34F80C" blockId="5.[151,1435,1943,2026]" pageId="5" pageNumber="522">
|
||
<emphasis id="2EE4EACFCE30F230A0F2F801FEE5F84B" bold="true" box="[151,274,1943,1965]" pageId="5" pageNumber="522">FIGURE 4.</emphasis>
|
||
Region of the lateral fold in (a)
|
||
<taxonomicName id="DB904D5ECE30F230A23CF801FCEAF84B" box="[601,797,1943,1965]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="522" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE30F230A23CF801FCEAF84B" box="[601,797,1943,1965]" italics="true" pageId="5" pageNumber="522">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99143) and (b)
|
||
<taxonomicName id="DB904D5ECE30F230A464F80EFB66F84B" box="[1025,1169,1943,1965]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="522" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE30F230A464F80EFB66F84B" box="[1025,1169,1943,1965]" italics="true" pageId="5" pageNumber="522">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
(SMF 26132). Lateral tail surface in (c)
|
||
<taxonomicName id="DB904D5ECE30F230A146F823FE10F82A" box="[291,487,1973,1996]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="522" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE30F230A146F823FE10F82A" box="[291,487,1973,1996]" italics="true" pageId="5" pageNumber="522">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99143) and (d)
|
||
<taxonomicName id="DB904D5ECE30F230A2AEF821FCAAF82D" box="[715,861,1973,1996]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="522" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE30F230A2AEF821FCAAF82D" box="[715,861,1973,1996]" italics="true" pageId="5" pageNumber="522">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
(SMF 26133). Scale bars equal 1.0 mm. Photographs by G.K.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="48EF6655CE33F233A0F2FA67FE9DF9A2" httpUri="https://zenodo.org/record/257258/files/figure.png" pageId="6" pageNumber="523" targetBox="[150,1435,193,1499]" targetPageId="6">
|
||
<paragraph id="1C2F36DDCE33F233A0F2FA67FE9DF9A2" blockId="6.[151,1436,1521,1604]" pageId="6" pageNumber="523">
|
||
<emphasis id="2EE4EACFCE33F233A0F2FA67FEE4F9E1" bold="true" box="[151,275,1521,1543]" pageId="6" pageNumber="523">FIGURE 5.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="DB904D5ECE33F233A1F9FA67FD97F9E1" box="[412,608,1521,1543]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="523" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE33F233A1F9FA67FD97F9E1" box="[412,608,1521,1543]" italics="true" pageId="6" pageNumber="523">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99143): (a) lateral view; (b) dorsal view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (a–c) and 1.0 mm in (d–f), respectively. Photographs by G.K.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C2F36DDCE33F233A0A2F9FAFA8DF82E" blockId="6.[151,1436,1644,1992]" pageId="6" pageNumber="523">
|
||
Coloration in life was as follows (
|
||
<figureCitation id="84AB2A58CE33F233A22CF9FAFD7AF962" box="[585,653,1644,1669]" captionStart="FIGURE 7" captionStartId="8.[151,250,1521,1543]" captionTargetBox="[150,1435,193,1499]" captionTargetId="figure@8.[150,1436,193,1500]" captionTargetPageId="8" captionText="FIGURE 7. Leiocephalus sixtoi in life. (a, b) SMF 99143 (holotype, male); (c) SMF 99150 (male); (d) SMF 99144 (female); (e) SMF 99150 (male); (d) SMF 99146 (female). Photographs by G. K." httpUri="https://zenodo.org/record/257260/files/figure.png" pageId="6" pageNumber="523">Fig. 7</figureCitation>
|
||
a, b): Dorsal surface of head Olive Brown (278) with Pale Cinnamon (55) suffusions; lateral surfaces of head Army Brown (46) above, grading into Straw Yellow (53) in labial regions, and with Olive Brown (278) and Greenish Glaucous (271) splotches and suffusions; ventral surface of head Smoky White (261); dorsal surface of neck Drab (19) suffused with Salmon Color (58); dorsal suface of body Drab (19) with Raw Umber (22) longitudinal dorsolateral bands with Creme Color (12) speckles; lateral surfaces of body with Sepia (286) transverse bands and Light Bluish Gray (288) speckles; ventral surface of body Smoke Gray (266) with transverse bands composed of Medium Paris White (140) and Pratt’s Rufous (72) scales; dorsal surfaces of limbs Drab (19) with Olive Horn Color (16) speckles; dorsal surface of tail Drab (19) with Light Bluish Gray (288) and Creme Color (12) speckles; caudal crest scales Light Bluish Gray (288) with Glaucous (289) suffusions; ventral surface of tail Medium Fawn Color (257), heavily suffused with Carmine (64); iris Orange-Rufous (56).
|
||
</paragraph>
|
||
<caption id="48EF6655CE32F232A0F2F82EFE9DF80A" httpUri="https://zenodo.org/record/257259/files/figure.png" pageId="7" pageNumber="524" targetBox="[371,1221,193,1951]" targetPageId="7">
|
||
<paragraph id="1C2F36DDCE32F232A0F2F82EFE9DF80A" blockId="7.[151,1436,1976,2028]" pageId="7" pageNumber="524">
|
||
<emphasis id="2EE4EACFCE32F232A0F2F82EFEE5F828" bold="true" box="[151,274,1976,1998]" pageId="7" pageNumber="524">FIGURE 6.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="DB904D5ECE32F232A1FDF82EFDABF828" box="[408,604,1976,1998]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="524" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE32F232A1FDF82EFDABF828" box="[408,604,1976,1998]" italics="true" pageId="7" pageNumber="524">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99143): (a) dorsal region (b) flank; (c) midventer. Scale bars equal 1.0 mm. Photographs by G.K.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="48EF6655CE3DF23DA0F2FA67FCCAF9C3" httpUri="https://zenodo.org/record/257260/files/figure.png" pageId="8" pageNumber="525" targetBox="[150,1435,193,1499]" targetPageId="8">
|
||
<paragraph id="1C2F36DDCE3DF23DA0F2FA67FCCAF9C3" blockId="8.[151,1436,1521,1573]" pageId="8" pageNumber="525">
|
||
<emphasis id="2EE4EACFCE3DF23DA0F2FA67FEE5F9E1" bold="true" box="[151,274,1521,1543]" pageId="8" pageNumber="525">FIGURE 7.</emphasis>
|
||
<taxonomicName id="DB904D5ECE3DF23DA17DFA67FE2CF9E1" box="[280,475,1521,1543]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="525" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE3DF23DA17DFA67FE2CF9E1" box="[280,475,1521,1543]" italics="true" pageId="8" pageNumber="525">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
in life. (a,b) SMF 99143 (holotype, male); (c) SMF 99150 (male); (d) SMF 99144 (female); (e) SMF 99150 (male); (d) SMF 99146 (female). Photographs by G.K.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C2F36DDCE3DF23DA0A2F9C4FD36F8A5" blockId="8.[151,1437,1618,2003]" pageId="8" pageNumber="525">
|
||
<emphasis id="2EE4EACFCE3DF23DA0A2F9C4FEC2F98D" bold="true" box="[199,309,1618,1643]" pageId="8" pageNumber="525">Variation</emphasis>
|
||
. The
|
||
<typeStatus id="C32B887FCE3DF23DA119F9C5FE10F98D" box="[380,487,1619,1643]" pageId="8" pageNumber="525" type="paratype">paratypes</typeStatus>
|
||
agree well with the
|
||
<typeStatus id="C32B887FCE3DF23DA2B1F9C4FCC1F98D" box="[724,822,1618,1643]" pageId="8" pageNumber="525" type="holotype">holotype</typeStatus>
|
||
in general appearance, morphometrics, and scalation. The largest male (SMF 99151) in the
|
||
<typeStatus id="C32B887FCE3DF23DA236F9E1FD44F969" box="[595,691,1655,1679]" pageId="8" pageNumber="525" type="paratype">paratype</typeStatus>
|
||
series has a SVL of
|
||
<quantity id="DB689B38CE3DF23DA3CBF9E1FBF5F968" box="[942,1026,1655,1679]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="8" pageNumber="525" unit="mm" value="88.0">88 mm</quantity>
|
||
, the largest female is
|
||
<quantity id="DB689B38CE3DF23DA561F9E1FAA1F968" box="[1284,1366,1655,1679]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="525" unit="mm" value="70.0">70 mm</quantity>
|
||
SVL. Variation in selected scalation characters: SAM in 68–79 (74.2 ± 3.4); median dorsal crest scales 72–80 (75.1 ± 2.5); number of IN is
|
||
<quantity id="DB689B38CE3DF23DA1EFF929FE43F931" box="[394,436,1726,1751]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="525" unit="in" value="3.0">3 in</quantity>
|
||
all, except SMF 99147 that has 4 IN; number of lorilabials anterior to enlarged subocular 3–4 (3.7 ± 0.4); number of SPL 4; number of IFL 5–6 (5.2 ± 0.4), number supraoculars 6–8 (7.0 ± 0.8); and subdigital lamellae on Phalanges II–IV of Toe
|
||
<date id="682E101DCE3DF23DA2DEF891FCDEF8F9" box="[699,809,1798,1823]" pageId="8" pageNumber="525" value="1925-04-28">IV 25–28</date>
|
||
(26.3 ± 0.9). Only one specimen (SMF 99147) has a complete tail with a ratio tail length/SVL of 1.64.
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE3DF23CA0A2F8D8FAFFFE6E" blockId="8.[151,1437,1618,2003]" lastBlockId="9.[151,1436,151,501]" lastPageId="9" lastPageNumber="526" pageId="8" pageNumber="525">
|
||
All specimens in the
|
||
<typeStatus id="C32B887FCE3DF23DA1D1F8D9FE13F881" box="[436,484,1871,1895]" pageId="8" pageNumber="525">type</typeStatus>
|
||
series have a distinct series of dark grayish brown blotches on the lateral surfaces of the body (see
|
||
<figureCitation id="84AB2A58CE3DF23DA152F8E4FE8BF86D" box="[311,380,1906,1931]" captionStart="FIGURE 7" captionStartId="8.[151,250,1521,1543]" captionTargetBox="[150,1435,193,1499]" captionTargetId="figure@8.[150,1436,193,1500]" captionTargetPageId="8" captionText="FIGURE 7. Leiocephalus sixtoi in life. (a, b) SMF 99143 (holotype, male); (c) SMF 99150 (male); (d) SMF 99144 (female); (e) SMF 99150 (male); (d) SMF 99146 (female). Photographs by G. K." httpUri="https://zenodo.org/record/257260/files/figure.png" pageId="8" pageNumber="525">Fig. 7</figureCitation>
|
||
). The ventral surface of head and neck is mostly immaculate in males, but with distinct dark spots in females. Coloration in life of an adult female (SMF 99144;
|
||
<figureCitation id="84AB2A58CE3DF23DA3ECF800FC35F849" box="[905,962,1942,1967]" captionStart="FIGURE 7" captionStartId="8.[151,250,1521,1543]" captionTargetBox="[150,1435,193,1499]" captionTargetId="figure@8.[150,1436,193,1500]" captionTargetPageId="8" captionText="FIGURE 7. Leiocephalus sixtoi in life. (a, b) SMF 99143 (holotype, male); (c) SMF 99150 (male); (d) SMF 99144 (female); (e) SMF 99150 (male); (d) SMF 99146 (female). Photographs by G. K." httpUri="https://zenodo.org/record/257260/files/figure.png" pageId="8" pageNumber="525">Fig 7</figureCitation>
|
||
d) was recorded as follows: Dorsal surface of head Grayish Horn Color (286) with a suffusion of Sepia (286), especially on supraoculars; lateral surfaces of head Drab (19) above, grading into Smoky White (261) in labial regions, and with Jet Black (300) blotches; ventral surface of head Smoky White (261); dorsal suface of Smoke Gray (267) with Sepia (286) paravertebral and lateral blotches with Pale Buff (1) speckles; ventral surface of body Smoky White (261) with transverse rows of Sepia (286) blotches; dorsal and lateral surfaces of limbs Grayish Horn Color (286) with Sepia (286) splotches and Pale Buff (1) speckles; ventral surfaces of limbs Smoky White (261); dorsal surface of tail Smoke Gray (267) anteriorly, grading into Ground Cinnamon (270) with Sepia (286) chevrons, bordered distally by Smoky White (261); ventral surface of tail Smoky White (261), heavily suffused with Pale Pinkish Buff (3); iris Warm Sepia (40).
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE3CF23CA0A2FE02FB9AFE13" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="526">
|
||
The almost completely everted hemipenis of SMF 99150 (
|
||
<figureCitation id="84AB2A58CE3CF23CA30FFE02FC58FE4A" box="[874,943,404,429]" captionStart="FIGURE 8" captionStartId="9.[151,250,1634,1656]" captionTargetBox="[151,1435,553,1612]" captionTargetId="figure@9.[151,1435,553,1613]" captionTargetPageId="9" captionText="FIGURE 8. Hemipenes of Leiocephalus sixtoi (SMF 99150): sulcate view left, asulcate view right. Scale bar equals 1.0 mm. Photographs by G. K." httpUri="https://zenodo.org/record/257261/files/figure.png" pageId="9" pageNumber="526">Fig. 8</figureCitation>
|
||
) is a stout, medium-sized, slightly bilobed organ; sulcus spermaticus bordered by well-developed sulcal lips and opening into a broad concave area at base of apex; distal region of apex covered by large calyces, lower apex has 4–5 large flounces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="48EF6655CE3CF23CA0F2F9F4FE9DF971" httpUri="https://zenodo.org/record/257261/files/figure.png" pageId="9" pageNumber="526" targetBox="[151,1435,553,1612]" targetPageId="9">
|
||
<paragraph id="1C2F36DDCE3CF23CA0F2F9F4FE9DF971" blockId="9.[151,1437,1634,1687]" pageId="9" pageNumber="526">
|
||
<emphasis id="2EE4EACFCE3CF23CA0F2F9F4FEE4F99E" bold="true" box="[151,275,1634,1656]" pageId="9" pageNumber="526">FIGURE 8.</emphasis>
|
||
Hemipenes of
|
||
<taxonomicName id="DB904D5ECE3CF23CA1D5F9F4FD81F99E" box="[432,630,1634,1656]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="526" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE3CF23CA1D5F9F4FD81F99E" box="[432,630,1634,1656]" italics="true" pageId="9" pageNumber="526">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 99150): sulcate view left, asulcate view right. Scale bar equals 1.0 mm. Photographs by G.K.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="548A6556CE3CF23FA0A2F928FC7FF952" lastPageId="10" lastPageNumber="527" pageId="9" pageNumber="526" type="etymology">
|
||
<paragraph id="1C2F36DDCE3CF23CA0A2F928FD30F881" blockId="9.[151,1436,1726,2003]" pageId="9" pageNumber="526">
|
||
<emphasis id="2EE4EACFCE3CF23CA0A2F928FEB0F931" bold="true" box="[199,327,1726,1751]" pageId="9" pageNumber="526">Etymology</emphasis>
|
||
. The name
|
||
<taxonomicName id="DB904D5ECE3CF23CA1B1F956FDE6F931" box="[468,529,1728,1751]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="526" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE3CF23CA1B1F956FDE6F931" box="[468,529,1728,1751]" italics="true" pageId="9" pageNumber="526">sixtoi</emphasis>
|
||
</taxonomicName>
|
||
is a patronym honoring our friend and colleague Sixto Incháustegui, who has contributed substantially to our knowledge of Hispaniolan amphibians and reptiles. Sixto is professor at the Universidad Autónoma de Santo
|
||
<collectingCountry id="6487764DCE3CF23CA279F890FD7EF8F9" box="[540,649,1798,1823]" name="Dominican Republic" pageId="9" pageNumber="526">Domingo</collectingCountry>
|
||
,
|
||
<collectingCountry id="6487764DCE3CF23CA2FDF890FC70F8F9" box="[664,903,1798,1823]" name="Dominican Republic" pageId="9" pageNumber="526">Dominican Republic</collectingCountry>
|
||
, where he teaches herpetology and history of biology. For more than 35 years, he has been involved as a major player in biological research and nature conservation on a national and international level.
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE3CF23CA0A2F8E4FEBAF849" blockId="9.[151,1436,1726,2003]" pageId="9" pageNumber="526">
|
||
<emphasis id="2EE4EACFCE3CF23CA0A2F8E4FE1BF86D" bold="true" box="[199,492,1906,1931]" pageId="9" pageNumber="526">Geographic distribution</emphasis>
|
||
.
|
||
<taxonomicName id="DB904D5ECE3CF23CA265F8E5FD29F86C" box="[512,734,1907,1931]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="526" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE3CF23CA265F8E5FD29F86C" box="[512,734,1907,1931]" italics="true" pageId="9" pageNumber="526">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
is presently only known from the surroundings of its
|
||
<typeStatus id="C32B887FCE3CF23CA509F8E5FA6BF86D" box="[1388,1436,1907,1931]" pageId="9" pageNumber="526">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="84AB2A58CE3CF23CA09FF800FEC9F849" box="[250,318,1942,1967]" captionStart="FIGURE 9" captionStartId="10.[151,250,1379,1401]" captionTargetBox="[156,1432,522,1353]" captionTargetId="figure@10.[150,1436,517,1358]" captionTargetPageId="10" captionText="FIGURE 9. Map indicating the geographic distribution of Leiocephalus sixtoi (yellow dot) and its closest relatives, L. inagua (blue) and L. schreibersii (red)." httpUri="https://zenodo.org/record/257262/files/figure.png" pageId="9" pageNumber="526">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE3CF23CA0A2F82CFA6BF835" blockId="9.[151,1436,1726,2003]" box="[199,1436,1978,2003]" pageId="9" pageNumber="526">
|
||
<emphasis id="2EE4EACFCE3CF23CA0A2F82CFE32F835" bold="true" box="[199,453,1978,2003]" pageId="9" pageNumber="526">Natural history notes</emphasis>
|
||
. The habitat at the
|
||
<typeStatus id="C32B887FCE3CF23CA2C4F82DFD26F835" box="[673,721,1979,2003]" pageId="9" pageNumber="526">type</typeStatus>
|
||
locality is sandy dunes with low Tropical Dry Forest (
|
||
<figureCitation id="84AB2A58CE3CF23CA55EF82CFA78F835" box="[1339,1423,1978,2003]" captionStart="FIGURE 10" captionStartId="11.[151,250,1353,1375]" captionTargetBox="[150,1436,483,1330]" captionTargetId="figure@11.[150,1436,483,1331]" captionTargetPageId="11" captionText="FIGURE 10. Habitat of Leiocephalus sixtoi at Dunas de Baní, near the village Las Salinas, Provincia de Peravia, Dominican Republic. Photograph by Eladio Fernández." httpUri="https://zenodo.org/record/257263/files/figure.png" pageId="9" pageNumber="526">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1C2F36DDCE3FF23FA0F2FF01FCC8FE36" blockId="10.[151,1437,151,464]" pageId="10" pageNumber="527">
|
||
This dune area streches about
|
||
<quantity id="DB689B38CE3FF23FA18DFF01FDD8FF56" box="[488,559,151,176]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="527" unit="km" value="15.0">15 km</quantity>
|
||
in length with a maximum width of about
|
||
<quantity id="DB689B38CE3FF23FA468FF0EFBBEFF56" box="[1037,1097,151,176]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="527" unit="km" value="3.0">3 km</quantity>
|
||
. The accumulation of sand for the formation of the dunes is a product of the waves from the Caribbean Sea and a more or less steady wind blowing mostly from the same direction. These dunes reach a maximum height of about
|
||
<specimenCount id="0A96FD54CE3FF23FA4E9FF49FB3AFF11" box="[1164,1229,223,248]" pageId="10" pageNumber="527" type="generic">
|
||
<quantity id="DB689B38CE3FF23FA4E9FF49FB3EFF11" box="[1164,1225,223,248]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="527" unit="m" value="35.0">35 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
The vegetation in the vicinity of the dunes consists mostly of trees (
|
||
<emphasis id="2EE4EACFCE3FF23FA2B6FE92FA6BFEFA" box="[723,1436,260,284]" italics="true" pageId="10" pageNumber="527">
|
||
<taxonomicName id="DB904D5ECE3FF23FA2B6FE92FBAEFEFA" authority="Coccoloba" authorityName="Coccoloba" box="[723,1113,260,284]" class="Magnoliopsida" family="Simaroubaceae" genus="Simarouba" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="527" phylum="Tracheophyta" rank="species" species="berteroana">Simarouba berteroana, Coccoloba</taxonomicName>
|
||
uvifera, Ziziphus reticulata,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="DB904D5ECE3FF23FA0A8FEBEFE4BFEA6" box="[205,444,296,320]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="527" phylum="Tracheophyta" rank="species" species="macracantha">
|
||
<emphasis id="2EE4EACFCE3FF23FA0A8FEBEFE4BFEA6" box="[205,444,296,320]" italics="true" pageId="10" pageNumber="527">Acacia macracantha</emphasis>
|
||
</taxonomicName>
|
||
), cacti (
|
||
<emphasis id="2EE4EACFCE3FF23FA243FEBEFB3BFE82" italics="true" pageId="10" pageNumber="527">
|
||
<taxonomicName id="DB904D5ECE3FF23FA243FEBEFC69FED9" authority="Opuntia" authorityName="Opuntia" box="[550,926,296,320]" class="Magnoliopsida" family="Cactaceae" genus="Consolea" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="527" phylum="Tracheophyta" rank="species" species="moniliformis">Consolea moniliformis, Opuntia</taxonomicName>
|
||
dillenii, Cylindropuntia caribaea, Opuntia antillana, Harrisia nashii, Lemairecereus hystrix, Pilocereus polygonus, Melocactus lemairei,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="DB904D5ECE3FF23FA56DFEDAFF0DFE6E" class="Magnoliopsida" family="Cactaceae" genus="Mammillaria" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="527" phylum="Tracheophyta" rank="species" species="prolifera">
|
||
<emphasis id="2EE4EACFCE3FF23FA56DFEDAFF0DFE6E" italics="true" pageId="10" pageNumber="527">Mammillaria prolifera</emphasis>
|
||
</taxonomicName>
|
||
), shrubs, and grasses (Anomymous 2016). The
|
||
<typeStatus id="C32B887FCE3FF23FA34AFEE6FCA8FE6E" box="[815,863,368,392]" pageId="10" pageNumber="527">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="DB904D5ECE3FF23FA468FEE6FB1DFE61" box="[1037,1258,368,392]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE3FF23FA468FEE6FB1DFE61" box="[1037,1258,368,392]" italics="true" pageId="10" pageNumber="527">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
were collected during the morning hours on and under rocks in a grassy area.
|
||
<collectingCountry id="6487764DCE3FF23FA33BFE02FC8AFE4A" box="[862,893,404,428]" name="American Samoa" pageId="10" pageNumber="527">As</collectingCountry>
|
||
the other species in this genus, this is a diurnal, terrestrial lizard that feeds mostly on a variety of arthropods.
|
||
</paragraph>
|
||
<caption id="48EF6655CE3FF23FA0F2FAF5FE23FA7E" httpUri="https://zenodo.org/record/257262/files/figure.png" pageId="10" pageNumber="527" targetBox="[156,1432,522,1353]" targetPageId="10">
|
||
<paragraph id="1C2F36DDCE3FF23FA0F2FAF5FE23FA7E" blockId="10.[151,1436,1379,1432]" pageId="10" pageNumber="527">
|
||
<emphasis id="2EE4EACFCE3FF23FA0F2FAF5FEE5FA9F" bold="true" box="[151,274,1379,1401]" pageId="10" pageNumber="527">FIGURE 9.</emphasis>
|
||
Map indicating the geographic distribution of
|
||
<taxonomicName id="DB904D5ECE3FF23FA294FAF5FC41FA9F" box="[753,950,1379,1401]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="sixtoi">
|
||
<emphasis id="2EE4EACFCE3FF23FA294FAF5FC41FA9F" box="[753,950,1379,1401]" italics="true" pageId="10" pageNumber="527">Leiocephalus sixtoi</emphasis>
|
||
</taxonomicName>
|
||
(yellow dot) and its closest relatives,
|
||
<taxonomicName id="DB904D5ECE3FF23FA55FFAF2FA6BFA9F" box="[1338,1436,1380,1401]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="inagua">
|
||
<emphasis id="2EE4EACFCE3FF23FA55FFAF2FA6BFA9F" box="[1338,1436,1380,1401]" italics="true" pageId="10" pageNumber="527">L. inagua</emphasis>
|
||
</taxonomicName>
|
||
(blue) and
|
||
<taxonomicName id="DB904D5ECE3FF23FA161FA15FE61FA7E" box="[260,406,1410,1432]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE3FF23FA161FA15FE61FA7E" box="[260,406,1410,1432]" italics="true" pageId="10" pageNumber="527">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
(red).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1C2F36DDCE3FF23FA0A2FA52FC7FF952" blockId="10.[151,1437,1476,1717]" pageId="10" pageNumber="527">
|
||
<emphasis id="2EE4EACFCE3FF23FA0A2FA52FB67FA3B" bold="true" box="[199,1168,1476,1501]" pageId="10" pageNumber="527">
|
||
Designation of a
|
||
<typeStatus id="C32B887FCE3FF23FA1CBFA52FDFCFA3B" box="[430,523,1476,1501]" pageId="10" pageNumber="527" type="neotype">neotype</typeStatus>
|
||
for
|
||
<taxonomicName id="DB904D5ECE3FF23FA237FA52FB7CFA3B" authority="Gravenhorst 1838" authorityName="Gravenhorst" authorityYear="1838" box="[594,1163,1476,1501]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE3FF23FA237FA52FC73FA3B" bold="true" box="[594,900,1476,1501]" italics="true" pageId="10" pageNumber="527">Leiocephalus schreibersii</emphasis>
|
||
(
|
||
<bibRefCitation id="78014B2CCE3FF23FA3F8FA52FB74FA3B" author="Gravenhorst" box="[925,1155,1476,1501]" pageId="10" pageNumber="527" refString="Gravenhorst, J. L. C. (1838) Beitrage zur genaueren Kenntniss einiger Eidechsengattungen. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum. Verhandlungen der Kaiserlich Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher, 18, 712 - 784. [1837]" type="journal article" year="1838">Gravenhorst 1838</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
According to
|
||
<bibRefCitation id="78014B2CCE3FF23FA534FA52FEFFF9E7" author="Pregill" pageId="10" pageNumber="527" refString="Pregill, G. K. (1992) Systematics of the West Indian lizard genus Leiocephalus (Squamata: Iguania: Tropiduridae). Miscellaneous Publication, University of Kansas, Museum of Natural History, 84, 1 - 169." type="journal article" year="1992">Pregill (1992:51)</bibRefCitation>
|
||
, the
|
||
<typeStatus id="C32B887FCE3FF23FA121FA7FFE83F9E7" box="[324,372,1513,1537]" pageId="10" pageNumber="527">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="DB904D5ECE3FF23FA26CFA7EFCDBF9E6" box="[521,812,1512,1536]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE3FF23FA26CFA7EFCDBF9E6" box="[521,812,1512,1536]" italics="true" pageId="10" pageNumber="527">Leiocephalus schreibersii</emphasis>
|
||
</taxonomicName>
|
||
was unlocatable, apparently deposited in the Breslau Museum (= now Museum of Natural History at Wrocław University). Upon request by G.K., the present curator of herpetology at the Museum of Natural History at Wrocław University, Andrzej Jablonski, informed us that the
|
||
<typeStatus id="C32B887FCE3FF23FA509F9A7FA6BF9AF" box="[1388,1436,1585,1609]" pageId="10" pageNumber="527">type</typeStatus>
|
||
of
|
||
<taxonomicName id="DB904D5ECE3FF23FA0D3F9C3FEA0F98B" box="[182,343,1621,1645]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE3FF23FA0D3F9C3FEA0F98B" box="[182,343,1621,1645]" italics="true" pageId="10" pageNumber="527">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
is not among the specimens in the collection under his care. Therefore, we consider it to be lost. We designate SMF 26228 (
|
||
<figureCitation id="84AB2A58CE3FF23FA1A8F9EEFDDFF977" box="[461,552,1656,1681]" captionStart="FIGURE 11" captionStartId="12.[151,250,1565,1587]" captionTargetBox="[150,1435,215,1521]" captionTargetId="figure@12.[150,1436,215,1522]" captionTargetPageId="12" captionText="FIGURE 11. Neotype of Leiocephalus schreibersii (SMF 26228): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (a – c) and 1.0 mm in (d – f), respectively. Photographs by G. K." httpUri="https://zenodo.org/record/257264/files/figure.png" pageId="10" pageNumber="527">Figs. 11</figureCitation>
|
||
and
|
||
<figureCitation id="84AB2A58CE3FF23FA204F9EFFD77F977" box="[609,640,1657,1681]" captionStart="FIGURE 12" captionStartId="13.[151,250,1974,1996]" captionTargetBox="[365,1216,193,1951]" captionTargetId="figure@13.[365,1216,193,1951]" captionTargetPageId="13" captionText="FIGURE 12. Neotype of Leiocephalus schreibersii (SMF 26228): (a) dorsal region; (b) flank; (c) midventer. Scale bars equal 1.0 mm. Photographs by G. K." httpUri="https://zenodo.org/record/257265/files/figure.png" pageId="10" pageNumber="527">12</figureCitation>
|
||
), an adult male from Saint Marc, Province Artibonite,
|
||
<collectingCountry id="6487764DCE3FF23FA495F9EEFADCF977" box="[1264,1323,1656,1681]" name="Haiti" pageId="10" pageNumber="527">Haiti</collectingCountry>
|
||
, collected
|
||
<date id="682E101DCE3FF23FA0F2F90BFEC3F953" box="[151,308,1692,1717]" pageId="10" pageNumber="527" value="1939-04-19">19 April 1939</date>
|
||
by Robert Mertens, as the
|
||
<typeStatus id="C32B887FCE3FF23FA207F90BFD4CF953" box="[610,699,1693,1717]" pageId="10" pageNumber="527" type="neotype">neotype</typeStatus>
|
||
of
|
||
<taxonomicName id="DB904D5ECE3FF23FA285F90BFC76F953" box="[736,897,1693,1717]" class="Reptilia" family="Leiocephalidae" genus="Leiocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="schreibersii">
|
||
<emphasis id="2EE4EACFCE3FF23FA285F90BFC76F953" box="[736,897,1693,1717]" italics="true" pageId="10" pageNumber="527">L. schreibersii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |