121 lines
11 KiB
XML
121 lines
11 KiB
XML
<document ENCODING="UTF8" ID-DOI="http://doi.org/10.5281/zenodo.6137116" ID-GBIF-Dataset="9aa410be-7687-45e7-a820-065d5b3281e8" ID-GBIF-Taxon="100060568" ID-HNS-Pub="27210" ID-Zenodo-Dep="6137116" checkinTime="1400715470232" checkinUser="donat" docAuthor="Bolton, Barry & Fisher, Brian" docDate="2014" docId="6DEBF1209DC6A4D6807E5E9965600BC7" docLanguage="en" docName="27210_4.xml" docOrigin="Zootaxa 3791 (1)" docTitle="Royidris shuckardi Forel, comb. n." docType="treatment" docVersion="10" lastPageNumber="66" masterDocId="94F02DEB689E84D4B189E46A16A44076" masterDocTitle="The Madagascan endemic myrmicine ants related to Eutetramorium (Hymenoptera: Formicidae): taxonomy of the genera Eutetramorium Emery, Malagidris nom. n., Myrmisaraka gen. n., Royidris gen. n., and Vitsika gen. n .." masterLastPageNumber="99" masterPageNumber="1" pageNumber="65" updateTime="1645179377042" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Madagascan endemic myrmicine ants related to Eutetramorium (Hymenoptera: Formicidae): taxonomy of the genera Eutetramorium Emery, Malagidris nom. n., Myrmisaraka gen. n., Royidris gen. n., and Vitsika gen. n ..</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bolton, Barry</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>3791</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>99</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="HNS-Pub">27210</mods:identifier>
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<mods:identifier type="DOI">10.11646/zootaxa.3791.1.1</mods:identifier>
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<mods:identifier type="ZooBank">1C6E3E9C-E1F4-4A0E-9481-B59B817C8355</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6137116" ID-GBIF-Taxon="100060568" ID-Zenodo-Dep="6137116" LSID="urn:lsid:plazi:treatment:6DEBF1209DC6A4D6807E5E9965600BC7" httpUri="http://treatment.plazi.org/id/6DEBF1209DC6A4D6807E5E9965600BC7" lastPageNumber="66" pageNumber="65">
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<subSubSection type="nomenclature">
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<paragraph pageId="64" pageNumber="65" type="mainText">
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<taxonomicName authority="Forel" authorityName="Forel" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="shuckardi">
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Royidris shuckardi (
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<authority>Forel</authority>
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)
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</taxonomicName>
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<taxonomicNameLabel>comb. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection type="description">
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<paragraph pageId="64" pageNumber="65" type="mainText">(Figs 75-77, Map 147)</paragraph>
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</subSubSection>
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<subSubSection type="reference_group">
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<paragraph pageId="64" pageNumber="65" type="mainText">
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<taxonomicName authority="Forel" authorityName="Forel" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="shuckardi">Monomorium shuckardi Forel</taxonomicName>
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, 1895: 251. Holotype queen, MADAGASCAR: Moramanga (Sikora) (MHNG) [examined].
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<taxonomicNameLabel>Comb. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection type="discussion">
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<paragraph lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="mainText">
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QUEEN (gyne). Alate when virgin. Among the few known queens of the
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<taxonomicName class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="gravipuncta">gravipuncta</taxonomicName>
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complex,
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<taxonomicName class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="shuckardi">shuckardi</taxonomicName>
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is isolated because with its head in full-face view the eyes are located in front of the midlength of the sides, and the
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<pageBreakToken pageId="65" pageNumber="66" start="start">sides</pageBreakToken>
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behind the eyes appear long and shallowly convex, convergent posteriorly through long, shallow curves, and with the posterior margin short and indented medially. In the worker-associated queen of
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<taxonomicName class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="peregrina">peregrina</taxonomicName>
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, and the two unassociated queens that are known in the complex, the eyes are at, or very close to, the midlength, and the sides behind the eyes round through narrow curves into a broad posterior margin that is shallowly concave across its width.
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</paragraph>
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</subSubSection>
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<subSubSection type="multiple">
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<caption pageId="65" pageNumber="66">
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<paragraph pageId="65" pageNumber="66" type="caption">
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FIGURES 75-77. Lateral, full face and dorsal view of body.
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="shuckardi">Royidris shuckardi</taxonomicName>
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holotype queen CASENT0010761.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection type="description">
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<paragraph pageId="65" pageNumber="66" type="mainText">WORKER and MALE: unknown.</paragraph>
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</subSubSection>
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<subSubSection type="diagnosis">
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<paragraph pageId="65" pageNumber="66" type="mainText">
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This species is known only from the holotype queen. While it certainly belongs in
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">Royidris</taxonomicName>
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, and because of its 4-segmented antennal club is referable to the
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="notorthotenes">notorthotenes</taxonomicName>
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group, it can not be matched with any other queen assigned to that group. In addition, as pointed out by Heterick (2006), its type-locality, Moramanga, is in Toamasina Province, hundreds of kilometers away from the range of, and with vegetation and conditions very different from, that inhabited by other species now in
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">Royidris</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection type="discussion">
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<paragraph pageId="65" pageNumber="66" type="mainText">
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Heterick (2006) associated the isolated
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="shuckardi">shuckardi</taxonomicName>
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holotype queen with workers of what is now considered a complex of three worker-based species,
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="gravipuncta">gravipuncta</taxonomicName>
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,
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="longiseta">longiseta</taxonomicName>
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, and the more recently discovered
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="peregrina">peregrina</taxonomicName>
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. He grouped the first two species together with the isolated queen, and regarded them as a single species, to which he applied the name
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="shuckardi">shuckardi</taxonomicName>
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. However, there is no evidence for this association. None of the few queens associated with, or assumed to belong to the same group as the three worker-based species, corresponds to the holotype of
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="shuckardi">shuckardi</taxonomicName>
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. Heterick (2006) suggests that the locality given for the
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="shuckardi">shuckardi</taxonomicName>
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queen may be in error. This could be correct, but even if true the queen can not be linked to any known species. Resolution of the identity of
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<taxonomicName authority="Bolton & Fisher, 2014" authorityName="Bolton & Fisher" authorityYear="2014" class="Insecta" family="Formicidae" genus="Royidris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="shuckardi">shuckardi</taxonomicName>
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will require the acquisition and comparison of more samples of worker-associated queens.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |