536 lines
97 KiB
XML
536 lines
97 KiB
XML
<document id="D9C5B61324A7E413269422E9211670C5" ID-DOI="10.5281/zenodo.211959" ID-GBIF-Dataset="3a1ff7c7-c93f-49c7-b2c6-61624fdd5684" ID-ISSN="1175-5326" ID-Zenodo-Dep="211959" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460553719715" checkinUser="plazi" docAuthor="Iniesta, Luiz Felipe Moretti, Ferreira, Rodrigo Lopes & Wesener, Thomas" docDate="2012" docId="5571F65F9C14DA2D05D88B9DFB3F6E4A" docLanguage="en" docName="zt03550p042.pdf" docOrigin="Zootaxa 3550" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Glomeridesmus spelaeus Iniesta & Wesener, new species" docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="A9488E279C10DA26054F8B0AFFFA6B28" masterDocTitle="The first troglobitic Glomeridesmus from Brazil, and a template for a modern taxonomic description of Glomeridesmida (Diplopoda)" masterLastPageNumber="42" masterPageNumber="26" pageNumber="30" updateTime="1698318761654" updateUser="plazi">
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<mods:mods id="D41600B7E739DA80CF8B55EC3D7045C8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4D029BB3468366E057C9F514878EADB7">
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<mods:title id="B7DD049F166BBE227CC6FC533A0F0B37">The first troglobitic Glomeridesmus from Brazil, and a template for a modern taxonomic description of Glomeridesmida (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name id="5B4187CA849641835B95FEC3F71D35B0" type="personal">
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<mods:role id="0444B5563CEB362E74D6D4711D6134D0">
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<mods:roleTerm id="980980373DD7334159420F6B750E893C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="578816EDC4C0F754F64E33077A0563AD">Iniesta, Luiz Felipe Moretti</mods:namePart>
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</mods:name>
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<mods:name id="8A9DEE089E4798F52FA0549A5F565207" type="personal">
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<mods:role id="10C2A4CD31524FA08F57C10D67B49819">
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<mods:roleTerm id="070B6484CEF765355C4E7EFB559EA022">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4C3D1B4A2CC539EF52EB095A8545AD42">Ferreira, Rodrigo Lopes</mods:namePart>
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</mods:name>
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<mods:name id="2D167B767201A758921E6AD5B984406B" type="personal">
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<mods:role id="EC2AF8E7EBD078C80B9DF649C4656E92">
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<mods:roleTerm id="29E915BE4F742260EB89351CFB0ABD16">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6AA2306F6D005998F8FC4863D401F7B1">Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="19AAAD245837B4CBE63BE5FA2640D962">text</mods:typeOfResource>
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<mods:relatedItem id="71F9C4CC58770156E4854F133E6C5024" type="host">
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<mods:titleInfo id="A5D57C7363E0924394B119E1671B9967">
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<mods:title id="786C479E851B63859B2A6C55902E7393">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3D74A0D54E77903BB2E8522278842715">
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<mods:date id="DC22739B6835449C18722C711C8F37B7">2012</mods:date>
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<mods:detail id="04288B725B3B06628B24D605339B254B" type="volume">
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||
<mods:number id="3C795BB3F48FE8DAF024FAB434D0C0E2">3550</mods:number>
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</mods:detail>
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<mods:extent id="D5045C56014D0408D7CD4D6AB42F4219" unit="page">
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<mods:start id="0DB0E7F5ADFA506DC38BBC31E7237636">26</mods:start>
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<mods:end id="F63FFF9A875BDB600F577DE4C7651922">42</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="A3E49939F64F2210CF53D9849A45A677">journal article</mods:classification>
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<mods:identifier id="5AB825CF133443A4C749D3659997B729" type="DOI">10.5281/zenodo.211959</mods:identifier>
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<mods:identifier id="325ACC91D8E4A500DEF0809B1ADE4E8B" type="GBIF-Dataset">3a1ff7c7-c93f-49c7-b2c6-61624fdd5684</mods:identifier>
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<mods:identifier id="8E5270AE125993271B4F9D65D8694FB7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7C26F779D9716B57329E277FEF8AB290" type="Zenodo-Dep">211959</mods:identifier>
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</mods:mods>
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<treatment id="5571F65F9C14DA2D05D88B9DFB3F6E4A" ID-DOI="http://doi.org/10.5281/zenodo.6179710" ID-GBIF-Taxon="119555238" ID-Zenodo-Dep="6179710" LSID="urn:lsid:plazi:treatment:5571F65F9C14DA2D05D88B9DFB3F6E4A" httpUri="http://treatment.plazi.org/id/5571F65F9C14DA2D05D88B9DFB3F6E4A" lastPageId="11" lastPageNumber="37" pageId="4" pageNumber="30">
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||
<subSubSection id="95C214C29C14DA2205D88B9DFCB16B99" box="[151,843,151,177]" pageId="4" pageNumber="30" type="nomenclature">
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||
<paragraph id="DD6747499C14DA2205D88B9DFCB16B99" blockId="4.[151,843,151,177]" box="[151,843,151,177]" pageId="4" pageNumber="30">
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||
<heading id="862FF0259C14DA2205D88B9DFCB16B99" bold="true" box="[151,843,151,177]" fontSize="11" level="1" pageId="4" pageNumber="30" reason="1">
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||
<emphasis id="EFAC9B5B9C14DA2205D88B9DFCB16B99" bold="true" box="[151,843,151,177]" pageId="4" pageNumber="30">
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<taxonomicName id="1AD83CCA9C14DA2205D88B9DFD556B99" ID-CoL="3GCN7" authority="Iniesta & Wesener" authorityName="Iniesta & Wesener" box="[151,687,151,177]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
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<emphasis id="EFAC9B5B9C14DA2205D88B9DFE3B6B99" bold="true" box="[151,449,151,177]" italics="true" pageId="4" pageNumber="30">Glomeridesmus spelaeus</emphasis>
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Iniesta & Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel id="F49F26209C14DA2207F48B9DFCB16B99" box="[699,843,151,177]" pageId="4" pageNumber="30" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="95C214C29C14DA2205D88BD4FEE769C6" pageId="4" pageNumber="30" type="materials_examined">
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<paragraph id="DD6747499C14DA2205D88BD4FBC76A32" blockId="4.[151,1437,222,2010]" pageId="4" pageNumber="30">
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<emphasis id="EFAC9B5B9C14DA2205D88BD4FE7F6BDF" bold="true" box="[151,389,222,247]" pageId="4" pageNumber="30">Material examined:</emphasis>
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<typeStatus id="0263F9EB9C14DA2204C28BD4FE006BDE" box="[397,506,222,246]" pageId="4" pageNumber="30" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="CBDE8CC09C14DA2207498BD4FDA26BDE" box="[518,600,222,246]" pageId="4" pageNumber="30" type="male">1 Male</specimenCount>
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(
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<collectingCountry id="A5CF07D99C14DA2207278BD4FD526BDE" box="[616,680,222,246]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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1906) SLQ31 EU,
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<collectingCountry id="A5CF07D99C14DA2206C58BD4FC2B6BDE" box="[906,977,222,246]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
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, Pará, Curionópolis, iron cave SL 31, E 0650189m, N
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<quantity id="1A20EAAC9C14DA2204778A08FE486A32" box="[312,434,258,282]" metricMagnitude="6" metricUnit="m" metricValue="9.339714" pageId="4" pageNumber="30" unit="m" value="9339714.0">9339714m</quantity>
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, in bat guano pile far from entrance, Zampaulo, R.A. coll.
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</paragraph>
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<paragraph id="DD6747499C14DA2205888A2DFB8C6AAE" blockId="4.[151,1437,222,2010]" pageId="4" pageNumber="30">
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<typeStatus id="0263F9EB9C14DA2205888A2DFEC36A17" box="[199,313,295,319]" pageId="4" pageNumber="30" type="paratype">Paratypes</typeStatus>
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: 1 M (fragmented,
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<collectionCode id="BBC9DF8C9C14DA2207418A2DFDA66A16" LSID="urn:lsid:biocol.org:col:34613" box="[526,604,295,318]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="30">ZFMK</collectionCode>
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<collectionCode id="BBC9DF8C9C14DA22072C8A2DFD596A16" box="[611,675,295,318]" pageId="4" pageNumber="30">MYR</collectionCode>
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0936), 1 F (
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<collectionCode id="BBC9DF8C9C14DA2206638A2CFC866A16" LSID="urn:lsid:biocol.org:col:34613" box="[812,892,294,318]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="30">ZFMK</collectionCode>
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<collectionCode id="BBC9DF8C9C14DA2206CC8A2DFC396A16" box="[899,963,295,318]" pageId="4" pageNumber="30">MYR</collectionCode>
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0937), '1758', same data as
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<typeStatus id="0263F9EB9C14DA2201B28A2CFA996A16" box="[1277,1379,294,318]" pageId="4" pageNumber="30" type="holotype">holotype</typeStatus>
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; 3 M (fragmented,
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<collectingCountry id="A5CF07D99C14DA22047F8A40FE946A4A" box="[304,366,330,354]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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1758,
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<collectingCountry id="A5CF07D99C14DA22048F8A40FE046A4A" box="[448,510,330,354]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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1759, fragmented
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<collectingCountry id="A5CF07D99C14DA2207968A40FCED6A4A" box="[729,791,330,354]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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3837); 5 F (
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<collectingCountry id="A5CF07D99C14DA2206E48A40FC116A4A" box="[939,1003,330,354]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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1928,
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<collectingCountry id="A5CF07D99C14DA2201728A40FB816A4A" box="[1085,1147,330,354]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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1929,
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<collectingCountry id="A5CF07D99C14DA2201828A40FAF16A4A" box="[1229,1291,330,354]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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1930,
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<collectingCountry id="A5CF07D99C14DA2200118A40FA666A4A" box="[1374,1436,330,354]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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1931, fragmented
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<collectingCountry id="A5CF07D99C14DA22042C8A64FE5B6AAE" box="[355,417,366,390]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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3838);
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||
<specimenCount id="CBDE8CC09C14DA2204B68A64FDCD6AAE" box="[505,567,366,390]" pageId="4" pageNumber="30" type="generic">4 juv.</specimenCount>
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||
(
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<collectingCountry id="A5CF07D99C14DA22070A8A64FD7F6AAE" box="[581,645,366,390]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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||
1932,
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<collectingCountry id="A5CF07D99C14DA22079D8A64FCEA6AAE" box="[722,784,366,390]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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||
1933,
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||
<collectingCountry id="A5CF07D99C14DA2206128A64FC616AAE" box="[861,923,366,390]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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||
1934,
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<collectingCountry id="A5CF07D99C14DA2206A78A64FBDC6AAE" box="[1000,1062,366,390]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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||
1935).
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||
</paragraph>
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||
<paragraph id="DD6747499C14DA2205888A98FEE769C6" blockId="4.[151,1437,222,2010]" pageId="4" pageNumber="30">
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<emphasis id="EFAC9B5B9C14DA2205888A98FE096A83" bold="true" box="[199,499,402,427]" pageId="4" pageNumber="30">Other material examined</emphasis>
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||
: 3 F (
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<collectingCountry id="A5CF07D99C14DA22077C8A98FD896A82" box="[563,627,402,426]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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||
3814,
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<collectingCountry id="A5CF07D99C14DA2207F08A98FD076A82" box="[703,765,402,426]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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3815,
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<collectingCountry id="A5CF07D99C14DA2206068A98FC7D6A82" box="[841,903,402,426]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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3816),
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<collectingCountry id="A5CF07D99C14DA2206928A98FBDE6A82" box="[989,1060,402,426]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
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, Pará, Curionópolis, iron cave SL 31, E 0652243m/N
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<quantity id="1A20EAAC9C14DA2204358ABCFE0F6AE6" box="[378,501,438,462]" metricMagnitude="6" metricUnit="m" metricValue="9.339216" pageId="4" pageNumber="30" unit="m" value="9339216.0">9339216m</quantity>
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, Zampaulo, R.A. coll. same data as
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<typeStatus id="0263F9EB9C14DA2206D88ABCFC076AE6" box="[919,1021,438,462]" pageId="4" pageNumber="30" type="holotype">holotype</typeStatus>
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; 2 F (
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<collectingCountry id="A5CF07D99C14DA22010D8ABCFB786AE6" box="[1090,1154,438,462]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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3817,
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<collectingCountry id="A5CF07D99C14DA22019C8ABCFAEB6AE6" box="[1235,1297,438,462]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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3818), 1 M (
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<collectingCountry id="A5CF07D99C14DA2205D18AD0FF246ADA" box="[158,222,474,498]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
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3819)
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<collectingCountry id="A5CF07D99C14DA22047E8AD0FE826ADA" box="[305,376,474,498]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
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, Pará, Curionópolis, iron cave SL 31, E 0652243m/N
|
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<quantity id="1A20EAAC9C14DA2206AC8AD0FBA76ADA" box="[995,1117,474,498]" metricMagnitude="6" metricUnit="m" metricValue="9.339216" pageId="4" pageNumber="30" unit="m" value="9339216.0">9339216m</quantity>
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||
, Zampaulo, R.A. coll. same data as
|
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<typeStatus id="0263F9EB9C14DA2205A28AF4FEA8693E" box="[237,338,510,534]" pageId="4" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
; 4 F (
|
||
<collectingCountry id="A5CF07D99C14DA2204DB8AF4FE2E693E" box="[404,468,510,534]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3824,
|
||
<collectingCountry id="A5CF07D99C14DA22076C8AF4FD9B693E" box="[547,609,510,534]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3825,
|
||
<collectingCountry id="A5CF07D99C14DA2207FE8AF4FD15693E" box="[689,751,510,534]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3826,
|
||
<collectingCountry id="A5CF07D99C14DA2206718AF4FC86693E" box="[830,892,510,534]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3827),
|
||
<collectingCountry id="A5CF07D99C14DA22069A8AF4FBE6693E" box="[981,1052,510,534]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
|
||
, Pará, Curionópolis, iron cave SL 58, E 0652243m/N
|
||
<quantity id="1A20EAAC9C14DA2204378928FE086912" box="[376,498,546,570]" metricMagnitude="6" metricUnit="m" metricValue="9.339216" pageId="4" pageNumber="30" unit="m" value="9339216.0">9339216m</quantity>
|
||
, Zampaulo, R.A. coll.
|
||
<date id="A96661899C14DA2207B88928FC996912" box="[759,867,546,570]" pageId="4" pageNumber="30" value="2011-01-29">29.i.2011</date>
|
||
; 1 F (
|
||
<collectingCountry id="A5CF07D99C14DA2206E98928FC1C6912" box="[934,998,546,570]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3820)
|
||
<collectingCountry id="A5CF07D99C14DA2201778928FB856912" box="[1080,1151,546,570]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
|
||
, Pará, Curionópolis, iron cave SL 61, E 0
|
||
<quantity id="1A20EAAC9C14DA22040A894CFE4A6976" box="[325,432,582,606]" metricMagnitude="5" metricUnit="m" metricValue="6.525119999999999" pageId="4" pageNumber="30" unit="m" value="652512.0">652512m</quantity>
|
||
/N
|
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<quantity id="1A20EAAC9C14DA22049C894CFDB66976" box="[467,588,582,606]" metricMagnitude="6" metricUnit="m" metricValue="9.338874" pageId="4" pageNumber="30" unit="m" value="9338874.0">9338874m</quantity>
|
||
, Zampaulo, R.A. coll.
|
||
<date id="A96661899C14DA220606894CFC4F6976" box="[841,949,582,606]" pageId="4" pageNumber="30" value="2011-01-10">10.i.2011</date>
|
||
; 2 F (
|
||
<collectingCountry id="A5CF07D99C14DA2206BD894CFBC86976" box="[1010,1074,582,606]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3828,
|
||
<collectingCountry id="A5CF07D99C14DA220131894CFB466976" box="[1150,1212,582,606]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3829),
|
||
<collectingCountry id="A5CF07D99C14DA22005E894CFAA26976" box="[1297,1368,582,606]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
|
||
, Pará, Curionópolis, iron cave SL 93, E 0
|
||
<quantity id="1A20EAAC9C14DA22077B8960FD6469AA" box="[564,670,618,642]" metricMagnitude="5" metricUnit="m" metricValue="6.513859999999999" pageId="4" pageNumber="30" unit="m" value="651386.0">651386m</quantity>
|
||
/N
|
||
<quantity id="1A20EAAC9C14DA22078A8960FCC569AA" box="[709,831,618,642]" metricMagnitude="6" metricUnit="m" metricValue="9.34193" pageId="4" pageNumber="30" unit="m" value="9341930.0">9341930m</quantity>
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||
, Zampaulo, R.A. coll.
|
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<date id="A96661899C14DA2201018960FB4369AA" box="[1102,1209,618,642]" pageId="4" pageNumber="30" value="2011-01-17">17.i.2011</date>
|
||
; 1 F, (
|
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<collectingCountry id="A5CF07D99C14DA2200468960FAB369AA" box="[1289,1353,618,642]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3823),
|
||
<collectingCountry id="A5CF07D99C14DA2205D88984FF24698E" box="[151,222,654,678]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
|
||
, Pará, Curionópolis, iron cave SL 95, E
|
||
<quantity id="1A20EAAC9C14DA2207EF8984FCF3698E" box="[672,777,654,678]" metricMagnitude="5" metricUnit="m" metricValue="6.51417" pageId="4" pageNumber="30" unit="m" value="651417.0">651417m</quantity>
|
||
/N
|
||
<quantity id="1A20EAAC9C14DA2206628984FC5D698E" box="[813,935,654,678]" metricMagnitude="6" metricUnit="m" metricValue="9.341905999999998" pageId="4" pageNumber="30" unit="m" value="9341906.0">9341906m</quantity>
|
||
, Zampaulo, R.A. coll.
|
||
<date id="A96661899C14DA2201EA8984FAE2698E" box="[1189,1304,654,678]" pageId="4" pageNumber="30" value="2011-02-03">03.ii.2011</date>
|
||
; 2 F, (
|
||
<collectingCountry id="A5CF07D99C14DA2200138984FA66698E" box="[1372,1436,654,678]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3821,
|
||
<collectingCountry id="A5CF07D99C14DA22059189B8FEE669E2" box="[222,284,690,714]" name="Iceland" pageId="4" pageNumber="30">ISLA</collectingCountry>
|
||
3822),
|
||
<collectingCountry id="A5CF07D99C14DA22043A89B8FE4669E2" box="[373,444,690,714]" name="Brazil" pageId="4" pageNumber="30">Brazil</collectingCountry>
|
||
, Pará, Curionópolis, iron cave SL 97, E 0651411m/N
|
||
<quantity id="1A20EAAC9C14DA22016A89B8FB5A69E2" box="[1061,1184,690,714]" metricMagnitude="6" metricUnit="m" metricValue="9.339212" pageId="4" pageNumber="30" unit="m" value="9339212.0">9339212m</quantity>
|
||
, Zampaulo, R.A. coll.
|
||
<date id="A96661899C14DA2205D889DCFEE369C6" box="[151,281,726,750]" pageId="4" pageNumber="30" value="2011-12-04">04.xii.2011</date>
|
||
;
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95C214C29C14DA22058889F0FAC96F26" pageId="4" pageNumber="30" type="diagnosis">
|
||
<paragraph id="DD6747499C14DA22058889F0FAC96F26" blockId="4.[151,1437,222,2010]" pageId="4" pageNumber="30">
|
||
<emphasis id="EFAC9B5B9C14DA22058889F0FEBB683B" bold="true" box="[199,321,762,787]" pageId="4" pageNumber="30">Diagnosis.</emphasis>
|
||
White, translucent body (
|
||
<figureCitation id="45E35BCC9C14DA22072A89F0FD52683A" box="[613,680,762,786]" captionStart="FIGURE 2" captionStartId="5.[151,250,1710,1733]" captionTargetBox="[241,1345,733,1679]" captionTargetId="figure@5.[241,1345,733,1679]" captionTargetPageId="5" captionText="FIGURE 2. Glomeridesmus spelaeus, living specimens. First published pictures of living Glomeridesmida. arrows point to eggs and and intestine. Note the large eggs. Intestine not straight tube but folded." httpUri="https://zenodo.org/record/211961/files/figure.png" pageId="4" pageNumber="30">Fig. 2</figureCitation>
|
||
) shared with
|
||
<taxonomicName id="1AD83CCA9C14DA22060E89F1FBAD683A" authority="Shear, 1974" authorityName="Shear" authorityYear="1974" box="[833,1111,762,787]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis id="EFAC9B5B9C14DA22060E89F1FC30683B" box="[833,970,763,787]" italics="true" pageId="4" pageNumber="30">G. sbordonii</emphasis>
|
||
<bibRefCitation id="B9493AB89C14DA22069D89F0FBAD683A" author="Shear" box="[978,1111,762,786]" pageId="4" pageNumber="30" refString="Shear, W. A. (1974) Millipeds (Diplopoda) from Mexican and Guatemalan caves. Accademia Nazionale Lincei Rendiconti Classe Scienze Fisiche Matematiche e Naturali, 171, 205 - 239." type="journal article" year="1974">Shear, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="A5CF07D99C14DA2201D389F0FB0E683A" box="[1180,1268,762,786]" name="Mexico" pageId="4" pageNumber="30">Mexico</collectingCountry>
|
||
, the only other known troglobitic species of the order.
|
||
<emphasis id="EFAC9B5B9C14DA2207188815FD2E681F" box="[599,724,798,823]" italics="true" pageId="4" pageNumber="30">G. s p e l a e u s</emphasis>
|
||
<emphasis id="EFAC9B5B9C14DA2207AE8815FCDB681F" bold="true" box="[737,801,799,823]" pageId="4" pageNumber="30">n. sp.</emphasis>
|
||
differs from
|
||
<taxonomicName id="1AD83CCA9C14DA2206F48815FBBF681E" box="[955,1093,798,822]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis id="EFAC9B5B9C14DA2206F48815FBBF681E" box="[955,1093,798,822]" italics="true" pageId="4" pageNumber="30">G. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
in the presence of sclerotized knobs with mechanoreceptorical(?) setae on tergites (smooth and glabrous in
|
||
<taxonomicName id="1AD83CCA9C14DA22015C8849FB646873" box="[1043,1182,835,859]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis id="EFAC9B5B9C14DA22015C8849FB646873" box="[1043,1182,835,859]" italics="true" pageId="4" pageNumber="30">G. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
), larger size (~
|
||
<quantity id="1A20EAAC9C14DA2200168848FA666872" box="[1369,1436,834,858]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="30" unit="mm" value="5.0">5 mm</quantity>
|
||
body length in
|
||
<taxonomicName id="1AD83CCA9C14DA220405886DFE2C6856" box="[330,470,870,894]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis id="EFAC9B5B9C14DA220405886DFE2C6856" box="[330,470,870,894]" italics="true" pageId="4" pageNumber="30">G. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
), antennae more densely pilose than in
|
||
<taxonomicName id="1AD83CCA9C14DA2206E5886DFBCC6856" box="[938,1078,870,894]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="sbordonii">
|
||
<emphasis id="EFAC9B5B9C14DA2206E5886DFBCC6856" box="[938,1078,870,894]" italics="true" pageId="4" pageNumber="30">G. sbordonii</emphasis>
|
||
</taxonomicName>
|
||
(which could be a perspective error), joint 3 of telopod in oral (anterior) view not overlapping joint 4, as well as other telopod characteristic (provided drawing of
|
||
<emphasis id="EFAC9B5B9C14DA2204DE88A5FDE368EE" box="[401,537,942,966]" italics="true" pageId="4" pageNumber="30">
|
||
S.
|
||
<taxonomicName id="1AD83CCA9C14DA2204E088A4FDE368EE" box="[431,537,942,966]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="sbordonii">sbordonii</taxonomicName>
|
||
</emphasis>
|
||
[
|
||
<bibRefCitation id="B9493AB89C14DA22076788A4FD0068EE" author="Shear" box="[552,762,942,966]" pageId="4" pageNumber="30" refString="Shear, W. A. (1974) Millipeds (Diplopoda) from Mexican and Guatemalan caves. Accademia Nazionale Lincei Rendiconti Classe Scienze Fisiche Matematiche e Naturali, 171, 205 - 239." type="journal article" year="1974">Shear 1974, fig. 2</bibRefCitation>
|
||
] difficult to compare with our SEM images). Location and shape of projection (immovable finger) of joint 3, posteriorly instead of laterally of joint 4, appears to be unique for the order (
|
||
<figureCitation id="45E35BCC9C14DA22044588FCFEB66F26" box="[266,332,1014,1038]" captionStart="FIGURE 9. G. s p e l a e u s n" captionStartId="13.[151,250,1753,1776]" captionTargetBox="[151,1435,527,1737]" captionTargetId="figure@13.[151,1436,527,1738]" captionTargetPageId="13" captionText="FIGURE 9. G. s p e l a e u s n. sp., paratype males, SEM, telopod, posterior view. A: right telopod, podomeres 2 – 4 (ZFMK Myr 0936); B: left telopod, podomeres 2 – 4 (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936); D: right telopod, detail of podomere 3 & 4 (ZFMK Myr 0936). Abbreviations: mem = membranous area; pro = process of podomere 3; roman numerals refer to podomere number. Scale bars: A = 100 µm; B = 100 µm; C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/211969/files/figure.png" pageId="4" pageNumber="30">Fig. 9</figureCitation>
|
||
A, B, D), albeit telopods are only known from <1/3 of described
|
||
<taxonomicName id="1AD83CCA9C14DA22016B88FCFB2E6F26" box="[1060,1236,1014,1038]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EFAC9B5B9C14DA22016B88FCFB2E6F26" box="[1060,1236,1014,1038]" italics="true" pageId="4" pageNumber="30">Glomeridesmus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95C214C29C14DA2205888F11FA666F1A" box="[199,1436,1050,1075]" pageId="4" pageNumber="30" type="etymology">
|
||
<paragraph id="DD6747499C14DA2205888F11FA666F1A" blockId="4.[151,1437,222,2010]" box="[199,1436,1050,1075]" pageId="4" pageNumber="30">
|
||
<emphasis id="EFAC9B5B9C14DA2205888F11FEB46F1B" bold="true" box="[199,334,1051,1075]" pageId="4" pageNumber="30">Etymology.</emphasis>
|
||
Species epithet is a noun in apposition, as a reference to the Latin word
|
||
<taxonomicName id="1AD83CCA9C14DA22013B8F11FB2E6F1B" box="[1140,1236,1051,1075]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis id="EFAC9B5B9C14DA22013B8F11FB2E6F1B" box="[1140,1236,1051,1075]" italics="true" pageId="4" pageNumber="30">spelaeus</emphasis>
|
||
</taxonomicName>
|
||
, meaning “cave”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95C214C29C14DA2D05888F34FB3F6E4A" lastPageId="11" lastPageNumber="37" pageId="4" pageNumber="30" type="description">
|
||
<paragraph id="DD6747499C14DA2205888F34FE836F52" blockId="4.[151,1437,222,2010]" pageId="4" pageNumber="30">
|
||
<emphasis id="EFAC9B5B9C14DA2205888F34FEA36F7F" bold="true" box="[199,345,1086,1111]" pageId="4" pageNumber="30">Description.</emphasis>
|
||
<emphasis id="EFAC9B5B9C14DA22042E8F4AFDF96F7F" box="[353,515,1088,1111]" italics="true" pageId="4" pageNumber="30">Measurements</emphasis>
|
||
: (largest) adult females (20+
|
||
<collectingCountry id="A5CF07D99C14DA2206008F34FC896F7E" box="[847,883,1086,1110]" name="American Samoa" pageId="4" pageNumber="30">AS</collectingCountry>
|
||
tergites, 35+1 leg pairs): length:
|
||
<quantity id="1A20EAAC9C14DA2201BC8F34FAB46F7E" box="[1267,1358,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="30" unit="mm" value="8.6">8.6 mm</quantity>
|
||
; width (midbody):
|
||
<quantity id="1A20EAAC9C14DA2204538F68FE8F6F52" box="[284,373,1122,1146]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="30" unit="mm" value="2.3">2.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DD6747499C14DA2205888F8CFCB56FEA" blockId="4.[151,1437,222,2010]" pageId="4" pageNumber="30">
|
||
Males (19+
|
||
<collectingCountry id="A5CF07D99C14DA2204098F8CFE966FB6" box="[326,364,1158,1182]" name="American Samoa" pageId="4" pageNumber="30">AS</collectingCountry>
|
||
tergites, 33+1+T leg pairs,
|
||
<typeStatus id="0263F9EB9C14DA2207E98F8CFCF26FB6" box="[678,776,1158,1182]" pageId="4" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
male): length: 7.2; width (midbody): 2.1; (32+1+T, n =2): length: 5.4, width: 1.6; (31+1+T, n=1): length: 6.3, width: 1.9.
|
||
</paragraph>
|
||
<paragraph id="DD6747499C14DA2205888FC4FE116FCE" blockId="4.[151,1437,222,2010]" box="[199,491,1230,1254]" pageId="4" pageNumber="30">Colour: translucent white.</paragraph>
|
||
<paragraph id="DD6747499C14DA2205888FF9FC276DF6" blockId="4.[151,1437,222,2010]" pageId="4" pageNumber="30">
|
||
<emphasis id="EFAC9B5B9C14DA2205888FF9FEF76E23" box="[199,269,1267,1291]" italics="true" pageId="4" pageNumber="30">Head:</emphasis>
|
||
General shape: typical for the family (
|
||
<figureCitation id="45E35BCC9C14DA2207F48FF8FCFB6E22" box="[699,769,1266,1290]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Figs 3</figureCitation>
|
||
C, E). Epicranium glabrous; frons, clypeus and labrum with several isolated setae. Both antennae widely separated by a distance (300 µm) longer than twice the width of the first antennal joint. Genae (area below the antennae) almost non-existant, but basal joint of mandible large and visible in dorsal view. Antennal base and organ of Tömösváry surrounded by sclerotized rim (
|
||
<figureCitation id="45E35BCC9C14DA2200458E54FAAB6E5E" box="[1290,1361,1374,1398]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Fig. 3</figureCitation>
|
||
D, E). Antennae consisting of seven joints, each covered with numerous setae (
|
||
<figureCitation id="45E35BCC9C14DA2206AC8E88FBD36EB2" box="[995,1065,1410,1434]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Fig. 3</figureCitation>
|
||
E). Joints 4–6 slightly flattened, apically much wider than basally, wider than long (
|
||
<figureCitation id="45E35BCC9C14DA2207BF8EACFCCF6E96" box="[752,821,1446,1470]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Fig. 3</figureCitation>
|
||
C, E). Apical disc carrying four long (40 µm) apical cones; sensilla basiconicae seem to be absent (
|
||
<figureCitation id="45E35BCC9C14DA2207E28EC0FD0B6ECA" box="[685,753,1482,1506]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Fig. 3</figureCitation>
|
||
F). Tömösváry organs slightly larger than antennal base. Of well-rounded oval shape, interior covered by sclerotized plate (
|
||
<figureCitation id="45E35BCC9C14DA22063F8EE4FC4F6D2E" box="[880,949,1518,1542]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Fig. 3</figureCitation>
|
||
D). Gnathochilarium with very broad gula (hypostoma). Cardines very small, separated from basal part of gnathochilarium. Mentum basally towards gula with a transverse elevated area, completely fused with laminae linguales (
|
||
<figureCitation id="45E35BCC9C14DA2206A68D3CFBBF6D66" box="[1001,1093,1590,1614]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Fig. 3G</figureCitation>
|
||
). Stipites laterally with single sclerotized ledge. Whole surface of gnathochilarium covered with few, isolated setae. Stipites apically with 2, lamello-mentum with 4 extra-long setae. Lateral palpi thin, carrying 4 or 5 sensory cones; inner palpi very wide, carrying several dozen long, tube-shaped sensory cones; central pads large, mesally touching one another, in ventral view also covered with numerous long, tube-shaped sensory cones.
|
||
</paragraph>
|
||
<paragraph id="DD6747499C14DA2205888DE0FB656C2A" blockId="4.[151,1437,222,2010]" box="[199,1183,1770,1794]" pageId="4" pageNumber="30">
|
||
Mandible: with single, densely pilose basal joint (
|
||
<figureCitation id="45E35BCC9C14DA2207BB8DE0FCCD6C2A" box="[756,823,1770,1794]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="4" pageNumber="30">Fig. 3</figureCitation>
|
||
C). Apical part not investigated.
|
||
</paragraph>
|
||
<paragraph id="DD6747499C14DA2205888C04FBCD6C0E" blockId="4.[151,1437,222,2010]" box="[199,1079,1806,1830]" pageId="4" pageNumber="30">
|
||
<emphasis id="EFAC9B5B9C14DA2205888C04FEEB6C0E" box="[199,273,1806,1830]" italics="true" pageId="4" pageNumber="30">Trunk:</emphasis>
|
||
Collum (tergite 1): wider than head, shape similar to following tergites.
|
||
</paragraph>
|
||
<paragraph id="DD6747499C14DA2305888C38FDF569AC" blockId="4.[151,1437,222,2010]" lastBlockId="5.[151,1436,151,644]" lastPageId="5" lastPageNumber="31" pageId="4" pageNumber="30">
|
||
Tergites 2–19 (20): very thin, decalcified and difficult to distinguish from one another. Size of tergites first increasing until tergite 8, than slowly decreasing more posteriorly. Each tergite divided by 8–10 longitudinal striae, each stria projecting with a toothed margin above more posterior stria (
|
||
<figureCitation id="45E35BCC9C14DA2206B08C70FBB36CBA" box="[1023,1097,1914,1938]" captionStart="FIGURE 6. G" captionStartId="9.[151,250,1729,1752]" captionTargetBox="[152,1435,179,1714]" captionTargetId="figure@9.[151,1436,179,1715]" captionTargetPageId="9" captionText="FIGURE 6. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM, midbody tergite. A: dorsal view; B: lateral view; C: sclerotized knob with setae; D: posterior margin, ' limbus'; E: anterior margin. Abbreviations: ant = anterior; s = (sensorial) seta; str = stria; arrow points to oral side. Scale bars: A = 500 µm; B = 100 µm; C = 10 µm; D = 100 µm; E = 200 µm." httpUri="https://zenodo.org/record/211966/files/figure.png" pageId="4" pageNumber="30">Fig. 6</figureCitation>
|
||
B). Distances between striae expanding towards posterior margin (
|
||
<figureCitation id="45E35BCC9C14DA22070B8C94FD736C9E" box="[580,649,1950,1974]" captionStart="FIGURE 6. G" captionStartId="9.[151,250,1729,1752]" captionTargetBox="[152,1435,179,1714]" captionTargetId="figure@9.[151,1436,179,1715]" captionTargetPageId="9" captionText="FIGURE 6. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM, midbody tergite. A: dorsal view; B: lateral view; C: sclerotized knob with setae; D: posterior margin, ' limbus'; E: anterior margin. Abbreviations: ant = anterior; s = (sensorial) seta; str = stria; arrow points to oral side. Scale bars: A = 500 µm; B = 100 µm; C = 10 µm; D = 100 µm; E = 200 µm." httpUri="https://zenodo.org/record/211966/files/figure.png" pageId="4" pageNumber="30">Fig. 6</figureCitation>
|
||
E). Posterior margin with single row of large sclerotized knobs (
|
||
<figureCitation id="45E35BCC9C14DA2200218C94FF5F6CF2" captionStart="FIGURE 6. G" captionStartId="9.[151,250,1729,1752]" captionTargetBox="[152,1435,179,1714]" captionTargetId="figure@9.[151,1436,179,1715]" captionTargetPageId="9" captionText="FIGURE 6. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM, midbody tergite. A: dorsal view; B: lateral view; C: sclerotized knob with setae; D: posterior margin, ' limbus'; E: anterior margin. Abbreviations: ant = anterior; s = (sensorial) seta; str = stria; arrow points to oral side. Scale bars: A = 500 µm; B = 100 µm; C = 10 µm; D = 100 µm; E = 200 µm." httpUri="https://zenodo.org/record/211966/files/figure.png" pageId="4" pageNumber="30">Fig. 6</figureCitation>
|
||
A, B, D). Knobs accompanied by longer sensorial setae (
|
||
<figureCitation id="45E35BCC9C14DA22066B8CC8FC926CF2" box="[804,872,1986,2010]" captionStart="FIGURE 6. G" captionStartId="9.[151,250,1729,1752]" captionTargetBox="[152,1435,179,1714]" captionTargetId="figure@9.[151,1436,179,1715]" captionTargetPageId="9" captionText="FIGURE 6. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM, midbody tergite. A: dorsal view; B: lateral view; C: sclerotized knob with setae; D: posterior margin, ' limbus'; E: anterior margin. Abbreviations: ant = anterior; s = (sensorial) seta; str = stria; arrow points to oral side. Scale bars: A = 500 µm; B = 100 µm; C = 10 µm; D = 100 µm; E = 200 µm." httpUri="https://zenodo.org/record/211966/files/figure.png" pageId="4" pageNumber="30">Fig. 6</figureCitation>
|
||
C). Surface of tergites covered by a sparse field of very short setae (
|
||
<figureCitation id="45E35BCC9C15DA2304138B9DFE586B87" box="[348,418,151,175]" captionStart="FIGURE 6. G" captionStartId="9.[151,250,1729,1752]" captionTargetBox="[152,1435,179,1714]" captionTargetId="figure@9.[151,1436,179,1715]" captionTargetPageId="9" captionText="FIGURE 6. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM, midbody tergite. A: dorsal view; B: lateral view; C: sclerotized knob with setae; D: posterior margin, ' limbus'; E: anterior margin. Abbreviations: ant = anterior; s = (sensorial) seta; str = stria; arrow points to oral side. Scale bars: A = 500 µm; B = 100 µm; C = 10 µm; D = 100 µm; E = 200 µm." httpUri="https://zenodo.org/record/211966/files/figure.png" pageId="5" pageNumber="31">Fig. 6</figureCitation>
|
||
B). Tergites at apico-lateral corner with 2 or 3 striae and denser rows of short setae (
|
||
<figureCitation id="45E35BCC9C15DA2300218B9DFF5C6BFC" captionStart="FIGURE 6. G" captionStartId="9.[151,250,1729,1752]" captionTargetBox="[152,1435,179,1714]" captionTargetId="figure@9.[151,1436,179,1715]" captionTargetPageId="9" captionText="FIGURE 6. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM, midbody tergite. A: dorsal view; B: lateral view; C: sclerotized knob with setae; D: posterior margin, ' limbus'; E: anterior margin. Abbreviations: ant = anterior; s = (sensorial) seta; str = stria; arrow points to oral side. Scale bars: A = 500 µm; B = 100 µm; C = 10 µm; D = 100 µm; E = 200 µm." httpUri="https://zenodo.org/record/211966/files/figure.png" pageId="5" pageNumber="31">Fig. 6</figureCitation>
|
||
B). Tergite 12 onwards posterior-lateral edge pronounced into a sharp-edged tip, becoming a longer, spine like process at second last tergites. Last tergite at posterior margin towards anal shield without (male) or with (female) single row of short spines. Limbus (posterior margin of tergite) smooth, without any structures (
|
||
<figureCitation id="45E35BCC9C15DA2300618A0EFA8D6A34" box="[1326,1399,260,284]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="5" pageNumber="31">Fig. 5</figureCitation>
|
||
D). Endotergum (underside of posterior margin of tergite) smooth area without any specific structures (
|
||
<figureCitation id="45E35BCC9C15DA23007F8A2DFA866A17" box="[1328,1404,295,319]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="5" pageNumber="31">Figs 5</figureCitation>
|
||
F). Anterior margin of tergite smooth, without any structures. (
|
||
<figureCitation id="45E35BCC9C15DA2306158A46FC5B6A4C" box="[858,929,332,356]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="5" pageNumber="31">Fig. 5</figureCitation>
|
||
D, E). Latero-tergites of rectangular shape, posterior margin overlapping first third of subsequent latero-tergite (
|
||
<figureCitation id="45E35BCC9C15DA2306EF8A65FC1E6AAF" box="[928,996,367,391]" captionStart="FIGURE 4. G. s p e l a e u s n" captionStartId="7.[151,250,1694,1717]" captionTargetBox="[151,1435,188,1673]" captionTargetId="figure@7.[151,1436,188,1674]" captionTargetPageId="7" captionText="FIGURE 4. G. s p e l a e u s n. sp., A – D paratype male (ZFMK Myr 0936), E paratype female (ISLA 3838) SEM. A: leg pairs 4 – 7 with latero-tergites; B: coxosternite 6 with stigma opening; C: coxal pouch 7; D: midbody latero-tergite of male; E: endbody latero-tergite female. Abbreviations: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number. Scale bars: A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm." httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="5" pageNumber="31">Fig. 4</figureCitation>
|
||
A). Anterior and posterior sides longer than mesal and lateral ones (
|
||
<figureCitation id="45E35BCC9C15DA23049B8A9EFDED6A84" box="[468,535,404,428]" captionStart="FIGURE 4. G. s p e l a e u s n" captionStartId="7.[151,250,1694,1717]" captionTargetBox="[151,1435,188,1673]" captionTargetId="figure@7.[151,1436,188,1674]" captionTargetPageId="7" captionText="FIGURE 4. G. s p e l a e u s n. sp., A – D paratype male (ZFMK Myr 0936), E paratype female (ISLA 3838) SEM. A: leg pairs 4 – 7 with latero-tergites; B: coxosternite 6 with stigma opening; C: coxal pouch 7; D: midbody latero-tergite of male; E: endbody latero-tergite female. Abbreviations: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number. Scale bars: A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm." httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="5" pageNumber="31">Fig. 4</figureCitation>
|
||
D, E). Mesal side with a rounded angle towards coxosternite, marginal brim here extraordinarily thick; midbody latero-tergite with an elevated marginal brim engulfing all but posterior margin (
|
||
<figureCitation id="45E35BCC9C15DA2305D08AD6FF216ADC" box="[159,219,476,500]" captionStart="FIGURE 4. G. s p e l a e u s n" captionStartId="7.[151,250,1694,1717]" captionTargetBox="[151,1435,188,1673]" captionTargetId="figure@7.[151,1436,188,1674]" captionTargetPageId="7" captionText="FIGURE 4. G. s p e l a e u s n. sp., A – D paratype male (ZFMK Myr 0936), E paratype female (ISLA 3838) SEM. A: leg pairs 4 – 7 with latero-tergites; B: coxosternite 6 with stigma opening; C: coxal pouch 7; D: midbody latero-tergite of male; E: endbody latero-tergite female. Abbreviations: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number. Scale bars: A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm." httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="5" pageNumber="31">Fig.4</figureCitation>
|
||
D, E). Posterior margin with a single row of short and slender spines. Marginal rim covering more than anterior third of latero-tergite, descending in 3 steps. Surface area not covered by marginal rim (posterior 2/3) reticulated. Whole surface of latero-tergite very sparsely covered by isolated setae (
|
||
<figureCitation id="45E35BCC9C15DA230126892EFB4B6914" box="[1129,1201,548,572]" captionStart="FIGURE 4. G. s p e l a e u s n" captionStartId="7.[151,250,1694,1717]" captionTargetBox="[151,1435,188,1673]" captionTargetId="figure@7.[151,1436,188,1674]" captionTargetPageId="7" captionText="FIGURE 4. G. s p e l a e u s n. sp., A – D paratype male (ZFMK Myr 0936), E paratype female (ISLA 3838) SEM. A: leg pairs 4 – 7 with latero-tergites; B: coxosternite 6 with stigma opening; C: coxal pouch 7; D: midbody latero-tergite of male; E: endbody latero-tergite female. Abbreviations: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number. Scale bars: A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm." httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="5" pageNumber="31">Fig. 4</figureCitation>
|
||
D, E). Latero-tergite similar to one another, but last latero-tergite on posterior margin with a well-rounded indentation, providing space for the movements of the last leg.
|
||
</paragraph>
|
||
<caption id="89A717C19C15DA2305D88DA4FC316DCB" httpUri="https://zenodo.org/record/211961/files/figure.png" pageId="5" pageNumber="31" targetBox="[241,1345,733,1679]" targetPageId="5">
|
||
<paragraph id="DD6747499C15DA2305D88DA4FC316DCB" blockId="5.[151,1436,1710,1763]" pageId="5" pageNumber="31">
|
||
<emphasis id="EFAC9B5B9C15DA2305D88DA4FEEF6DEB" bold="true" box="[151,277,1710,1733]" pageId="5" pageNumber="31">FIGURE 2.</emphasis>
|
||
<taxonomicName id="1AD83CCA9C15DA2304518DA4FDE66DEC" box="[286,540,1710,1732]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis id="EFAC9B5B9C15DA2304518DA4FDE66DEC" box="[286,540,1710,1732]" italics="true" pageId="5" pageNumber="31">Glomeridesmus spelaeus</emphasis>
|
||
</taxonomicName>
|
||
, living specimens. First published pictures of living
|
||
<taxonomicName id="1AD83CCA9C15DA2301098DA4FB086DEC" box="[1094,1266,1710,1732]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glomeridesmida" pageId="5" pageNumber="31" phylum="Arthropoda" rank="order">Glomeridesmida</taxonomicName>
|
||
. arrows point to eggs and and intestine. Note the large eggs. Intestine not straight tube but folded.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89A717C19C16DA2005D88DE3FABE6CDC" httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="6" pageNumber="32" targetBox="[152,1435,193,1758]" targetPageId="6">
|
||
<paragraph id="DD6747499C16DA2005D88DE3FABE6CDC" blockId="6.[151,1436,1769,2037]" pageId="6" pageNumber="32">
|
||
<emphasis id="EFAC9B5B9C16DA2005D88DE3FEE86DD6" bold="true" box="[151,274,1769,1792]" pageId="6" pageNumber="32">FIGURE 3.</emphasis>
|
||
<emphasis id="EFAC9B5B9C16DA2004578DE0FE7D6DD7" box="[280,391,1769,1791]" italics="true" pageId="6" pageNumber="32">G. s p e l a e u s</emphasis>
|
||
<emphasis id="EFAC9B5B9C16DA2004DF8DE0FE326DD7" bold="true" box="[400,456,1770,1791]" pageId="6" pageNumber="32">n. sp.</emphasis>
|
||
, paratype female (ISLA 3837), SEM.
|
||
<emphasis id="EFAC9B5B9C16DA2006068DE3FCA16DD6" bold="true" box="[841,859,1769,1790]" pageId="6" pageNumber="32">A</emphasis>
|
||
: midbody segment, anterior (oral) view;
|
||
<emphasis id="EFAC9B5B9C16DA2001B68DE0FAF06DD7" bold="true" box="[1273,1290,1770,1791]" pageId="6" pageNumber="32">B</emphasis>
|
||
: coxosternites and prefemur of 2nd leg with ovipositor, posterior view;
|
||
<emphasis id="EFAC9B5B9C16DA2007908C0DFD0B6C36" bold="true" box="[735,753,1799,1822]" pageId="6" pageNumber="32">C</emphasis>
|
||
: head, ventral view;
|
||
<emphasis id="EFAC9B5B9C16DA2006858C02FC266C35" bold="true" box="[970,988,1800,1821]" pageId="6" pageNumber="32">D</emphasis>
|
||
: right organ of Tömösváry, lateral view;
|
||
<emphasis id="EFAC9B5B9C16DA2000CB8C02FA6F6C35" bold="true" box="[1412,1429,1800,1821]" pageId="6" pageNumber="32">E</emphasis>
|
||
: head, dorsal view;
|
||
<emphasis id="EFAC9B5B9C16DA2004138C2DFE916C14" bold="true" box="[348,363,1831,1852]" pageId="6" pageNumber="32">F</emphasis>
|
||
: antenna, antennomere 7 and disc, lateral view;
|
||
<emphasis id="EFAC9B5B9C16DA20062C8C2CFC8D6C15" bold="true" box="[867,887,1830,1853]" pageId="6" pageNumber="32">G</emphasis>
|
||
: gnathochilarium, underside.
|
||
<emphasis id="EFAC9B5B9C16DA2001E48C2CFABE6C15" bold="true" box="[1195,1348,1830,1853]" pageId="6" pageNumber="32">Abbreviations</emphasis>
|
||
: C&P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca1 = cardo, basal mandible joint 1; Ca2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres.
|
||
<emphasis id="EFAC9B5B9C16DA20047F8CD4FE5D6CDC" bold="true" box="[304,423,2014,2037]" pageId="6" pageNumber="32">Scale bars:</emphasis>
|
||
A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89A717C19C17DA2105D88D94FDFE6C07" httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="7" pageNumber="33" targetBox="[151,1435,188,1673]" targetPageId="7">
|
||
<paragraph id="DD6747499C17DA2105D88D94FDFE6C07" blockId="7.[151,1436,1694,1839]" pageId="7" pageNumber="33">
|
||
<emphasis id="EFAC9B5B9C17DA2105D88D94FEE96D9C" bold="true" box="[151,275,1694,1717]" pageId="7" pageNumber="33">FIGURE 4.</emphasis>
|
||
<emphasis id="EFAC9B5B9C17DA2104568DAAFE706D9D" box="[281,394,1695,1717]" italics="true" pageId="7" pageNumber="33">G. s p e l a e u s</emphasis>
|
||
<emphasis id="EFAC9B5B9C17DA2104DC8D95FE366D9C" bold="true" box="[403,460,1695,1716]" pageId="7" pageNumber="33">n. sp.</emphasis>
|
||
, A–D paratype male (ZFMK Myr0936), E paratype female (ISLA 3838) SEM.
|
||
<emphasis id="EFAC9B5B9C17DA2101B68D95FAF16D9C" bold="true" box="[1273,1291,1695,1716]" pageId="7" pageNumber="33">A</emphasis>
|
||
: leg pairs 4–7 with latero-tergites;
|
||
<emphasis id="EFAC9B5B9C17DA2104298DB4FE8D6DFB" bold="true" box="[358,375,1726,1747]" pageId="7" pageNumber="33">B</emphasis>
|
||
: coxosternite 6 with stigma opening;
|
||
<emphasis id="EFAC9B5B9C17DA2107B28DB7FCF56DFC" bold="true" box="[765,783,1725,1748]" pageId="7" pageNumber="33">C</emphasis>
|
||
: coxal pouch 7;
|
||
<emphasis id="EFAC9B5B9C17DA2106F38DB4FC346DFB" bold="true" box="[956,974,1726,1747]" pageId="7" pageNumber="33">D</emphasis>
|
||
: midbody latero-tergite of male;
|
||
<emphasis id="EFAC9B5B9C17DA21006A8DB4FACC6DFB" bold="true" box="[1317,1334,1726,1747]" pageId="7" pageNumber="33">E</emphasis>
|
||
: endbody latero-tergite female.
|
||
<emphasis id="EFAC9B5B9C17DA21043E8DD6FDF06DDB" bold="true" box="[369,522,1756,1779]" pageId="7" pageNumber="33">Abbreviations</emphasis>
|
||
: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number.
|
||
<emphasis id="EFAC9B5B9C17DA2101488DF0FB856C38" bold="true" box="[1031,1151,1786,1809]" pageId="7" pageNumber="33">Scale bars:</emphasis>
|
||
A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89A717C19C18DA2E05D88D66FC476DD5" httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="8" pageNumber="34" targetBox="[151,1435,190,1618]" targetPageId="8">
|
||
<paragraph id="DD6747499C18DA2E05D88D66FC476DD5" blockId="8.[151,1436,1644,1789]" pageId="8" pageNumber="34">
|
||
<emphasis id="EFAC9B5B9C18DA2E05D88D66FEEE6DAA" bold="true" box="[151,276,1644,1667]" pageId="8" pageNumber="34">FIGURE 5.</emphasis>
|
||
<taxonomicName id="1AD83CCA9C18DA2E04548D64FE756DAB" box="[283,399,1645,1667]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
|
||
<emphasis id="EFAC9B5B9C18DA2E04548D64FE756DAB" box="[283,399,1645,1667]" italics="true" pageId="8" pageNumber="34">G. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F49F26209C18DA2E04D88D67FE2B6DAA" box="[407,465,1645,1666]" pageId="8" pageNumber="34" rank="species">
|
||
<emphasis id="EFAC9B5B9C18DA2E04D88D67FE2B6DAA" bold="true" box="[407,465,1645,1666]" pageId="8" pageNumber="34">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype male (ZFMK Myr0936), SEM.
|
||
<emphasis id="EFAC9B5B9C18DA2E06CD8D67FC6E6DAA" bold="true" box="[898,916,1645,1666]" pageId="8" pageNumber="34">A</emphasis>
|
||
: right male leg 2, posterior view;
|
||
<emphasis id="EFAC9B5B9C18DA2E01BA8D67FAFC6DAA" bold="true" box="[1269,1286,1645,1666]" pageId="8" pageNumber="34">B</emphasis>
|
||
: gonopore;
|
||
<emphasis id="EFAC9B5B9C18DA2E00CC8D66FA6F6DAB" bold="true" box="[1411,1429,1644,1667]" pageId="8" pageNumber="34">C</emphasis>
|
||
: tarsus 2 with tarsal claw, apical spine and paranychium;
|
||
<emphasis id="EFAC9B5B9C18DA2E064F8D86FCE86D89" bold="true" box="[768,786,1676,1697]" pageId="8" pageNumber="34">D</emphasis>
|
||
: midbody tergite, underside;
|
||
<emphasis id="EFAC9B5B9C18DA2E011D8D86FB996D89" bold="true" box="[1106,1123,1676,1697]" pageId="8" pageNumber="34">E</emphasis>
|
||
: detail of anterior margin of underside;
|
||
<emphasis id="EFAC9B5B9C18DA2E04428DA0FEE66D97" bold="true" box="[269,284,1706,1727]" pageId="8" pageNumber="34">F</emphasis>
|
||
: detail of posterior margin of underside, 'endotergum'.
|
||
<emphasis id="EFAC9B5B9C18DA2E06338DA3FBEF6DE8" bold="true" box="[892,1045,1705,1728]" pageId="8" pageNumber="34">Abbreviations</emphasis>
|
||
: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia.
|
||
<emphasis id="EFAC9B5B9C18DA2E002B8DC2FF356DD5" bold="true" pageId="8" pageNumber="34">Scale bars:</emphasis>
|
||
A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89A717C19C19DA2F05D88DCBFAB46C3D" httpUri="https://zenodo.org/record/211966/files/figure.png" pageId="9" pageNumber="35" targetBox="[152,1435,179,1714]" targetPageId="9">
|
||
<paragraph id="DD6747499C19DA2F05D88DCBFAB46C3D" blockId="9.[151,1436,1729,1814]" pageId="9" pageNumber="35">
|
||
<emphasis id="EFAC9B5B9C19DA2F05D88DCBFEEE6DFF" bold="true" box="[151,276,1729,1752]" pageId="9" pageNumber="35">FIGURE 6.</emphasis>
|
||
<taxonomicName id="1AD83CCA9C19DA2F04538DC9FE756DF0" box="[284,399,1730,1752]" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" order="Glomeridesmida" pageId="9" pageNumber="35" phylum="Arthropoda" rank="species" species="spelaeus" status="sp. nov.">
|
||
<emphasis id="EFAC9B5B9C19DA2F04538DC9FE756DF0" box="[284,399,1730,1752]" italics="true" pageId="9" pageNumber="35">G. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="EFAC9B5B9C19DA2F04D78DC8FE286DFF" bold="true" box="[408,466,1730,1751]" pageId="9" pageNumber="35">
|
||
<taxonomicNameLabel id="F49F26209C19DA2F04D78DC8FE286DFF" box="[408,466,1730,1751]" pageId="9" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male (ZFMK Myr0936), SEM, midbody tergite.
|
||
<emphasis id="EFAC9B5B9C19DA2F017A8DC8FBBD6DFF" bold="true" box="[1077,1095,1730,1751]" pageId="9" pageNumber="35">A</emphasis>
|
||
: dorsal view;
|
||
<emphasis id="EFAC9B5B9C19DA2F01938DC8FB176DFF" bold="true" box="[1244,1261,1730,1751]" pageId="9" pageNumber="35">B</emphasis>
|
||
: lateral view;
|
||
<emphasis id="EFAC9B5B9C19DA2F00CC8DCBFA6F6DF0" bold="true" box="[1411,1429,1729,1752]" pageId="9" pageNumber="35">C</emphasis>
|
||
: sclerotized knob with setae;
|
||
<emphasis id="EFAC9B5B9C19DA2F04F08DEBFE2B6DDE" bold="true" box="[447,465,1761,1782]" pageId="9" pageNumber="35">D</emphasis>
|
||
: posterior margin, 'limbus';
|
||
<emphasis id="EFAC9B5B9C19DA2F07B98DEBFCFD6DDE" bold="true" box="[758,775,1761,1782]" pageId="9" pageNumber="35">E</emphasis>
|
||
: anterior margin.
|
||
<emphasis id="EFAC9B5B9C19DA2F068B8DEAFBA76DDF" bold="true" box="[964,1117,1760,1783]" pageId="9" pageNumber="35">Abbreviations</emphasis>
|
||
: ant = anterior; s = (sensorial) seta; str = stria; arrow points to oral side.
|
||
<emphasis id="EFAC9B5B9C19DA2F07748DF5FD486C3D" bold="true" box="[571,690,1791,1814]" pageId="9" pageNumber="35">Scale bars:</emphasis>
|
||
A = 500 µm; B = 100 µm; C = 10 µm; D = 100 µm; E = 200 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89A717C19C1ADA2C05D88DC6FD376C17" httpUri="https://zenodo.org/record/211967/files/figure.png" pageId="10" pageNumber="36" targetBox="[159,1435,177,1719]" targetPageId="10">
|
||
<paragraph id="DD6747499C1ADA2C05D88DC6FD376C17" blockId="10.[151,1436,1740,1855]" pageId="10" pageNumber="36">
|
||
<emphasis id="EFAC9B5B9C1ADA2C05D88DC6FEEF6DCA" bold="true" box="[151,277,1740,1763]" pageId="10" pageNumber="36">FIGURE 7.</emphasis>
|
||
<emphasis id="EFAC9B5B9C1ADA2C04528DC4FE6A6DCB" box="[285,400,1741,1763]" italics="true" pageId="10" pageNumber="36">G. s p e l a e u s</emphasis>
|
||
<emphasis id="EFAC9B5B9C1ADA2C04D48DC7FE2D6DCA" bold="true" box="[411,471,1741,1762]" pageId="10" pageNumber="36">n. sp.</emphasis>
|
||
, paratype male (ZFMK Myr0936), SEM.
|
||
<emphasis id="EFAC9B5B9C1ADA2C06C08DC7FC5B6DCA" bold="true" box="[911,929,1741,1762]" pageId="10" pageNumber="36">A</emphasis>
|
||
: last pair of walking legs (34) in oral view with sternite and subanal plate;
|
||
<emphasis id="EFAC9B5B9C1ADA2C04E38DE6FE476C29" bold="true" box="[428,445,1772,1793]" pageId="10" pageNumber="36">B</emphasis>
|
||
: anal shield, dorsal view, line marks part covered by last tergite (19).
|
||
<emphasis id="EFAC9B5B9C1ADA2C01D88DE1FACA6C2A" bold="true" box="[1175,1328,1771,1794]" pageId="10" pageNumber="36">Abbreviations</emphasis>
|
||
: Cx-St? = coxosternite; F = femur; poF = postfemur; prF = prefemur; s = field of small setae; S = isolated long setae, sp = spine-like claw; St = sternite; sub = subanal plate.
|
||
<emphasis id="EFAC9B5B9C1ADA2C04A18C22FDA66C16" bold="true" box="[494,604,1832,1855]" pageId="10" pageNumber="36">Scale bars</emphasis>
|
||
= 200 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD6747499C1ADA2D05888C6CFD486ADC" blockId="10.[151,1436,1894,2027]" lastBlockId="11.[151,1437,151,1378]" lastPageId="11" lastPageNumber="37" pageId="10" pageNumber="36">
|
||
<emphasis id="EFAC9B5B9C1ADA2C05888C6CFE656C56" box="[199,415,1894,1918]" italics="true" pageId="10" pageNumber="36">Trunk appendages:</emphasis>
|
||
Coxosternite(?) serrated at apico-lateral margin (
|
||
<figureCitation id="45E35BCC9C1ADA2C06838C6CFBF56C56" box="[972,1039,1894,1918]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="10" pageNumber="36">Fig. 5</figureCitation>
|
||
A, 4A). Sternal part more elevated, located basal and lateral of coxal part. Spiracle opening of anterior side in baso-lateral corner of coxosternite (
|
||
<figureCitation id="45E35BCC9C1ADA2C00208C80FF5C6CEE" captionStart="FIGURE 4. G. s p e l a e u s n" captionStartId="7.[151,250,1694,1717]" captionTargetBox="[151,1435,188,1673]" captionTargetId="figure@7.[151,1436,188,1674]" captionTargetPageId="7" captionText="FIGURE 4. G. s p e l a e u s n. sp., A – D paratype male (ZFMK Myr 0936), E paratype female (ISLA 3838) SEM. A: leg pairs 4 – 7 with latero-tergites; B: coxosternite 6 with stigma opening; C: coxal pouch 7; D: midbody latero-tergite of male; E: endbody latero-tergite female. Abbreviations: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number. Scale bars: A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm." httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="10" pageNumber="36">Fig. 4</figureCitation>
|
||
A, B). Starting at coxosternite 7, each odd-numbered coxa up to pair 29 (male) or 31 (female) apically with an eversible coxal pouch posteriorly of prefemur (
|
||
<figureCitation id="45E35BCC9C1ADA2C07E38CD8FD0F6CC2" box="[684,757,2002,2026]" captionStart="FIGURE 4. G. s p e l a e u s n" captionStartId="7.[151,250,1694,1717]" captionTargetBox="[151,1435,188,1673]" captionTargetId="figure@7.[151,1436,188,1674]" captionTargetPageId="7" captionText="FIGURE 4. G. s p e l a e u s n. sp., A – D paratype male (ZFMK Myr 0936), E paratype female (ISLA 3838) SEM. A: leg pairs 4 – 7 with latero-tergites; B: coxosternite 6 with stigma opening; C: coxal pouch 7; D: midbody latero-tergite of male; E: endbody latero-tergite female. Abbreviations: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number. Scale bars: A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm." httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="10" pageNumber="36">Figs 4</figureCitation>
|
||
A, C). Legs elongated and slender (
|
||
<figureCitation id="45E35BCC9C1ADA2C01C88CD8FB316CC2" box="[1159,1227,2002,2026]" captionStart="FIGURE 4. G. s p e l a e u s n" captionStartId="7.[151,250,1694,1717]" captionTargetBox="[151,1435,188,1673]" captionTargetId="figure@7.[151,1436,188,1674]" captionTargetPageId="7" captionText="FIGURE 4. G. s p e l a e u s n. sp., A – D paratype male (ZFMK Myr 0936), E paratype female (ISLA 3838) SEM. A: leg pairs 4 – 7 with latero-tergites; B: coxosternite 6 with stigma opening; C: coxal pouch 7; D: midbody latero-tergite of male; E: endbody latero-tergite female. Abbreviations: cp = coxal pouch; Cx-St = coxosternite; F = femur; LT = latero-tergite; poF = postfemur; prF = prefemur; st = stigma; Ta = tarsus; Ti = Tibia; numbers refer to leg pair number. Scale bars: A = 500 µm; B = 80 µm; C = 30 µm; D = 100 µm; E = 100 µm." httpUri="https://zenodo.org/record/211964/files/figure.png" pageId="10" pageNumber="36">Fig. 4</figureCitation>
|
||
A, 5A). Femur 2.5 times longer than wide, tarsus 5.5 times longer than wide. Coxa apically and mesally with few longer setae; prefemur apically, mesally and laterally with 3 or 4 longer setae; prefemur on mesal margin at midpoint with one very long seta, around 20 smaller setae oriented towards mesal margin; station of postfemur and tibia similar to prefemur (
|
||
<figureCitation id="45E35BCC9C1BDA2D04408A0EFEA96A34" box="[271,339,260,284]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="11" pageNumber="37">Fig. 5</figureCitation>
|
||
A). Tarsus with larger setae only in basal half, a pair of long spines present, tip with one apical spine, an elongated, weakly curved apical claw with basal flexible paranychium, twice as long as claw (
|
||
<figureCitation id="45E35BCC9C1BDA2D007A8A2DFA836A17" box="[1333,1401,295,319]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="11" pageNumber="37">Fig. 5</figureCitation>
|
||
C). Penultimate leg with coxosternite narrow, with stigma opening and sternal part located below instead of lateral of coxal part. Ultimate leg pair modified, consisting of coxosternite and 3 podomeres (
|
||
<figureCitation id="45E35BCC9C1BDA2D01068A65FB766AAF" box="[1097,1164,367,391]" captionStart="FIGURE 7. G. s p e l a e u s n" captionStartId="10.[151,250,1740,1763]" captionTargetBox="[159,1435,177,1719]" captionTargetId="figure@10.[151,1436,177,1719]" captionTargetPageId="10" captionText="FIGURE 7. G. s p e l a e u s n. sp., paratype male (ZFMK Myr 0936), SEM. A: last pair of walking legs (34) in oral view with sternite and subanal plate; B: anal shield, dorsal view, line marks part covered by last tergite (19). Abbreviations: Cx-St? = coxosternite; F = femur; poF = postfemur; prF = prefemur; s = field of small setae; S = isolated long setae, sp = spine-like claw; St = sternite; sub = subanal plate. Scale bars = 200 µm." httpUri="https://zenodo.org/record/211967/files/figure.png" pageId="11" pageNumber="37">Fig. 7</figureCitation>
|
||
A). Last two podomeres extending postero-laterally (confused with telopod by previous authors). Both legs widely separated from each other, connected by large plate (sternite?). Coxosternite large, almost as long as 3 podomeres combined. Podomere three apically with a large, long spine (
|
||
<figureCitation id="45E35BCC9C1BDA2D07058AD6FD746ADC" box="[586,654,476,500]" captionStart="FIGURE 7. G. s p e l a e u s n" captionStartId="10.[151,250,1740,1763]" captionTargetBox="[159,1435,177,1719]" captionTargetId="figure@10.[151,1436,177,1719]" captionTargetPageId="10" captionText="FIGURE 7. G. s p e l a e u s n. sp., paratype male (ZFMK Myr 0936), SEM. A: last pair of walking legs (34) in oral view with sternite and subanal plate; B: anal shield, dorsal view, line marks part covered by last tergite (19). Abbreviations: Cx-St? = coxosternite; F = femur; poF = postfemur; prF = prefemur; s = field of small setae; S = isolated long setae, sp = spine-like claw; St = sternite; sub = subanal plate. Scale bars = 200 µm." httpUri="https://zenodo.org/record/211967/files/figure.png" pageId="11" pageNumber="37">Fig. 7</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="DD6747499C1BDA2D0588890AFAF86914" blockId="11.[151,1437,151,1378]" pageId="11" pageNumber="37">
|
||
<emphasis id="EFAC9B5B9C1BDA2D0588890AFEBC6930" box="[199,326,512,536]" italics="true" pageId="11" pageNumber="37">Anal shield</emphasis>
|
||
: glabrous, with well-rounded edge (
|
||
<figureCitation id="45E35BCC9C1BDA2D07918AF5FCD8693F" box="[734,802,511,535]" captionStart="FIGURE 7. G. s p e l a e u s n" captionStartId="10.[151,250,1740,1763]" captionTargetBox="[159,1435,177,1719]" captionTargetId="figure@10.[151,1436,177,1719]" captionTargetPageId="10" captionText="FIGURE 7. G. s p e l a e u s n. sp., paratype male (ZFMK Myr 0936), SEM. A: last pair of walking legs (34) in oral view with sternite and subanal plate; B: anal shield, dorsal view, line marks part covered by last tergite (19). Abbreviations: Cx-St? = coxosternite; F = femur; poF = postfemur; prF = prefemur; s = field of small setae; S = isolated long setae, sp = spine-like claw; St = sternite; sub = subanal plate. Scale bars = 200 µm." httpUri="https://zenodo.org/record/211967/files/figure.png" pageId="11" pageNumber="37">Fig. 7</figureCitation>
|
||
B). On lateral side with slightly serrated margin and line of small (sensorial?) setae (
|
||
<figureCitation id="45E35BCC9C1BDA2D0485892EFDF76914" box="[458,525,548,572]" captionStart="FIGURE 7. G. s p e l a e u s n" captionStartId="10.[151,250,1740,1763]" captionTargetBox="[159,1435,177,1719]" captionTargetId="figure@10.[151,1436,177,1719]" captionTargetPageId="10" captionText="FIGURE 7. G. s p e l a e u s n. sp., paratype male (ZFMK Myr 0936), SEM. A: last pair of walking legs (34) in oral view with sternite and subanal plate; B: anal shield, dorsal view, line marks part covered by last tergite (19). Abbreviations: Cx-St? = coxosternite; F = femur; poF = postfemur; prF = prefemur; s = field of small setae; S = isolated long setae, sp = spine-like claw; St = sternite; sub = subanal plate. Scale bars = 200 µm." httpUri="https://zenodo.org/record/211967/files/figure.png" pageId="11" pageNumber="37">Fig. 7</figureCitation>
|
||
B). Subanal plate located behind last pair of legs, large and hyaline.
|
||
</paragraph>
|
||
<paragraph id="DD6747499C1BDA2D05888942FD266861" blockId="11.[151,1437,151,1378]" pageId="11" pageNumber="37">
|
||
<emphasis id="EFAC9B5B9C1BDA2D05888942FE636948" box="[199,409,584,608]" italics="true" pageId="11" pageNumber="37">Sexual characters:</emphasis>
|
||
Female: second coxae on posterior side with prominent ovipositors of medium length (
|
||
<figureCitation id="45E35BCC9C1BDA2D0021894DFF5F69A9" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="11" pageNumber="37">Fig. 3</figureCitation>
|
||
B). Coxa protruding mesally as a short lobe. Ovipositors basally supported by an undivided plate (sternite?). Ovipositors of medium length, in largest females protruding posteriorly up to leg 7. Ovipositor divided into a basal part consisting of several layers of (eversible?) half-rings, each carrying isolated setae in a regular distance to one another, and an apical part with the opening (
|
||
<figureCitation id="45E35BCC9C1BDA2D07EE89C7FD1F69CD" box="[673,741,717,741]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="11" pageNumber="37">Fig. 3</figureCitation>
|
||
B). In the basal part, half rings are anteriorly and posteriorly interrupted by two different, more slender tubes, also consisting of rings, running along the whole length of the ovipositor. Apical part complex, consisting of apparently four larger plates, each arising out of a tube of rings. All four apical plates covered by longer setae (
|
||
<figureCitation id="45E35BCC9C1BDA2D0738883BFD436861" box="[631,697,817,841]" captionStart="FIGURE 3. G. s p e l a e u s n" captionStartId="6.[151,250,1769,1792]" captionTargetBox="[152,1435,193,1758]" captionTargetId="figure@6.[151,1436,193,1762]" captionTargetPageId="6" captionText="FIGURE 3. G. s p e l a e u s n. sp., paratype female (ISLA 3837), SEM. A: midbody segment, anterior (oral) view; B: coxosternites and prefemur of 2 nd leg with ovipositor, posterior view; C: head, ventral view; D: right organ of Tömösváry, lateral view; E: head, dorsal view; F: antenna, antennomere 7 and disc, lateral view; G: gnathochilarium, underside. Abbreviations: C & P = tarsal claw and paranychium; Ca = cardines of gnathochilarium; Ca 1 = cardo, basal mandible joint 1; Ca 2? = either cardo (joint 1) or stipites (joint 2) of mandible; cP = central pads, modified central palpi; Cx-St = coxosternite; F = femur; Gn = gnathochilarium; iP = inner palpi; La = labrum; LL = lamella linguales; LP = lateral palpi; LT = latero-tergite; Me = mentum; O = ovipositor; Pl = plate covering organ of TO; poF = postfemur; prF = prefemur; St = stipes of gnathochilarium; sc = apical cones; St? = sternite?; T = tergite; Ta = tarsus; Ti = Tibia; TO = organ of Tömösváry; roman numerals refer to number of antennomeres. Scale bars: A = 200 µm; B = 100 µm; C = 200 µm; D = 20 µm; E = 200 µm; F = 20 µm; G = 100 µm." httpUri="https://zenodo.org/record/211962/files/figure.png" pageId="11" pageNumber="37">Fig. 3</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="DD6747499C1BDA2D05888858FCA86F4E" blockId="11.[151,1437,151,1378]" pageId="11" pageNumber="37">
|
||
Male: second coxa with gonopore located mesally (
|
||
<figureCitation id="45E35BCC9C1BDA2D06648858FC886842" box="[811,882,850,874]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="11" pageNumber="37">Fig. 5</figureCitation>
|
||
A, B). Gonopore consisting of two sclerotized, apically intertwined plates forming a slit-like opening in basal half (
|
||
<figureCitation id="45E35BCC9C1BDA2D06DE887CFC2E68A6" box="[913,980,886,910]" captionStart="FIGURE 5. G" captionStartId="8.[151,250,1644,1667]" captionTargetBox="[151,1435,190,1618]" captionTargetId="figure@8.[151,1436,190,1618]" captionTargetPageId="8" captionText="FIGURE 5. G. spelaeus n. sp., paratype male (ZFMK Myr 0936), SEM. A: right male leg 2, posterior view; B: gonopore; C: tarsus 2 with tarsal claw, apical spine and paranychium; D: midbody tergite, underside; E: detail of anterior margin of underside; F: detail of posterior margin of underside, ' endotergum'. Abbreviations: Apo = tracheal apodeme; Cx-St = coxosternite; F = femur; Go = gonopore; Par = paranychium; poF = postfemur; prF = prefemur; Ta = tarsus; Ti = Tibia. Scale bars: A = 300 µm; B = 30 µm; C = 60 µm; D = 200 µm; E = 20 µm; F = 8 µm." httpUri="https://zenodo.org/record/211965/files/figure.png" pageId="11" pageNumber="37">Fig. 5</figureCitation>
|
||
B). Apical part of both plates with 2 or 3 longer setae. Male telopod consisting of syncoxite with inner horns and 4 podomeres (
|
||
<figureCitation id="45E35BCC9C1BDA2D01FE8890FB00689A" box="[1201,1274,922,946]" captionStart="FIGURE 8. G. s p e l a e u s n" captionStartId="12.[151,250,1570,1593]" captionTargetBox="[151,1435,361,1548]" captionTargetId="figure@12.[151,1436,361,1549]" captionTargetPageId="12" captionText="FIGURE 8. G. s p e l a e u s n. sp., paratype males, SEM, telopod, anterior view. A: left telopod with syncoxite and inner horns (ZFMK Myr 0936); B: right telopod, podomeres 2 – 4, (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936). Abbreviations: IH = inner horns; mem = membranous area; roman numerals refer to podomere number. Scale bars: A = 300 µm; B = 30 µm; C = 50 µm." httpUri="https://zenodo.org/record/211968/files/figure.png" pageId="11" pageNumber="37">Fig. 8</figureCitation>
|
||
A). Syncoxite engulfing the basal podomeres laterally and in anterior view, medially rising into a smooth glabrous process (
|
||
<figureCitation id="45E35BCC9C1BDA2D002088B4FF5F68D2" captionStart="FIGURE 8. G. s p e l a e u s n" captionStartId="12.[151,250,1570,1593]" captionTargetBox="[151,1435,361,1548]" captionTargetId="figure@12.[151,1436,361,1549]" captionTargetPageId="12" captionText="FIGURE 8. G. s p e l a e u s n. sp., paratype males, SEM, telopod, anterior view. A: left telopod with syncoxite and inner horns (ZFMK Myr 0936); B: right telopod, podomeres 2 – 4, (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936). Abbreviations: IH = inner horns; mem = membranous area; roman numerals refer to podomere number. Scale bars: A = 300 µm; B = 30 µm; C = 50 µm." httpUri="https://zenodo.org/record/211968/files/figure.png" pageId="11" pageNumber="37">Fig. 8</figureCitation>
|
||
A). Process apically with two long inner horns; horns basally only divided by suture, more apically completely separated, diverging and later overlapping one another (
|
||
<figureCitation id="45E35BCC9C1BDA2D06698F0CFC976F36" box="[806,877,1030,1054]" captionStart="FIGURE 8. G. s p e l a e u s n" captionStartId="12.[151,250,1570,1593]" captionTargetBox="[151,1435,361,1548]" captionTargetId="figure@12.[151,1436,361,1549]" captionTargetPageId="12" captionText="FIGURE 8. G. s p e l a e u s n. sp., paratype males, SEM, telopod, anterior view. A: left telopod with syncoxite and inner horns (ZFMK Myr 0936); B: right telopod, podomeres 2 – 4, (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936). Abbreviations: IH = inner horns; mem = membranous area; roman numerals refer to podomere number. Scale bars: A = 300 µm; B = 30 µm; C = 50 µm." httpUri="https://zenodo.org/record/211968/files/figure.png" pageId="11" pageNumber="37">Fig. 8</figureCitation>
|
||
A). Each horn apically consisting of sclerotized lateral margins that work like a frame holding more membranous folds covering the median part of the tip (
|
||
<figureCitation id="45E35BCC9C1BDA2D00208F20FF596F4E" captionStart="FIGURE 8. G. s p e l a e u s n" captionStartId="12.[151,250,1570,1593]" captionTargetBox="[151,1435,361,1548]" captionTargetId="figure@12.[151,1436,361,1549]" captionTargetPageId="12" captionText="FIGURE 8. G. s p e l a e u s n. sp., paratype males, SEM, telopod, anterior view. A: left telopod with syncoxite and inner horns (ZFMK Myr 0936); B: right telopod, podomeres 2 – 4, (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936). Abbreviations: IH = inner horns; mem = membranous area; roman numerals refer to podomere number. Scale bars: A = 300 µm; B = 30 µm; C = 50 µm." httpUri="https://zenodo.org/record/211968/files/figure.png" pageId="11" pageNumber="37">Fig. 8</figureCitation>
|
||
A).Whole surface of horns covered by isolated, minute setae.
|
||
</paragraph>
|
||
<paragraph id="DD6747499C1BDA2D05888F78FB3F6E4A" blockId="11.[151,1437,151,1378]" pageId="11" pageNumber="37">
|
||
Podomere 1 largest and most massive. Podomere 2 basally wide, apically tapering, mesally with a large, swollen, membranous area (
|
||
<figureCitation id="45E35BCC9C1BDA2D04948F9CFDE56F86" box="[475,543,1174,1198]" captionStart="FIGURE 8. G. s p e l a e u s n" captionStartId="12.[151,250,1570,1593]" captionTargetBox="[151,1435,361,1548]" captionTargetId="figure@12.[151,1436,361,1549]" captionTargetPageId="12" captionText="FIGURE 8. G. s p e l a e u s n. sp., paratype males, SEM, telopod, anterior view. A: left telopod with syncoxite and inner horns (ZFMK Myr 0936); B: right telopod, podomeres 2 – 4, (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936). Abbreviations: IH = inner horns; mem = membranous area; roman numerals refer to podomere number. Scale bars: A = 300 µm; B = 30 µm; C = 50 µm." httpUri="https://zenodo.org/record/211968/files/figure.png" pageId="11" pageNumber="37">Fig. 8</figureCitation>
|
||
A, B, 9A, B). Membranous area well rounded, in anterior view consisting of several inverse membranous folds (
|
||
<figureCitation id="45E35BCC9C1BDA2D07688FB0FD906FFA" box="[551,618,1210,1234]" captionStart="FIGURE 8. G. s p e l a e u s n" captionStartId="12.[151,250,1570,1593]" captionTargetBox="[151,1435,361,1548]" captionTargetId="figure@12.[151,1436,361,1549]" captionTargetPageId="12" captionText="FIGURE 8. G. s p e l a e u s n. sp., paratype males, SEM, telopod, anterior view. A: left telopod with syncoxite and inner horns (ZFMK Myr 0936); B: right telopod, podomeres 2 – 4, (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936). Abbreviations: IH = inner horns; mem = membranous area; roman numerals refer to podomere number. Scale bars: A = 300 µm; B = 30 µm; C = 50 µm." httpUri="https://zenodo.org/record/211968/files/figure.png" pageId="11" pageNumber="37">Fig. 8</figureCitation>
|
||
C), in posterior aspect with scale like and well-rounded sclerotized teeth (
|
||
<figureCitation id="45E35BCC9C1BDA2D05D08FD4FF196FDE" box="[159,227,1246,1270]" captionStart="FIGURE 9. G. s p e l a e u s n" captionStartId="13.[151,250,1753,1776]" captionTargetBox="[151,1435,527,1737]" captionTargetId="figure@13.[151,1436,527,1738]" captionTargetPageId="13" captionText="FIGURE 9. G. s p e l a e u s n. sp., paratype males, SEM, telopod, posterior view. A: right telopod, podomeres 2 – 4 (ZFMK Myr 0936); B: left telopod, podomeres 2 – 4 (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936); D: right telopod, detail of podomere 3 & 4 (ZFMK Myr 0936). Abbreviations: mem = membranous area; pro = process of podomere 3; roman numerals refer to podomere number. Scale bars: A = 100 µm; B = 100 µm; C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/211969/files/figure.png" pageId="11" pageNumber="37">Fig. 9</figureCitation>
|
||
C). Podomere 3 long and slender, 2 times longer than wide, in anterior view smooth and glabrous (
|
||
<figureCitation id="45E35BCC9C1BDA2D00718FD4FA7A6FDE" box="[1342,1408,1246,1270]" captionStart="FIGURE 8. G. s p e l a e u s n" captionStartId="12.[151,250,1570,1593]" captionTargetBox="[151,1435,361,1548]" captionTargetId="figure@12.[151,1436,361,1549]" captionTargetPageId="12" captionText="FIGURE 8. G. s p e l a e u s n. sp., paratype males, SEM, telopod, anterior view. A: left telopod with syncoxite and inner horns (ZFMK Myr 0936); B: right telopod, podomeres 2 – 4, (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936). Abbreviations: IH = inner horns; mem = membranous area; roman numerals refer to podomere number. Scale bars: A = 300 µm; B = 30 µm; C = 50 µm." httpUri="https://zenodo.org/record/211968/files/figure.png" pageId="11" pageNumber="37">Fig. 8</figureCitation>
|
||
A, B). Posterior aspect of podomere
|
||
<quantity id="1A20EAAC9C1BDA2D07628E08FDA76E32" box="[557,605,1282,1306]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="11" pageNumber="37" unit="in" value="3.0">3 in</quantity>
|
||
apical part with slender, finger-shaped and well-rounded process, the immovable finger (
|
||
<figureCitation id="45E35BCC9C1BDA2D043E8E2CFE4F6E16" box="[369,437,1318,1342]" captionStart="FIGURE 9. G. s p e l a e u s n" captionStartId="13.[151,250,1753,1776]" captionTargetBox="[151,1435,527,1737]" captionTargetId="figure@13.[151,1436,527,1738]" captionTargetPageId="13" captionText="FIGURE 9. G. s p e l a e u s n. sp., paratype males, SEM, telopod, posterior view. A: right telopod, podomeres 2 – 4 (ZFMK Myr 0936); B: left telopod, podomeres 2 – 4 (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936); D: right telopod, detail of podomere 3 & 4 (ZFMK Myr 0936). Abbreviations: mem = membranous area; pro = process of podomere 3; roman numerals refer to podomere number. Scale bars: A = 100 µm; B = 100 µm; C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/211969/files/figure.png" pageId="11" pageNumber="37">Fig. 9</figureCitation>
|
||
A, B, D). Immovable finger protruding up to 3/4 of length of podomere 4, the movable finger (
|
||
<figureCitation id="45E35BCC9C1BDA2D05A68E40FED76E4A" box="[233,301,1354,1378]" captionStart="FIGURE 9. G. s p e l a e u s n" captionStartId="13.[151,250,1753,1776]" captionTargetBox="[151,1435,527,1737]" captionTargetId="figure@13.[151,1436,527,1738]" captionTargetPageId="13" captionText="FIGURE 9. G. s p e l a e u s n. sp., paratype males, SEM, telopod, posterior view. A: right telopod, podomeres 2 – 4 (ZFMK Myr 0936); B: left telopod, podomeres 2 – 4 (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936); D: right telopod, detail of podomere 3 & 4 (ZFMK Myr 0936). Abbreviations: mem = membranous area; pro = process of podomere 3; roman numerals refer to podomere number. Scale bars: A = 100 µm; B = 100 µm; C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/211969/files/figure.png" pageId="11" pageNumber="37">Fig. 9</figureCitation>
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D). Movable finger slender and glabrous, at apex with single long spine (
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<figureCitation id="45E35BCC9C1BDA2D01118E40FB5B6E4A" box="[1118,1185,1354,1378]" captionStart="FIGURE 9. G. s p e l a e u s n" captionStartId="13.[151,250,1753,1776]" captionTargetBox="[151,1435,527,1737]" captionTargetId="figure@13.[151,1436,527,1738]" captionTargetPageId="13" captionText="FIGURE 9. G. s p e l a e u s n. sp., paratype males, SEM, telopod, posterior view. A: right telopod, podomeres 2 – 4 (ZFMK Myr 0936); B: left telopod, podomeres 2 – 4 (ISLA 3838); C: right telopod, podomere 2, detail of swollen membranous area (ZFMK Myr 0936); D: right telopod, detail of podomere 3 & 4 (ZFMK Myr 0936). Abbreviations: mem = membranous area; pro = process of podomere 3; roman numerals refer to podomere number. Scale bars: A = 100 µm; B = 100 µm; C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/211969/files/figure.png" pageId="11" pageNumber="37">Fig. 9</figureCitation>
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D).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |