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<document id="754D6F76419EB178DAF8FAB3C627EBD8" ENCODING="UTF-8" ID-GBIF-Dataset="913a65fa-131f-40af-b8d4-786873482385" ModsDocAuthor="Mueller, U. G." ModsDocID="21361" checkinTime="1244213718300" checkinUser="thomas" docAuthor="Rabeling, Ch., Cover, S. P., Johnson, R. A. &amp; Mueller, U. G." docDate="2007" docId="1DC94E957608D014A3764553992F8457" docLanguage="en" docName="21361_gg2.xml" docOrigin="Zootaxa 1664" docSource="http://www.antbase.org/ants/publications/21361/21361.pdf" docTitle="Trachymyrmex turrifex" docType="treatment" docVersion="12" lastPageNumber="24" masterDocId="DC6B9C504F79C1B58750E8E5B0F9A502" masterDocTitle="A review of the North American species of the fungus-gardening ant genus Trachymyrmex (Hymenoptera: Formicidae)." masterLastPageNumber="53" masterPageNumber="1" pageNumber="22" updateTime="1701311538475" updateUser="plazi">
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<mods:title id="AA5730E6FA9A69FF3DA536ED7621C0FF">A review of the North American species of the fungus-gardening ant genus Trachymyrmex (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="A48033F47B4824BFDFA1863AA7AFC1EF">Cover, S. P.</mods:namePart>
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<treatment id="1DC94E957608D014A3764553992F8457" ID-DOI="http://doi.org/10.5281/zenodo.6241408" ID-GBIF-Taxon="100125548" ID-Zenodo-Dep="6241408" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457" httpUri="http://treatment.plazi.org/id/1DC94E957608D014A3764553992F8457" lastPageNumber="24" pageNumber="22">
<subSubSection id="592FFCCBAAB05D3564A865C594AB8368" type="nomenclature">
<paragraph id="789721FABA81CB260A2262ECBA7D48D2" pageNumber="22">
<taxonomicName id="A4FA204D31E5317C00B107EC7D2C6521" ID-CoL="7CKT6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1903" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="22" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex (Wheeler)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7A2B2341323DFEF45C24D0F41F00CE38" type="reference_group">
<paragraph id="F2395B0FEA9BA85CB9EE24B3141C1B70" pageNumber="22">
<taxonomicName id="25902474ED7926224D0822BF5E1E40D6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145468" authority="Wheeler" authorityName="Wheeler" authorityYear="1903" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Atta (Trachymyrmex) turrifex Wheeler" order="Hymenoptera" pageNumber="22" phylum="Arthropoda" rank="species" species="turrifex" subGenus="Trachymyrmex">Atta (Trachymyrmex) turrifex Wheeler</taxonomicName>
, 1903: 100.
<typeStatus id="98E0FDE9AC7B3189EC1E07D14A3DDA91">Syntype</typeStatus>
workers and queens,
<materialsCitation id="11C46B30A33272783AF668F43AF35967" ID-GBIF-Occurrence="923899384" collectionCode="AMNH, MCZC, USNM" country="U. S. A." county="Presidio County" location="Marfa" stateProvince="Texas" typeStatus="Syntypes">
<location id="2B362856F9E40EFB4825C6866621A16C" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:2B362856F9E40EFB4825C6866621A16C" county="Presidio County" name="Marfa">Marfa</location>
,
<collectingCounty id="22BE4D09A9584D5E38013831B0C2EBCF">Presidio County</collectingCounty>
</materialsCitation>
;
<materialsCitation id="976C415A68A539EED5DB32B7111AFCD3" ID-GBIF-Occurrence="923899386" collectionCode="AMNH, MCZC, USNM" country="U. S. A." county="Pecos County" location="Fort Stockton" stateProvince="Texas" typeStatus="Syntypes">
<location id="338DA4E7EE048164247AA8AB206BFEE2" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:338DA4E7EE048164247AA8AB206BFEE2" county="Pecos County" name="Fort Stockton">Fort Stockton</location>
,
<collectingCounty id="7A2F895D4BABF0E646DE070FC52F93CA">Pecos County</collectingCounty>
</materialsCitation>
;
<materialsCitation id="315FA16D71A44CC05A42CED01ED8ED21" ID-GBIF-Occurrence="923899383" collectionCode="AMNH, MCZC, USNM" country="U. S. A." county="Valverde County" location="Del Rio and Langtry" stateProvince="Texas" typeStatus="Syntypes">
<location id="223545151DB36EAE1A66CC281C0CFCC5" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:223545151DB36EAE1A66CC281C0CFCC5" county="Valverde County" name="Del Rio and Langtry">Del Rio and Langtry</location>
,
<collectingCounty id="279A6C5AC232DCD4691C2CEA64710E83">Valverde County</collectingCounty>
</materialsCitation>
;
<materialsCitation id="C4D601612CFE4A5A1217F6A67EF35BE8" ID-GBIF-Occurrence="923899396" collectionCode="AMNH, MCZC, USNM" country="U. S. A." county="Burnet County" location="Marble Falls" stateProvince="Texas" typeStatus="Syntypes">
<location id="BCE92AE9F761B2A964009DC05D916B2F" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:BCE92AE9F761B2A964009DC05D916B2F" county="Burnet County" name="Marble Falls">Marble Falls</location>
,
<collectingCounty id="6399702F0801004D77D0BE5DB7B14A51">Burnet County</collectingCounty>
</materialsCitation>
;
<materialsCitation id="A153D733AFD4707A3F6B383BDBF3F743" ID-GBIF-Occurrence="923899380" collectionCode="AMNH, MCZC, USNM" country="U. S. A." county="Travis County" location="Austin" stateProvince="Texas" typeStatus="Syntypes">
<location id="E2C75E5F2B172437B4812E70E28D8650" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:E2C75E5F2B172437B4812E70E28D8650" county="Travis County" name="Austin">Austin</location>
,
<collectingCounty id="6D4A72958F4C902DC0B3AC046927D797">Travis County</collectingCounty>
</materialsCitation>
;
<collectingRegion id="D0FE42A16F36BC19B7F9EE76413CECB2">Texas</collectingRegion>
,
<collectingCountry id="71BC110DBDDF82DE7E559726BA022F7B">U.S.A.</collectingCountry>
(
<collectionCode id="5FC3E55BB99D4445B1BC449B6ED13AFB" collectionName="USA, New York, New York, American Museum of Natural History">AMNH</collectionCode>
,
<collectionCode id="F9293D9FE823B958714F54B5C313F77A" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
,
<collectionCode id="0D13A1866C88DC649AD2D0BF74B31AAF" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
) [examined]
</paragraph>
<paragraph id="79B037B54065159F9769C469EE135D1E" pageNumber="23">
<taxonomicName id="7DF5BE9618E31D721099809F4088F590" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" authority="Wheeler" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">Trachymyrmex turrifex Wheeler</taxonomicName>
; Gallardo 1916: 242 [Combination in
<taxonomicName id="3824898400DB456B268C8B48F0562BB3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2397" authorityName="Forel" authorityYear="1893" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="genus">Trachymyrmex</taxonomicName>
]
</paragraph>
<paragraph id="0A0F6359F0F718C700993CEEFB3012E0" pageNumber="23">
<taxonomicName id="DC21776AEED385B0D1134D3FC240BE03" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145467" authority="Wheeler" authorityName="Wheeler" authorityYear="1911" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Atta (Trachymyrmex) turrifex subsp. caroli Wheeler" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="turrifex" subGenus="Trachymyrmex" subSpecies="caroli">Atta (Trachymyrmex) turrifex subsp. caroli Wheeler</taxonomicName>
, 1911: 248.
<materialsCitation id="7831A1D87C250013A557CF7EC0AE7DA5" ID-GBIF-Occurrence="923899377" country="U. S. A." county="Walker County" location="Huntsville" stateProvince="Texas" typeStatus="Syntypes">
<typeStatus id="7514C8545EE94F1D9B205FCD37B3F91E">Syntype</typeStatus>
workers,
<location id="6BF11C3B0B90F66254218261392233E8" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:6BF11C3B0B90F66254218261392233E8" country="U. S. A." county="Walker County" name="Huntsville" stateProvince="Texas">Huntsville</location>
,
<collectingCounty id="705A671E3A346B9EC0E5759CB986D019">Walker County</collectingCounty>
,
<collectingRegion id="0432C2A96FD18A219BE8F80792B51EFC">Texas</collectingRegion>
,
<collectingCountry id="00485CF3847DF4DD2E02C32D9FF3D0D3">U.S.A.</collectingCountry>
(repository unknown)
</materialsCitation>
[not examined]
<taxonomicNameLabel id="2AC02A2C6FAC581A350BBC635C8E54BB">syn. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="7441A4B4E36EBBC805CC1C6FAE0939B3" pageNumber="23">
<taxonomicName id="14DCFC927FBD577D422D0F0FC2C221CA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152435" authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex caroli (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="turrifex" subSpecies="caroli">Trachymyrmex turrifex caroli (Wheeler)</taxonomicName>
, Creighton 1950: 324 [Combination in
<taxonomicName id="01C75755A6C4990FBCA98B6C2E21186C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2397" authorityName="Forel" authorityYear="1893" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="genus">Trachymyrmex</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="74CE01503B304EDB62076669AB495CDA" type="diagnosis">
<paragraph id="7E8FB142D18326A74259DFE12F4EF1BE" pageNumber="23">Diagnosis</paragraph>
<paragraph id="DC7A3F611D6DF22E12382BB933EE82EC" pageNumber="23">
Worker: HL 0.88-1.0, HW 0.8-0.96, CI 91-96, SL 0.72-0.8, SI 83-91, ML 1.2-1.4. A medium sized, species (HL 0.88-1.0, HW 0.8-0.96) with the shortest antennal scapes (SI 83-91) relative to head width of all US
<taxonomicName id="8B9170C2A425624EBD4DB7E863CF723C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2397" authorityName="Forel" authorityYear="1893" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="genus">Trachymyrmex</taxonomicName>
species. The scapes reach or pass the posterior corner of the head by half its maximum diameter at most. Head slightly longer than broad (CI 91-96) with its maximum width close to the posterior border of head, tapering gradually from point of broadest width to mandibular insertion. Posterior margin moderately concave. Both preocular and frontal carinae long and subparallel, reaching back towards the posterior margin of head, forming well-developed antennal scrobes. In full face view, frontal lobes broad, rounded, describing a half circle. Anterolateral promesonotal teeth short, thin, in dorsal view sharply pointed, directed anterolaterally and upwards. Bases of median pronotal teeth fused, but the peaks of each tooth still distinguishable. Propodeal teeth pointed, almost twice as long as the distance between their bases. Dorsal body surface strongly tuberculate, tuberculi well developed, on first gastric tergite, sometimes connected through ridges. Tuberculi bearing long, dark, strongly recurved setae. Side of mesosoma less tuberculate, tuberculi smaller. Color is variable from yellowish brown to medium reddish brown.
</paragraph>
<paragraph id="D0F0B82CFCCE29D7E318305111401C93" pageNumber="23">Queen: HL 1.1-1.2, HW 1.1-1.2, CI 100, SL 0.85-1.0, SI 71-87, ML 1.7-1.8. As in the worker diagnosis, except for case-specific structures of the mesosoma related to wing-bearing. Dorsoventral pronotal teeth pointed, broadly triangular in dorsal view; ventrolateral pronotal teeth small, lobelike. Mesoscutum with moderately abundant, short, coarse, longitudinal rugulae and abundant small tuberculi, each bearing a stiff, more or less recurved suberect or decumbent seta. Rear border of scutellum projecting posteriorly as two lobelike teeth. First gastric tergite covered with small tubercles interconnected by a network of fine rugulae forming an almost geometric pattern; each tubercle bearing a stiff, recurved, decumbent setae.</paragraph>
<paragraph id="B11A0D5227D1B90856FA81942080A808" pageNumber="23">
Male: HL 0.7-0.75, HW 0.6, CI 80-86, SL 0.7-0.75, SI 117-125, ML 1.5-1.6. A distinctive
<taxonomicName id="0F003B64C64F02A2FC8A5BB04E87EC32" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2397" authorityName="Forel" authorityYear="1893" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="genus">Trachymyrmex</taxonomicName>
male easily recognized by the general lack of tuberculi on all body surfaces. In addition, the rear border of the antennal scrobe is poorly defined or absent. Dorsoventral pronotal teeth small, triangular, usually sharply pointed; ventrolateral pronotal teeth absent. Mesoscutum sparsely to moderately longitudinally rugulose, interrugal spaces finely granulate. In dorsal view, rear border of scutellum forming two triangular teeth that project posteriorly. First gastric tergite finely granulate with scattered punctures bearing fine, appressed recurved setae.
</paragraph>
</subSubSection>
<subSubSection id="EBDE48123A87FBA99926F599FD514599" type="discussion">
<paragraph id="80C9D3EAFA43C6C773CE12A4EEB36545" pageNumber="23">Discussion</paragraph>
<paragraph id="2B0D951E9C54BC4FDC3F8AE796AF6D7B" pageNumber="23">
<taxonomicName id="644C4DAD6A02BFECF254D76BEE4FEC99" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">Trachymyrmex turrifex</taxonomicName>
is distinguished from other North American species by its well-developed antennal scrobes. Only the allopatric
<taxonomicName id="3F5FDDF37D8894E946AB6351F0008714" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37414" baseAuthorityName="Andre" baseAuthorityYear="1893" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex jamaicensis (André)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="jamaicensis">T. jamaicensis</taxonomicName>
has comparably developed antennal scrobes, but it may be easily distinguished from
<taxonomicName id="22FF5492B69B22282D8CA2D52675619B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
by its larger size, reddish-black coloration and shorter clypeal pilosity (as described in the key).
</paragraph>
<paragraph id="BB2D1CD10DE3D6B6DD193130F70D9C61" lastPageNumber="24" pageNumber="23">
<taxonomicName id="5A60AC4F108BA005AFBA766890BD708C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">Trachymyrmex turrifex</taxonomicName>
is distributed throughout Texas, northeastern Mexico, western Louisiana, and barely reaches southern Oklahoma. Individuals are somewhat variable in size, and color varies from light yellowish brown to dark reddish-brown. Wheeler (1903) described
<taxonomicName id="437D51218CC822CA23D595FEA87AC0BE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">turrifex</taxonomicName>
from several localities in Central Texas, and
<taxonomicName id="95850152B86B278071C5A7816C302DCF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152435" baseAuthorityName="Wheeler" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex caroli (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="turrifex" subSpecies="caroli">T. turrifex caroli</taxonomicName>
as a subspecies of
<taxonomicName id="D2174CCD5A726B8DD1187A01553200A2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
from Huntsville, Texas (1911). Wheeler 's reasoning was that
<taxonomicName id="37ACD2A0C0A1480DC60D6373C33A7A23" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152435" baseAuthorityName="Wheeler" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex caroli (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="turrifex" subSpecies="caroli">caroli</taxonomicName>
&quot;represents a depauperate, arenicolous race ranging considerably eastwards of the typical
<taxonomicName id="6897DAAD8A20DF40105CDCE788E05A66" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">turrifex</taxonomicName>
&quot; (Wheeler 1911, p. 248). Wheeler distinguished between these two &quot;subspecies&quot; based on the smaller body size, brownish yellow coloration, smaller tubercles, and lack of longitudinal impression on first gastric tergite in
<taxonomicName id="17D2F4B89016C199143AECAFD29FAFEA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152435" baseAuthorityName="Wheeler" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex caroli (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="turrifex" subSpecies="caroli">T. turrifex caroli</taxonomicName>
. In our experience, all of these character states fall well within the variation observed within and among colonies of typical
<taxonomicName id="B480BDE1754F3B7BD6AC9F68115A4004" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="turrifex">turrifex</taxonomicName>
, as seen throughout its known geographic range, which now extends to western Louisiana and places Huntsville towards the center of
<taxonomicName id="4D422D77A5E4BC8BE61010E1D1671967" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
's distribution (Figure 24C). Since we were not able to locate Wheeler 's type specimens of
<taxonomicName id="24490A39CFE509B4C7F9C5ECA71921D9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152435" baseAuthorityName="Wheeler" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex caroli (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="turrifex" subSpecies="caroli">T. turrifex caroli</taxonomicName>
, we collected fresh material at the type locality in Huntsville, Texas. These specimens were morphologically not different from other examined
<taxonomicName id="AD2868F61E0B2AC6C53F581399D4B002" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">turrifex</taxonomicName>
. Lastly, our DNA sequence information indicates that
<taxonomicName id="ED755592BD6844BD2D67B42AB6A5826A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152435" baseAuthorityName="Wheeler" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex caroli (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="turrifex" subSpecies="caroli">caroli</taxonomicName>
is also genetically not distinct from
<taxonomicName id="7922076F7721D1AB6A376CD0B355A48C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">turrifex</taxonomicName>
(Figure 21). We therefore synonymize
<taxonomicName id="83A2423A672AE7477CA4E1CACB0D1322" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152435" baseAuthorityName="Wheeler" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex caroli (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="turrifex" subSpecies="caroli">T. turrifex caroli</taxonomicName>
with
<taxonomicName id="BED1C41E19057D1DBBF73D463D74307D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="009CE2559383E366F039880719565A17" type="etymology">
<paragraph id="E870B5241C91DB8020E355693BCD3038" pageNumber="24">Etymology</paragraph>
<paragraph id="FD6E7438BDFD030FB0D8C855ACED60CA" pageNumber="24">
The epithet for
<taxonomicName id="FE5FE466D44B82BDA095F211C2F9064D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
is derived from its characteristic turretlike nest entrances, which are built from soil particles and leaf litter debris.
</paragraph>
</subSubSection>
<subSubSection id="5C73F7D23087319F0FB3CBFBB517EBEB" type="biology_ecology">
<paragraph id="19BA017596199E4256AA4D21868947BB" pageNumber="24">Biology</paragraph>
</subSubSection>
<subSubSection id="846C1D0B9DA8AE5851CD5CBE7369BC1A" type="distribution">
<paragraph id="87EBAC6745F1C608B69BA2A447F0B133" pageNumber="24">
The geographic range of
<taxonomicName id="8C8F0B24BBB2A7E88C48767B4B3457A7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
is centered in Texas and extends into southern Oklahoma, western Louisiana, and south into the Mexican States of Chihuahua and Tamaulipas (Figure 24C).
</paragraph>
</subSubSection>
<subSubSection id="1F37D9C2218C7E91208D6EF4A679E557" type="biology_ecology">
<paragraph id="D999811E60F2C0E511BD51CF71FADD57" pageNumber="24">
<taxonomicName id="4457CF594D18E1EB472065D8D03A73FF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
is abundant in open desert habitats of west Texas, the black clay soils of central Texas and is somewhat less abundant in the sandy soils of eastern Texas and western Louisiana. Dense populations occur in southern Texas, particularly in the Rio Grande Valley. When occurring sympatrically in sandy soils with
<taxonomicName id="ABE0CF18A0E3FD56F08A67B6C0DE6026" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37429" baseAuthorityName="McCook" baseAuthorityYear="1881" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex septentrionalis (McCook)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="septentrionalis">T. septentrionalis</taxonomicName>
,
<taxonomicName id="21053A2FA0E8F1B2E642A2AA9402AA7A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">T. turrifex</taxonomicName>
is less abundant, but the reverse pattern occurs in black clay soils where
<taxonomicName id="1E6628B16A5F992F0223EE2E27194C06" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">turrifex</taxonomicName>
is usually more common than
<taxonomicName id="8B8FDD5A382A12154B059175544FACCF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37429" baseAuthorityName="McCook" baseAuthorityYear="1881" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex septentrionalis (McCook)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="septentrionalis">septentrionalis</taxonomicName>
. The characteristic turretlike nest entrances, which are built from soil particles and plant litter debris during the wetter parts of the year, are a diagnostic character for
<taxonomicName id="16EAD06C26D347D70B694DD63D6A53A4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:37434" class="Insecta" family="Formicidae" genus="Trachymyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Trachymyrmex turrifex (Wheeler)" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="turrifex">turrifex</taxonomicName>
in the field. Turrets are absent during winter dormancy and are reduced or absent in summer, particularly after a period of drought. Nests consist of 1-5 chambers, which are connected by vertical tunnels. Colonies are monogynous and contain up to 300 workers.
</paragraph>
</subSubSection>
<subSubSection id="1F6A59E240962DD12C6C11175B3C5801" type="materials_examined">
<paragraph id="A6FD9C11DA6E5CA66DF810977A886729" pageNumber="24">
Additional material examined:
<materialsCitation id="06C48FA78B21033C553F4E670F786D50" ID-GBIF-Occurrence="923899391" collectorName="WF Buren" country="U. S. A." county="Beauregard Parish" location="De Ridder" stateProvince="Louisiana">
<collectingCountry id="5E0F521145565C6BEA79C93F889233FB">U.S.A.</collectingCountry>
:
<collectingRegion id="F51617415DAD1A409EDFD8558A338678">Louisiana</collectingRegion>
,
<collectingCounty id="0C835FE285D977EEF3F774253B88AD81">Beauregard Parish</collectingCounty>
:
<location id="D2B354E205C0E35FBD146D0B0E1452FA" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:D2B354E205C0E35FBD146D0B0E1452FA" country="U. S. A." name="De Ridder" stateProvince="Louisiana">De Ridder</location>
(
<collectorName id="77C3696ABB9F6F6D142CD0CEA8B41EE2">WF Buren</collectorName>
)
</materialsCitation>
;
<materialsCitation id="995FA438AC317FB75AE1000DDF289E58" ID-GBIF-Occurrence="923899395" collectorName="WF Buren" country="U. S. A." county="Rapides Parish" location="Alexandria" stateProvince="Louisiana">
<collectingCounty id="7F5131592BDA5F23E0DA348BDF2245CC">Rapides Parish</collectingCounty>
:
<location id="E1DD1D4EE9DAB20008587962FBD79F58" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:E1DD1D4EE9DAB20008587962FBD79F58" country="U.s.a." name="Alexandria" stateProvince="Louisiana">Alexandria</location>
(
<collectorName id="32D75A8E667CF4CA0215561CC111F631">WF Buren</collectorName>
)
</materialsCitation>
;
<materialsCitation id="A28472719D4F78947967ADAE36D11048" ID-GBIF-Occurrence="923899389" collectorName="UG Mueller" country="U. S. A." county="Vernon Parish" location="Pickering" locationDeviation="7mi E" stateProvince="Louisiana">
<collectingCounty id="593C827BEC648E18C99D92557E63C709">Vernon Parish</collectingCounty>
:
<locationDeviation id="9ECF670F818BD06A95AF0F5B3206903F">7mi E</locationDeviation>
<location id="E8B33484D696C048DB2829BC3010481A" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:E8B33484D696C048DB2829BC3010481A" country="U.s.a." name="Pickering" stateProvince="Louisiana">Pickering</location>
(
<collectorName id="07EF5F9C05A3211CD95498B5BA96CD5F">UG Mueller</collectorName>
)
</materialsCitation>
;
<materialsCitation id="EFCF3662FE67C89565BAF72D149A40B3" ID-GBIF-Occurrence="923899390" collectorName="UG Mueller" country="U. S. A." county="Webster Parish" location="Sibley" locationDeviation="3mi W" stateProvince="Louisiana">
<collectingCounty id="CE5D8A6EE57CD1EB86CF6F0F05BFF815">Webster Parish</collectingCounty>
:
<locationDeviation id="6F95AD8626B565C26DF61F7BED7B7771">3mi W</locationDeviation>
<location id="246D21A8037687BA29C0BEE64ED4FF60" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:246D21A8037687BA29C0BEE64ED4FF60" country="U.s.a." name="Sibley" stateProvince="Louisiana">Sibley</location>
(
<collectorName id="A68E9D79A6119A98A96E3A14FD9C8303">UG Mueller</collectorName>
)
</materialsCitation>
;
<materialsCitation id="B1E1461503238C19EABE5100562F160D" ID-GBIF-Occurrence="923899392" collectorName="UG Mueller" country="U. S. A." county="Love County" location="I 35 and Red River" locationDeviation="200m N" stateProvince="Oklahoma">
<collectingRegion id="4CD73F4111FAD96A6AF572A028C123E7">Oklahoma</collectingRegion>
,
<collectingCounty id="F878F30A24FADD3F342F5891411F00FE">Love County</collectingCounty>
:
<location id="4136B1BE6B0918961FF423A6B401A732" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:4136B1BE6B0918961FF423A6B401A732" country="U.s.a." county="Love County" name="I 35 &amp; Red River" stateProvince="Oklahoma">I35 &amp; Red River</location>
,
<locationDeviation id="69818B4EE30367122812FA5B5ADCAE69">200m N</locationDeviation>
(
<collectorName id="272883134A255007DEB86A2C2735D24B">UG Mueller</collectorName>
)
</materialsCitation>
;
<materialsCitation id="AC707EF6CE8FE52366D70CD0367B961B" ID-GBIF-Occurrence="923899387" collectorName="UG Mueller" country="U. S. A." county="Austin County" location="Rt. 3013 crossing San Bernard River" stateProvince="Texas">
<collectingRegion id="75FA93008EB5DFBA85C73259CF68D7B5">Texas</collectingRegion>
,
<collectingCounty id="966A8A492EE8EDB7A72F9136B1DE62B9">Austin County</collectingCounty>
:
<location id="78D1CE4C0DD25F39EC323D1A624ED4CA" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:78D1CE4C0DD25F39EC323D1A624ED4CA" country="U.s.a." county="Austin County" name="Rt. 3013 crossing San Bernard River" stateProvince="Texas">Rt. 3013 crossing San Bernard River</location>
(
<collectorName id="52DD0DA3E102A5C39EDF65C759A46404">UG Mueller</collectorName>
)
</materialsCitation>
;
<materialsCitation id="BFC74321C2775624A2E59EBF42B15882" ID-GBIF-Occurrence="923899381" collectorName="UG Mueller, C Rabeling" country="U. S. A." county="Bastrop County" location="Stengl Biological Station" stateProvince="Texas">
<collectingCounty id="3D0FFE56F82E9DF2BC8EB0D4AE204A1A">Bastrop County</collectingCounty>
:
<location id="81F30E71FEB75DB2C6F936A1E5E77331" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:81F30E71FEB75DB2C6F936A1E5E77331" country="U.s.a." county="Bastrop County" name="Stengl Biological Station" stateProvince="Texas">Stengl Biological Station</location>
(
<collectorName id="EAB0F08C9FCB58A4329C84E6C947A9B7">UG Mueller, C Rabeling</collectorName>
)
</materialsCitation>
;
<materialsCitation id="7082A2561DEB2B2E65099FC3DA42CF4B" ID-GBIF-Occurrence="923899375" collectorName="UG Mueller" country="U. S. A." county="Baylor County" location="Round Timber" stateProvince="Texas">
<collectingCounty id="E76EBA28BDFBF8F1D2F33DE0E5A3C979">Baylor County</collectingCounty>
:
<location id="7AC7E57637E8037EED801A71EE815327" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:7AC7E57637E8037EED801A71EE815327" country="U.s.a." county="Baylor County" name="Round Timber" stateProvince="Texas">Round Timber</location>
(
<collectorName id="A1832EABD7509AAE3BD6172346473820">UG Mueller</collectorName>
)
</materialsCitation>
;
<materialsCitation id="034AAB65552C17A439F0EEE9FA45CBC1" ID-GBIF-Occurrence="923899394" collectorName="UG Mueller" country="U. S. A." county="Blanco County" location="Rt. 3232, 3 mi N Rt. 290" stateProvince="Texas">
<collectingCounty id="074C7414B382D36AD5B77AAFFFBEABD1">Blanco County</collectingCounty>
:
<location id="05DE0603A4154A3DA35EC8F4C77F2077" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:05DE0603A4154A3DA35EC8F4C77F2077" country="U.s.a." county="Blanco County" name="Rt. 3232, 3 mi N Rt. 290" stateProvince="Texas">Rt. 3232, 3mi N Rt. 290</location>
(
<collectorName id="55D670C04320DC732204894321FCAA58">UG Mueller</collectorName>
)
</materialsCitation>
,
<materialsCitation id="AEB2EFE62F10DE0AA9988C919CE88FD3" ID-GBIF-Occurrence="923899374" collectorName="UG Mueller, C Rabeling" country="U. S. A." county="Blanco County" location="Pedernales State Park" stateProvince="Texas">
<location id="9876D2514642E5D8DAFF5B82C3577A6A" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:9876D2514642E5D8DAFF5B82C3577A6A" country="U.s.a." name="Pedernales State Park" stateProvince="Texas">Pedernales State Park</location>
(
<collectorName id="E261CFB293CA8D68AB86238FE96A7546">UG Mueller, C Rabeling</collectorName>
)
</materialsCitation>
;
<materialsCitation id="59542D8A343A126A648C7C136C71DDE7" ID-GBIF-Occurrence="923899397" collectorName="OF Francke, JV Moody, TB Hall" country="U. S. A." county="Brewster County" location="Marathon" locationDeviation="18mi NE" stateProvince="Texas">
<collectingCounty id="370854843B688ADA307CC435909D17EF">Brewster County</collectingCounty>
:
<locationDeviation id="0303B738E4479CBE0E041AC3652E151B">18mi NE</locationDeviation>
<location id="DE532B4F93B1EA8A428E30E1582E0259" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:DE532B4F93B1EA8A428E30E1582E0259" country="U.s.a." county="Brewster County" name="Marathon" stateProvince="Texas">Marathon</location>
(
<collectorName id="730E160D7746A52FA50678A962A5AB03">OF Francke, JV Moody &amp; TB Hall</collectorName>
)
</materialsCitation>
,
<materialsCitation id="00930D3EC944F4C659DC70A9F65A6340" ID-GBIF-Occurrence="923899378" collectorName="OF Francke, JV Moody, TB Hall" country="U. S. A." county="Brewster County" location="Alpine" locationDeviation="6mi E" stateProvince="Texas">
<locationDeviation id="3D90CD86026728E2D94C0D4616138A8A">6mi E</locationDeviation>
<location id="08CEA71882CDE18858EC8102415BF753" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:08CEA71882CDE18858EC8102415BF753" country="U.s.a." name="Alpine" stateProvince="Texas">Alpine</location>
(
<collectorName id="21BFC1CD7ADBA1C6F17AE4A81CCCDCD1">OF Francke, JV Moody &amp; TB Hall</collectorName>
)
</materialsCitation>
,
<materialsCitation id="61D1D9A21D2F7CF419F5805CFFDCFBF2" ID-GBIF-Occurrence="923899388" collectorName="OF Francke, JV Moody, TB Hall" country="U. S. A." county="Brewster County" location="Panther Junction" locationDeviation="6mi SE" stateProvince="Texas">
<locationDeviation id="2F8354996288CF9450F0B68BD12D306B">6mi SE</locationDeviation>
<location id="2443D2FEE8BF34DB0D35D1C7DA68AAF5" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:2443D2FEE8BF34DB0D35D1C7DA68AAF5" country="U.s.a." name="Panther Junction" stateProvince="Texas">Panther Junction</location>
(
<collectorName id="1E089B11B5BB29D74E9F367C7C8F69A4">OF Francke, JV Moody &amp; TB Hall</collectorName>
)
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(
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(
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(
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(
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(
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(
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(
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<collectingCounty id="6826AD0DB18DB5DC316719434D698C5E">Refugio County</collectingCounty>
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(
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:
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(
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:
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:
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(
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<collectingCounty id="DF583D70EF8465524824F8CA118AB2CE">Smith County</collectingCounty>
:
<location id="A4753F2A67A6A25AAEE8916653606774" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:A4753F2A67A6A25AAEE8916653606774" country="U.s.a." county="Smith County" name="Tyler State Park" stateProvince="Texas">Tyler State Park</location>
(
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<collectingCounty id="27FF4FCDFA375FDD47EDE1C08385CAD9">Starr County</collectingCounty>
:
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:
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<location id="CA4BE8C1EE9AD5FB311691CE85F14ED1" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:CA4BE8C1EE9AD5FB311691CE85F14ED1" country="U.s.a." county="Tom Green County" name="San Angelo" stateProvince="Texas">San Angelo</location>
(
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(
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<location id="57179601A01D569C1C5AD5F123149ED6" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:57179601A01D569C1C5AD5F123149ED6" country="U.s.a." name="Bull Creek Park" stateProvince="Texas">Bull Creek Park</location>
(
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,
<materialsCitation id="653FB2DD45F45E2A24C0DE816F438F73" ID-GBIF-Occurrence="923899465" collectorName="UG Mueller, C Rabeling" country="U. S. A." county="Travis County" location="Hamilton Pool Reserve" stateProvince="Texas">
<location id="7DD9D06D3C9FA0F3A8FF3BFE01AD0F2A" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:7DD9D06D3C9FA0F3A8FF3BFE01AD0F2A" country="U.s.a." name="Hamilton Pool Reserve" stateProvince="Texas">Hamilton Pool Reserve</location>
(
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<collectingCounty id="B8DBC45A34B41EC21929CD5268D7388B">Upshur County</collectingCounty>
:
<location id="AAE8444464869CF102D1BE019A474990" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:AAE8444464869CF102D1BE019A474990" country="U.s.a." county="Upshur County" name="Rhonesboro" stateProvince="Texas">Rhonesboro</location>
(
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<materialsCitation id="1CCD92B28EE27272E1C9A44293CD9A2D" ID-GBIF-Occurrence="923899457" collectorName="ACF Hung, UG Mueller" country="U. S. A." county="Uvalde County" location="Uvalde" stateProvince="Texas">
<collectingCounty id="C789B25B35ECCDF5BBD63ABBA40F7BCF">Uvalde County</collectingCounty>
:
<location id="B80C2DA5E1C345F2D658BCF2397197C9" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:B80C2DA5E1C345F2D658BCF2397197C9" country="U.s.a." county="Uvalde County" name="Uvalde" stateProvince="Texas">Uvalde</location>
(
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,
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<collectingCounty id="3445819CA91C5C7B9BCD6F81D4A42EE7">Val Verde County</collectingCounty>
:
<location id="733F34F2B4C4FA2D978EDD825AE8F75F" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:733F34F2B4C4FA2D978EDD825AE8F75F" country="U.s.a." county="Val Verde County" name="Del Rio" stateProvince="Texas">Del Rio</location>
(
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<collectingCounty id="D2E3B20B6BDE8F2523B50BA6BBB6901B">Walker County</collectingCounty>
:
<location id="C1E6AED01BA61E03BE9B5EA9A95F328B" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:C1E6AED01BA61E03BE9B5EA9A95F328B" country="U.s.a." county="Walker County" name="Huntsville" stateProvince="Texas">Huntsville</location>
(
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;
<materialsCitation id="6F9893D65DEB2DE39DFB7FA01A3A1CE9" ID-GBIF-Occurrence="923899461" collectorName="UG Mueller" country="U. S. A." county="Washington County" location="Washington-on-the-Brazos State Park" stateProvince="Texas">
<collectingCounty id="DA4E5B07F0289F17D4176AA5C607AE8F">Washington County</collectingCounty>
:
<location id="E30D4865B8714C37121A0DECACBBD871" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:E30D4865B8714C37121A0DECACBBD871" country="U.s.a." county="Washington County" name="Washington-onthe-Brazos State Park" stateProvince="Texas">Washington-on-the-Brazos State Park</location>
(
<collectorName id="F56020567BF961169D60FAACD7B57631">UG Mueller</collectorName>
)
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;
<materialsCitation id="6986DEB83DDC0542B4BCE616CD853108" ID-GBIF-Occurrence="923899456" collectorName="OF Francke, JV Moody, TB Hall" country="U. S. A." county="Webb County" location="Laredo" locationDeviation="52.5mi N" stateProvince="Texas">
<collectingCounty id="56CA87A4928E1A0DBD9947B3EA3E1285">Webb County</collectingCounty>
:
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<location id="EEA9380C36CA8035E3E9298972FB259D" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:EEA9380C36CA8035E3E9298972FB259D" country="U.s.a." county="Webb County" name="Laredo" stateProvince="Texas">Laredo</location>
(
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;
<materialsCitation id="DC36D2E7307F029A3B43A200C727A5A1" ID-GBIF-Occurrence="923899451" collectorName="UG Mueller" country="U. S. A." county="Willacy County" location="Kenedy County border" locationDeviation="0.7mi S" stateProvince="Texas">
<collectingCounty id="A6D4135DB4FA24505E836F05193E0B32">Willacy County</collectingCounty>
:
<locationDeviation id="7EDBA838DCA42199CBB7FB8F00590396">0.7mi S</locationDeviation>
<location id="A136DF96955CD094C9CD25C831DC8897" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:A136DF96955CD094C9CD25C831DC8897" country="U.s.a." county="Willacy County" name="Kenedy County border" stateProvince="Texas">
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border
</location>
(
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;
<materialsCitation id="1F7054C411F9FF8C489AEDDC1F89AB43" ID-GBIF-Occurrence="923899468" collectorName="JV Moody" country="U. S. A." county="Wise County" location="Bridgeport" locationDeviation="7.5mi SW" stateProvince="Texas">
<collectingCounty id="229300FE7D140F87B981A158F87F28FA">Wise County</collectingCounty>
:
<locationDeviation id="E4C36AAC9E8BF06FF0953878E46E1822">7.5mi SW</locationDeviation>
<location id="4D9A3ABEF694822097DD9137C53504F2" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:4D9A3ABEF694822097DD9137C53504F2" country="U.s.a." county="Wise County" name="Bridgeport" stateProvince="Texas">Bridgeport</location>
(
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;
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<collectingCounty id="F9D7BB337612E54B5E6E4D263B585BE2">Wood County</collectingCounty>
:
<location id="1C774B7FBE281737F36CF820AB2571D6" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:1C774B7FBE281737F36CF820AB2571D6" country="U.s.a." county="Wood County" name="9 mi E Minneola on US 80" stateProvince="Texas">9mi E Minneola on US 80</location>
(
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;
<materialsCitation id="5653012C1FD8CB902995BBD9D73FAA5F" ID-GBIF-Occurrence="923899469" collectorName="UG Mueller" country="U. S. A." county="Young County" location="Fort Belknap" stateProvince="Texas">
<collectingCounty id="603F2AE9447A276501521898748B27FB">Young County</collectingCounty>
:
<location id="90EAFB909835DBB0503CFD9138A699E7" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:90EAFB909835DBB0503CFD9138A699E7" country="U.s.a." county="Young County" name="Fort Belknap" stateProvince="Texas">Fort Belknap</location>
(
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;
<materialsCitation id="59C13DEC4E17318B557CA3DCCFAA4D69" ID-GBIF-Occurrence="923899452" collectorName="UG Mueller" country="U. S. A." county="Zapata County" location="Falcon State Park" stateProvince="Texas">
<collectingCounty id="838F0C4FF942A53D51EC0DA0FD48A8A5">Zapata County</collectingCounty>
:
<location id="9C983AA811372E1BB8F70BD7110C95D0" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:9C983AA811372E1BB8F70BD7110C95D0" country="U.s.a." county="Zapata County" name="Falcon State Park" stateProvince="Texas">Falcon State Park</location>
(
<collectorName id="C103C00B8F312F431E91F0A00015B101">UG Mueller</collectorName>
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;
<materialsCitation id="86FADE3AA4DE40628FAACE4814F3BD36" ID-GBIF-Occurrence="923899462" collectorName="UG Mueller" country="U. S. A." county="Zava County" location="Pryor" stateProvince="Texas">
<collectingCounty id="C4772B57CA40C3ABB0C47C8CE0721CF7">Zava County</collectingCounty>
:
<location id="9CD0B3004B98256FD8BBBDC626DE0529" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:9CD0B3004B98256FD8BBBDC626DE0529" country="U.s.a." county="Zava County" name="Pryor" stateProvince="Texas">Pryor</location>
(
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)
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;
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:
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,
<location id="AAABC21C60818B3949EBD7BFAF99A824" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:AAABC21C60818B3949EBD7BFAF99A824" country="MEXICO" name="Vallecillo" stateProvince="Nuevo Leon">Vallecillo</location>
(
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,
<location id="1922F2249AE1BBA600C7DD82C3BA32E2" LSID="urn:lsid:plazi:treatment:1DC94E957608D014A3764553992F8457:1922F2249AE1BBA600C7DD82C3BA32E2" country="Mexico" name="near Cuidad Victoria" stateProvince="Tamaulipas">near Cuidad Victoria</location>
(
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)
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.
</paragraph>
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