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<document id="FAF1E05C36DEA4986037EA14BA0DC292" ID-DOI="10.37520/aemnp.2023.014" ID-ISSN="1804-6487" ID-Zenodo-Dep="10134093" ID-ZooBank="D982BDAC-CB44-41BE-A35C-92D630F96BF7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699897763845" checkinUser="felipe" docAuthor="Bilton, David T." docDate="2023" docId="03F6746EFFDFB62BFC69FCCAFD80B431" docLanguage="en" docName="ActaEntMusNatPra.63.2.249-264.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2023.014" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Helophorus (Ropalohelophorus) cooperi Orchymont 1947, stat. nov." docType="treatment" docVersion="2" lastPageNumber="258" masterDocId="FFCF0C16FFD9B622FFE4FFB6FFE2B204" masterDocTitle="Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)" masterLastPageNumber="264" masterPageNumber="249" pageNumber="255" updateTime="1700044289635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="B8A8AA53C0F50AD847C7217DBE2F94A4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BBE7B21A7DE9A35C0A6BF79761DC8F40">
<mods:title id="FD88B054AFD7D643119591AFC46D2975">Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)</mods:title>
</mods:titleInfo>
<mods:name id="0191AE10E714FF28F06D29A58B6CCABF" type="personal">
<mods:role id="94B3B382E4E3DCF0F119CC2202AC70E9">
<mods:roleTerm id="9C299815E22F177F2E0B74507A31890E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BC6E97228E89B92C522FA3DE792547E">Bilton, David T.</mods:namePart>
<mods:affiliation id="0C1B384E6ED3A816CCCFF68014DBC192">Marine Biology and Ecology Research Centre, School of Biological and Marine Sciences, University of Plymouth, Drake Circus Plymouth PL &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, 2006 Johannesburg, South Africa</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D90D5B1EA8D5ABB2D5A0D2B532FED7C1">text</mods:typeOfResource>
<mods:relatedItem id="E75B600E1E217C5FECC5BBEE7A70B576" type="host">
<mods:titleInfo id="EDC4F8CC5ABA065146841136A07EA572">
<mods:title id="84E6240B41444A7BF1AB35B54C6500CF">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="AF09B9CFE40C68793778631E635169DB">
<mods:date id="BF5E48E4FDBA7AAA957F3675F6BE6B69">2023</mods:date>
<mods:detail id="866F6514DA2D3EC19188364ADCE6AF56" type="series">
<mods:title id="DFCE1EFA405AE02401777F0A7F32CE4B">Acta. Ent. Mus. Natl. Pragae</mods:title>
</mods:detail>
<mods:detail id="7B32DF88763BBCC7C3A716F636E13211" type="pubDate">
<mods:number id="01964A9FC826A8734E5928DCA452A2F1">2023-09-10</mods:number>
</mods:detail>
<mods:detail id="85111D50151C059CEF1989B5C9A7DFFC" type="volume">
<mods:number id="8071359CC1DD06360D387AB755B41135">63</mods:number>
</mods:detail>
<mods:detail id="89CE12D91CD88A2944C38E7A7869F9F2" type="issue">
<mods:number id="41AB2705AD505C5FB993F547CE2690F5">2</mods:number>
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<mods:extent id="F64F42B5202F0280945E39B56BB70B12" unit="page">
<mods:start id="E9703E64857EEEDF35E0A41BD8DC7461">249</mods:start>
<mods:end id="2462C5525973A7274E8F1088D397AB4D">264</mods:end>
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<mods:location id="4D4ACF3864755DCCB36E00430AADF007">
<mods:url id="FE2BE781D5AD8F557AE115089A194229">http://dx.doi.org/10.37520/aemnp.2023.014</mods:url>
</mods:location>
<mods:classification id="B4B6D9C91BED6FF8A6B5F17DF84A7590">journal article</mods:classification>
<mods:identifier id="9030BCD5E900592662A3AF9A2F7C3EB8" type="DOI">10.37520/aemnp.2023.014</mods:identifier>
<mods:identifier id="B8E99E4C7BB8F1FCA90ABAFCB282762C" type="ISSN">1804-6487</mods:identifier>
<mods:identifier id="7B0A42D3C54F80BCF1AEE261D8CA79C1" type="Zenodo-Dep">10134093</mods:identifier>
<mods:identifier id="D4B487633EE4AEB768AFE0BF1180E1BC" type="ZooBank">D982BDAC-CB44-41BE-A35C-92D630F96BF7</mods:identifier>
</mods:mods>
<treatment id="03F6746EFFDFB62BFC69FCCAFD80B431" LSID="urn:lsid:plazi:treatment:03F6746EFFDFB62BFC69FCCAFD80B431" httpUri="http://treatment.plazi.org/id/03F6746EFFDFB62BFC69FCCAFD80B431" lastPageId="9" lastPageNumber="258" pageId="6" pageNumber="255">
<subSubSection id="C34596F3FFDFB624FC69FCCAFAEBB1B0" pageId="6" pageNumber="255" type="nomenclature">
<paragraph id="8BE0C578FFDFB624FC69FCCAFAEBB1B0" blockId="6.[909,1354,892,948]" pageId="6" pageNumber="255">
<heading id="D0A87214FFDFB624FC69FCCAFAEBB1B0" centered="true" fontSize="10" level="2" pageId="6" pageNumber="255" reason="3">
<emphasis id="B92B196AFFDFB624FC69FCCAFAEBB1B0" bold="true" pageId="6" pageNumber="255">
<taxonomicName id="4C5FBEFBFFDFB624FC69FCCAFB75B1B0" ID-CoL="3KHJ3" authority="Orchymont, 1947" authorityName="Orchymont" authorityYear="1947" baseAuthorityName="Orchymont" baseAuthorityYear="1948" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="cooperi" status="stat. nov." subGenus="Ropalohelophorus">
<emphasis id="B92B196AFFDFB624FC69FCCAFBEDB190" bold="true" box="[909,1039,892,916]" italics="true" pageId="6" pageNumber="255">Helophorus</emphasis>
(
<emphasis id="B92B196AFFDFB624FBFBFCCAFB08B190" bold="true" box="[1055,1258,892,916]" italics="true" pageId="6" pageNumber="255">Ropalohelophorus</emphasis>
)
<emphasis id="B92B196AFFDFB624FB1DFCCBFAA8B191" bold="true" box="[1273,1354,893,917]" italics="true" pageId="6" pageNumber="255">cooperi</emphasis>
Orchymont, 1947
</taxonomicName>
,
<taxonomicNameLabel id="A218A411FFDFB624FB47FC2BFAEBB1B0" box="[1187,1289,925,948]" pageId="6" pageNumber="255">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34596F3FFDFB624FBF0FC0AFB26B1CB" box="[1044,1220,956,975]" pageId="6" pageNumber="255" type="description">
<paragraph id="8BE0C578FFDFB624FBF0FC0AFB26B1CB" blockId="6.[828,1302,956,1009]" box="[1044,1220,956,975]" pageId="6" pageNumber="255">
(
<figureCitation id="1364D9FDFFDFB624FBFEFC0AFB82B1CB" box="[1050,1120,956,975]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="6" pageNumber="255">Figs 1D</figureCitation>
,
<figureCitation id="1364D9FDFFDFB624FB8EFC0AFB64B1CB" box="[1130,1158,956,975]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="6" pageNumber="255">2G</figureCitation>
,
<figureCitation id="1364D9FDFFDFB624FB6BFC0AFB4BB1CB" box="[1167,1193,956,975]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="6" pageNumber="255">3C</figureCitation>
,
<figureCitation id="1364D9FDFFDFB624FB56FC0AFB5FB1CB" box="[1202,1213,956,975]" captionStart="Fig" captionStartId="14.[1132,1165,1128,1147]" captionTargetBox="[152,1098,191,1200]" captionTargetId="figure-437@14.[152,1098,191,1200]" captionTargetPageId="14" captionText="Fig. 5. Afrotropical Helophorus, known distributions.Map by https:// mapswire.com." figureDoi="http://doi.org/10.5281/zenodo.10134103" httpUri="https://zenodo.org/record/10134103/files/figure.png" pageId="6" pageNumber="255">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34596F3FFDFB624FCD8FC68FAF4B1F5" box="[828,1302,990,1009]" pageId="6" pageNumber="255" type="reference_group">
<paragraph id="8BE0C578FFDFB624FCD8FC68FAF4B1F5" blockId="6.[828,1302,956,1009]" box="[828,1302,990,1009]" pageId="6" pageNumber="255">
<taxonomicName id="4C5FBEFBFFDFB624FCD8FC68FAF4B1F5" ID-CoL="7H76S" authority="Orchymont, 1947: 722" authorityName="Orchymont" authorityPageNumber="722" authorityYear="1947" box="[828,1302,990,1009]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="aethiopicus" subSpecies="cooperi">
<emphasis id="B92B196AFFDFB624FCD8FC68FBACB1F5" box="[828,1102,990,1009]" italics="true" pageId="6" pageNumber="255">Helophorus aethiopicus cooperi</emphasis>
Orchymont, 1947: 722
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C34596F3FFDFB624FCD8FBBEFA72B744" pageId="6" pageNumber="255" type="materials_examined">
<paragraph id="8BE0C578FFDFB624FCD8FBBEFBC8B681" blockId="6.[828,1438,1032,1344]" pageId="6" pageNumber="255">
<emphasis id="B92B196AFFDFB624FCD8FBBEFC59B61F" bold="true" box="[828,955,1032,1051]" pageId="6" pageNumber="255">
<typeStatus id="54E47BDAFFDFB624FCD8FBBEFC88B61F" box="[828,874,1032,1051]" pageId="6" pageNumber="255">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F34885E8FFDFB624FC20FBBEFBF3B61F" box="[964,1041,1032,1051]" name="Ethiopia" pageId="6" pageNumber="255">Ethiopia</collectingCountry>
,
<collectingRegion id="499B0B9AFFDFB624FBF9FBBEFB57B61F" box="[1053,1205,1032,1051]" country="Ethiopia" name="Oromiya" pageId="6" pageNumber="255">Oromiya Region</collectingRegion>
, Wourambouchi, close to Djem-Djem Forest, ca.
<quantity id="4CA7689DFFDFB624FBF1FB94FBADB631" box="[1045,1103,1058,1077]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="255" unit="km" value="60.0">60 km</quantity>
W of Addis-Abbaba, approximately
<geoCoordinate id="EE6BA3BFFFDFB624FCD8FB8BFC9AB654" box="[828,888,1085,1104]" degrees="9" direction="north" minutes="03" orientation="latitude" pageId="6" pageNumber="255" precision="925" value="9.05">9°03′N</geoCoordinate>
<geoCoordinate id="EE6BA3BFFFDFB624FC9BFB8BFC27B654" box="[895,965,1085,1104]" degrees="38" direction="east" minutes="09" orientation="longitude" pageId="6" pageNumber="255" precision="925" value="38.15">38°09′E</geoCoordinate>
, ca.
<quantity id="4CA7689DFFDFB624FC0AFB8BFBD5B654" box="[1006,1079,1085,1104]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="6" pageNumber="255" unit="m" value="2750.0">2,750 m</quantity>
. Exact details imprecise. Wouramboulchi is a frequent locality on historical specimens from the region and was close to Djem-Djem Forest.
</paragraph>
<paragraph id="8BE0C578FFDFB624FCD8FB3BFA72B744" blockId="6.[828,1438,1032,1344]" pageId="6" pageNumber="255">
<emphasis id="B92B196AFFDFB624FCD8FB3BFC59B6A4" bold="true" box="[828,955,1165,1184]" pageId="6" pageNumber="255">Type material.</emphasis>
<materialsCitation id="3B37CF25FFDFB624FC5AFB3BFA72B744" collectingDate="1926-10-02" collectingDateMax="1926-10-07" collectingDateMin="1926-10-02" collectionCode="BMNH, RBINS" collectorName="J. Omer-Cooper." country="Ethiopia" determinerName="A. d'Orchymont" elevation="2743" location="Wouramboulchi" pageId="6" pageNumber="255" specimenCount="4" specimenCount-female="4" typeStatus="holotype">
<typeStatus id="54E47BDAFFDFB624FC5AFB3BFBF5B6A4" box="[958,1047,1165,1184]" pageId="6" pageNumber="255" type="holotype">HOLOTYPE</typeStatus>
: J, labelled:“J” “
<collectingCountry id="F34885E8FFDFB624FB54FB3BFAE7B6A4" box="[1200,1285,1165,1184]" name="Ethiopia" pageId="6" pageNumber="255">Abyssinia</collectingCountry>
//
<location id="8E8093A3FFDFB624FAF3FB3BFA7FB6A4" LSID="urn:lsid:plazi:treatment:03F6746EFFDFB62BFC69FCCAFD80B431:8E8093A3FFDFB624FAF3FB3BFA7FB6A4" box="[1303,1437,1165,1184]" country="Ethiopia" name="Wouramboulchi" pageId="6" pageNumber="255">Wouramboulchi</location>
//
<quantity id="4CA7689DFFDFB624FCAAFB1EFC70B6BF" box="[846,914,1192,1211]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="6" pageNumber="255" unit="ft" value="9000.0">
<elevation id="0072224BFFDFB624FCAAFB1EFC70B6BF" box="[846,914,1192,1211]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="6" pageNumber="255" unit="ft" value="9000.0">9,000ft.</elevation>
</quantity>
//
<date id="FFE1E3B8FFDFB624FC4EFB1EFBEBB6BF" box="[938,1033,1192,1211]" pageId="6" pageNumber="255" value="1926-10-02" valueMax="1926-10-07" valueMin="1926-10-02">
<collectingDate id="EFA51A50FFDFB624FC4EFB1EFBEBB6BF" box="[938,1033,1192,1211]" pageId="6" pageNumber="255" value="1926-10-02" valueMax="1926-10-07" valueMin="1926-10-02">2-7.x.1926</collectingDate>
</date>
. //
<collectorName id="26AAA0AEFFDFB624FBC1FB1EFB55B6BF" box="[1061,1207,1192,1211]" pageId="6" pageNumber="255">J. Omer-Cooper.</collectorName>
” “TYPE” [pink label] “
<determinerName id="7E3C29BCFFDFB624FA63FB1EFC4DB6D1" pageId="6" pageNumber="255">A. d’Orchymont</determinerName>
det. //
<taxonomicName id="4C5FBEFBFFDFB624FC02FB74FBA8B6D1" authorityName="Fabricius" authorityYear="1775" box="[998,1098,1218,1237]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
(s.str.) aethio // picus cooperi m.” [name Hw.] (
<collectionCode id="ED4E5DBDFFDFB624FC96FB6BFC55B6F4" box="[882,951,1245,1264]" pageId="6" pageNumber="255">BMNH</collectionCode>
).
<typeStatus id="54E47BDAFFDFB624FC22FB6BFBFDB6F4" box="[966,1055,1245,1264]" pageId="6" pageNumber="255" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D590EF1FFDFB624FBCFFB6BFBA9B6F4" box="[1067,1099,1239,1264]" count="1" pageId="6" pageNumber="255" type="female">1 ♀</specimenCount>
, “
<specimenCount id="9D590EF1FFDFB624FB84FB61FB92B6F4" box="[1120,1136,1239,1264]" count="1" pageId="6" pageNumber="255" type="female">♀</specimenCount>
” otherwise same data as Holotype (
<collectionCode id="ED4E5DBDFFDFB624FCA6FB4EFC64B70F" box="[834,902,1272,1291]" pageId="6" pageNumber="255">BMNH</collectionCode>
);
<specimenCount id="9D590EF1FFDFB624FC76FB4EFC4DB70E" box="[914,943,1265,1291]" count="1" pageId="6" pageNumber="255" type="female">1♀</specimenCount>
, “
<specimenCount id="9D590EF1FFDFB624FC26FB47FC30B70E" box="[962,978,1265,1290]" count="1" pageId="6" pageNumber="255" type="female">♀</specimenCount>
” “
<collectingCountry id="F34885E8FFDFB624FC03FB4EFBDFB70F" box="[999,1085,1272,1291]" name="Ethiopia" pageId="6" pageNumber="255">Abyssinia</collectingCountry>
// Wouramboulchi //
<quantity id="4CA7689DFFDFB624FB0CFB4EFAC9B70F" box="[1256,1323,1272,1291]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="6" pageNumber="255" unit="ft" value="9000.0">
<elevation id="0072224BFFDFB624FB0CFB4EFAC9B70F" box="[1256,1323,1272,1291]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="6" pageNumber="255" unit="ft" value="9000.0">9,000ft.</elevation>
</quantity>
//
<date id="FFE1E3B8FFDFB624FAD8FB4EFA7BB70F" box="[1340,1433,1272,1291]" pageId="6" pageNumber="255" value="1926-10-02" valueMax="1926-10-07" valueMin="1926-10-02">
<collectingDate id="EFA51A50FFDFB624FAD8FB4EFA7BB70F" box="[1340,1433,1272,1291]" pageId="6" pageNumber="255" value="1926-10-02" valueMax="1926-10-07" valueMin="1926-10-02">2-7.x.1926</collectingDate>
</date>
. //
<collectorName id="26AAA0AEFFDFB624FCAAFAA4FC02B721" box="[846,992,1298,1317]" pageId="6" pageNumber="255">J. Omer-Cooper.</collectorName>
” “
<determinerName id="7E3C29BCFFDFB624FC11FAA4FB66B721" box="[1013,1156,1298,1317]" pageId="6" pageNumber="255">A. d’Orchymont</determinerName>
det. //
<taxonomicName id="4C5FBEFBFFDFB624FB26FAA4FAC4B721" authorityName="Fabricius" authorityYear="1775" box="[1218,1318,1298,1317]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
(s.str.) aethio // picus cooperi m.” [name Hw.] “
<typeStatus id="54E47BDAFFDFB624FBBFFA9BFB68B744" box="[1115,1162,1325,1344]" pageId="6" pageNumber="255" type="paratype">Para-</typeStatus>
// type” [pink label) (
<collectionCode id="ED4E5DBDFFDFB624FAA6FA9BFA67B744" box="[1346,1413,1325,1344]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="6" pageNumber="255" type="Museum">RBINS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34596F3FFDFB62BFCD8FAEFFD72B6D1" lastPageId="9" lastPageNumber="258" pageId="6" pageNumber="255" type="description">
<paragraph id="8BE0C578FFDFB624FCD8FAEFFC31B594" blockId="6.[828,1437,1368,2064]" pageId="6" pageNumber="255">
<emphasis id="B92B196AFFDFB624FCD8FAEFFBCEB774" bold="true" box="[828,1068,1368,1392]" pageId="6" pageNumber="255">
Redescription.
<emphasis id="B92B196AFFDFB624FC16FAEEFBCEB774" bold="true" box="[1010,1068,1368,1392]" italics="true" pageId="6" pageNumber="255">Male</emphasis>
</emphasis>
(
<figureCitation id="1364D9FDFFDFB624FBDBFAEFFB79B774" box="[1087,1179,1369,1392]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="6" pageNumber="255">Fig. 1D</figureCitation>
).
<emphasis id="B92B196AFFDFB624FB50FAEFFB0DB774" box="[1204,1263,1369,1392]" italics="true" pageId="6" pageNumber="255">Head</emphasis>
dark brown to black, with green aeneous reflection; shining. Granules evident over entire frons and clypeus, but partly or fully coalescent with adjacent granules in places. Y-groove relatively shallow; floor shining, with dense, shallow, coarse, rugose punctures; stem weakly diverging anteriorly; lateral branches narrow, particularly over apical 0.5, straight. Clypeus strongly elevated in centre; anterior margin weakly arcuate. Labrum with shining, impunctate, basal portion with anterior margin produced and rounded at centre; anterior portion with coarse, rugose punctures, each puncture bearing a short, stout, whitish semi-erect seta; anterior margin arcuate with shallow anteromedian emargination. Antennae 9-segmented, yellowish brown, with cupule and club segments darker. Maxillary palpi yellow, apex of terminal segment infuscated; moderately elongate, terminal segment longest, curved and clearly asymmetrical.
</paragraph>
<paragraph id="8BE0C578FFDFB625FCBEF82FFBEFB7FB" blockId="6.[828,1437,1368,2064]" lastBlockId="7.[828,1437,1448,2079]" lastPageId="7" lastPageNumber="256" pageId="6" pageNumber="255">
<emphasis id="B92B196AFFDFB624FCBEF82FFC25B5B4" box="[858,967,1945,1968]" italics="true" pageId="6" pageNumber="255">Pronotum</emphasis>
(
<figureCitation id="1364D9FDFFDFB624FC3EF82FFBD0B5B4" box="[986,1074,1945,1968]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="6" pageNumber="255">Fig. 3C</figureCitation>
) pale reddish brown, with pale green aeneous reflections weak and restricted to sublateral groove; anterior and lateral borders paler, but contrast with remainder of pronotum indistinct; shining over entire dorsal surface. Broadest between middle and anterior margins; weakly arched in frontal and dorsal views and somewhat flattened over internal intervals, particularly anteriorly. Anterior margin slightly concave over central 0.5, then sinuate to broadly rounded front angles; posterior margin broadly bisinuate around centre. Sides straight over posterior 0.4, then weakly arcuate around broadest point to anterior angles. Lateral margins setose and crenulated to serrate. Internal and middle intervals slightly arched, strongly shining; granules reduced to pore and seta. External interval strongly shining with weak granules evident, particularly anteriorly. Grooves smooth and shining, with sparse punctures and wrinkles, particularly in sublateral and lateral. Median groove narrowed anteriorly, more so posteriorly, open at both ends more widely so anteriorly. Submedian and submarginal grooves broadly open anteriorly and posteriorly. Submedian grooves relatively narrow, weakly sinuate. Submarginal grooves almost straight, much broader than submedian. Marginal grooves curved parallel to the side of pronotum, width uniform.All grooves relatively deep, with intervals curving down towards their margins, which are well-defined but somewhat sloping rather than vertical.
</paragraph>
<caption id="DF2095F0FFDEB625FF73FA90FA7FB750" ID-DOI="http://doi.org/10.5281/zenodo.10134099" ID-Zenodo-Dep="10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="7" pageNumber="256" startId="7.[151,182,1318,1337]" targetBox="[244,1341,192,1289]" targetPageId="7" targetType="figure">
<paragraph id="8BE0C578FFDEB625FF73FA90FA7FB750" blockId="7.[150,1437,1318,1364]" pageId="7" pageNumber="256">
Fig. 3. Afrotropical
<taxonomicName id="4C5FBEFBFFDEB625FED9FA90FE40B73D" authorityName="Fabricius" authorityYear="1775" box="[317,418,1318,1337]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="B92B196AFFDEB625FED9FA90FE40B73D" box="[317,418,1318,1337]" italics="true" pageId="7" pageNumber="256">Helophorus</emphasis>
</taxonomicName>
, pronota.A –
<taxonomicName id="4C5FBEFBFFDEB625FDF2FA90FCC4B73D" authority="Regimbart, 1907" authorityName="Regimbart" authorityYear="1907" box="[534,806,1318,1337]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B92B196AFFDEB625FDF2FA90FD73B73D" box="[534,657,1318,1337]" italics="true" pageId="7" pageNumber="256">H. aethiopicus</emphasis>
<bibRefCitation id="EFCEB889FFDEB625FD71FA90FCC4B73D" author="REGIMBART M." box="[661,806,1318,1337]" pageId="7" pageNumber="256" pagination="126 - 128" refId="ref16606" refString="REGIMBART M. 1907: Collections recueillis par M. Maurice de Rothschild dans l'Afrique orientale. Insectes coleopteres: Dytiscides et hydrophilides. Bulletin du Museum National d'Histoire Naturelle Paris 13: 126 - 128." type="journal article" year="1907">Régimbart, 1907</bibRefCitation>
</taxonomicName>
, lectotype; B –
<taxonomicName id="4C5FBEFBFFDEB625FC43FA90FB02B73D" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" box="[935,1248,1318,1337]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="aethiops">
<emphasis id="B92B196AFFDEB625FC43FA90FBE5B73D" box="[935,1031,1318,1337]" italics="true" pageId="7" pageNumber="256">H.aethiops</emphasis>
J.
<bibRefCitation id="EFCEB889FFDEB625FBFAFA90FB02B73D" author="BALFOUR-BROWNE J." box="[1054,1248,1318,1337]" pageId="7" pageNumber="256" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
</taxonomicName>
, holotype; C –
<taxonomicName id="4C5FBEFBFFDEB625FAB9FA90FEB7B750" authority="Orchymont, 1947" authorityName="Orchymont" authorityYear="1947" baseAuthorityName="Orchymont" baseAuthorityYear="1948" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="B92B196AFFDEB625FAB9FA90FF5AB750" italics="true" pageId="7" pageNumber="256">H. cooperi</emphasis>
Orchymont, 1947
</taxonomicName>
, holotype; D –
<taxonomicName id="4C5FBEFBFFDEB625FE30FAF7FD8CB750" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[468,622,1345,1364]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
<emphasis id="B92B196AFFDEB625FE30FAF7FD8CB750" box="[468,622,1345,1364]" italics="true" pageId="7" pageNumber="256">H. brumopluvialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218A411FFDEB625FD97FAF7FD55B750" box="[627,695,1345,1364]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; E –
<taxonomicName id="4C5FBEFBFFDEB625FCD6FAF7FC31B750" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[818,979,1345,1364]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="nyandaruaensis" status="sp. nov.">
<emphasis id="B92B196AFFDEB625FCD6FAF7FC31B750" box="[818,979,1345,1364]" italics="true" pageId="7" pageNumber="256">H. nyandaruaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218A411FFDEB625FC33FAF7FBF9B750" box="[983,1051,1345,1364]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
, holotype; F –
<taxonomicName id="4C5FBEFBFFDEB625FB71FAF7FB1FB750" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1173,1277,1345,1364]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="simiensis" status="sp. nov.">
<emphasis id="B92B196AFFDEB625FB71FAF7FB1FB750" box="[1173,1277,1345,1364]" italics="true" pageId="7" pageNumber="256">H. simiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218A411FFDEB625FAE5FAF7FAA7B750" box="[1281,1349,1345,1364]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
, holotype.
</paragraph>
</caption>
<paragraph id="8BE0C578FFDEB625FCBEF9BEFB74B4BB" blockId="7.[828,1437,1448,2079]" pageId="7" pageNumber="256">
<emphasis id="B92B196AFFDEB625FCBEF9BEFC7FB41B" box="[858,925,1544,1567]" italics="true" pageId="7" pageNumber="256">Elytra</emphasis>
(
<figureCitation id="1364D9FDFFDEB625FC4AF9BEFBE1B41B" box="[942,1027,1544,1567]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="7" pageNumber="256">Fig. 1D</figureCitation>
) brown, with diffuse paler markings, particularly posteriorly. Shape elongate oval, sides rounded to bluntly rounded apex. In lateral profile flat over anterior 0.5, then gradually descending to apex. Puncture rows weakly striate, more so laterally; intervals slightly arched anteriorly and flatter posteriorly.
</paragraph>
<paragraph id="8BE0C578FFDEB625FCBEF97EFC68B51B" blockId="7.[828,1437,1448,2079]" pageId="7" pageNumber="256">
<emphasis id="B92B196AFFDEB625FCBEF97EFC6EB4DB" box="[858,908,1736,1759]" italics="true" pageId="7" pageNumber="256">Legs</emphasis>
yellowish-brown, apex of last tarsal segment and claws infuscated. Swimming hairs distinct and clearly visible.
</paragraph>
<paragraph id="8BE0C578FFDEB62BFCBEF89EFEB6B131" blockId="7.[828,1437,1448,2079]" lastBlockId="9.[151,761,190,1589]" lastPageId="9" lastPageNumber="258" pageId="7" pageNumber="256">
<emphasis id="B92B196AFFDEB625FCBEF89EFC7FB53B" box="[858,925,1832,1855]" italics="true" pageId="7" pageNumber="256">Venter</emphasis>
black, with pronotal hypomeron, antennal pocket and pseudepipleura and epipleura brown to yellow. Mentum shining, centre without microreticulation, with sparse, medium punctures; lateral marginal areas with weak isodiametric microreticulation centrally; margins with fine yellow setae, denser towards posterolateral angles. Submentum shining, without microreticulation; anterior half with sparse, shallow, coarse punctures and fine, hair-like vestiture. Gula and genae rugose, with dense hydrofuge vestiture. Gular sutures well marked as pits anteriorly. Vestiture only absent around margin of compound eyes and their posterolateral corners. Prosternum dull, rugose, with dense hydrofuge vestiture and sparse, long, erect setae approximately 2× length of vestiture. Pronotal hypomeron dull, with dense hydrofuge vestiture. Pronotal epipleura shining, glabrous, with sparse shallow punctures. Anterior antennal pocket well developed, with long, dense setae internally and long, curved setae at posterolateral margins. Elytral epipleura dull, glabrous, narrowing posteriorly at level of metaventrite and continuing until posterior margin of ventrite. Elytral flanks very narrowly visible from below. Ventral surface of mesothorax, metathorax and abdomen dull, with dense hydrofuge vestiture and longer, erect setae; longer setae concentrated on centre of metaventrite and abdominal ventrites 2–5; setae on ventrites arising from longitudinal tubercles, which sometimes appear as shining patches. Hind margin of abdominal ventrite 5 weakly crenulate.
</paragraph>
<caption id="DF2095F0FFD1B62AFF73F85DFA7CBA1C" ID-DOI="http://doi.org/10.5281/zenodo.10134101" ID-Zenodo-Dep="10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="8" pageNumber="257" startId="8.[151,182,2027,2046]" targetBox="[203,1385,189,1962]" targetPageId="8" targetType="figure">
<paragraph id="8BE0C578FFD1B62AFF73F85DFA7CBA1C" blockId="8.[151,1438,2027,2072]" pageId="8" pageNumber="257">
Fig. 4. South African
<taxonomicName id="4C5FBEFBFFD1B62AFEAEF85DFE4DB5FA" authorityName="Fabricius" authorityYear="1775" box="[330,431,2027,2046]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="B92B196AFFD1B62AFEAEF85DFE4DB5FA" box="[330,431,2027,2046]" italics="true" pageId="8" pageNumber="257">Helophorus</emphasis>
</taxonomicName>
, SEMs.A, C, E &amp; G –
<taxonomicName id="4C5FBEFBFFD1B62AFD88F85DFC46B5FA" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" box="[620,932,2027,2046]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="257" phylum="Arthropoda" rank="species" species="aethiops">
<emphasis id="B92B196AFFD1B62AFD88F85DFD2FB5FA" box="[620,717,2027,2046]" italics="true" pageId="8" pageNumber="257">H. aethiops</emphasis>
J.
<bibRefCitation id="EFCEB889FFD1B62AFD06F85DFC46B5FA" author="BALFOUR-BROWNE J." box="[738,932,2027,2046]" pageId="8" pageNumber="257" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
</taxonomicName>
, Kokstad, South Africa; B, D, F &amp; H –
<taxonomicName id="4C5FBEFBFFD1B62AFB02F85DFA62B5FA" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1254,1408,2027,2046]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="257" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
<emphasis id="B92B196AFFD1B62AFB02F85DFA62B5FA" box="[1254,1408,2027,2046]" italics="true" pageId="8" pageNumber="257">H. brumopluvialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218A411FFD1B62AFA60F85DFF5CBA1C" pageId="8" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
, Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex.
</paragraph>
</caption>
<paragraph id="8BE0C578FFD0B62BFF51FC88FDD4B611" blockId="9.[151,761,190,1589]" pageId="9" pageNumber="258">
<emphasis id="B92B196AFFD0B62BFF51FC88FEFFB151" box="[181,285,830,853]" italics="true" pageId="9" pageNumber="258">Aedeagus</emphasis>
(
<figureCitation id="1364D9FDFFD0B62BFECFFC88FE9FB151" box="[299,381,830,853]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="9" pageNumber="258">Fig. 2G</figureCitation>
) elongate,
<quantity id="4CA7689DFFD0B62BFE16FC88FDADB151" box="[498,591,830,853]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="258" unit="mm" value="0.65">0.65 mm</quantity>
in length. Parameres slightly shorter than basal piece; outer margins of parameres sinuate approximately 0.5 from apex; interior margins of parameres initially sloping, then rather straight, apices acuminated. Median lobe narrow, with weakly pointed apex; sides diverging posteriorly to base of struts, weakly concave; struts as long as tube.
</paragraph>
<paragraph id="8BE0C578FFD0B62BFF51FBABFE0CB631" blockId="9.[151,761,190,1589]" box="[181,494,1053,1077]" pageId="9" pageNumber="258">
<emphasis id="B92B196AFFD0B62BFF51FBABFEEDB631" bold="true" box="[181,271,1053,1077]" italics="true" pageId="9" pageNumber="258">Female.</emphasis>
Externally as males.
</paragraph>
<paragraph id="8BE0C578FFD0B62BFF51FB88FDDFB691" blockId="9.[151,761,190,1589]" pageId="9" pageNumber="258">
<emphasis id="B92B196AFFD0B62BFF51FB88FEBFB651" bold="true" box="[181,349,1086,1109]" italics="true" pageId="9" pageNumber="258">Measurements.</emphasis>
Males:
<typeStatus id="54E47BDAFFD0B62BFE54FB88FDF5B651" box="[432,535,1086,1109]" pageId="9" pageNumber="258" type="holotype">Holotype</typeStatus>
: BL =
<quantity id="4CA7689DFFD0B62BFDBEFB88FD59B651" box="[602,699,1086,1109]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" pageId="9" pageNumber="258" unit="mm" value="3.45">3.45 mm</quantity>
; EL = 2.30; EW =
<quantity id="4CA7689DFFD0B62BFEFFFBE8FE9CB671" box="[283,382,1118,1141]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="258" unit="mm" value="1.4">1.40 mm</quantity>
. Females: BL = 3.50–4.00 mm; EL =
<quantity id="4CA7689DFFD0B62BFF49FBC8FEA9B691" box="[173,331,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="2.575" metricValueMax="2.65" metricValueMin="2.5" pageId="9" pageNumber="258" unit="mm" value="2.575" valueMax="2.65" valueMin="2.5">2.50–2.65 mm</quantity>
; EW =
<quantity id="4CA7689DFFD0B62BFE7FFBC8FDDBB691" box="[411,569,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.65" metricValueMin="1.45" pageId="9" pageNumber="258" unit="mm" value="1.5499999999999998" valueMax="1.65" valueMin="1.45">1.45–1.65 mm</quantity>
.
</paragraph>
<paragraph id="8BE0C578FFD0B62BFF73FB28FD72B6D1" blockId="9.[151,761,190,1589]" pageId="9" pageNumber="258">
<emphasis id="B92B196AFFD0B62BFF73FB28FEE5B6B1" bold="true" box="[151,263,1182,1205]" pageId="9" pageNumber="258">Variation.</emphasis>
Limited, other than in size – note that only
<specimenCount id="9D590EF1FFD0B62BFD27FB28FEE4B6D1" count="3" pageId="9" pageNumber="258" type="generic">three specimens</specimenCount>
of this taxon are available, however.
</paragraph>
</subSubSection>
<subSubSection id="C34596F3FFD0B62BFF73FB68FEDEB751" pageId="9" pageNumber="258" type="diagnosis">
<paragraph id="8BE0C578FFD0B62BFF73FB68FEDEB751" blockId="9.[151,761,190,1589]" pageId="9" pageNumber="258">
<emphasis id="B92B196AFFD0B62BFF73FB68FE77B6F1" bold="true" box="[151,405,1246,1269]" pageId="9" pageNumber="258">Differential diagnosis.</emphasis>
<taxonomicName id="4C5FBEFBFFD0B62BFE7BFB68FD98B6F1" authorityName="Orchymont" authorityYear="1947" baseAuthorityName="Orchymont" baseAuthorityYear="1948" box="[415,634,1246,1269]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="B92B196AFFD0B62BFE7BFB68FD98B6F1" box="[415,634,1246,1269]" italics="true" pageId="9" pageNumber="258">Helophorus cooperi</emphasis>
</taxonomicName>
has a very distinctive pronotal sculpture, with smooth internal and middle intervals, 9-segmented antennae and a characteristic aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34596F3FFD0B62BFF73FAE8FD80B431" pageId="9" pageNumber="258" type="distribution">
<paragraph id="8BE0C578FFD0B62BFF73FAE8FD80B431" blockId="9.[151,761,190,1589]" pageId="9" pageNumber="258">
<emphasis id="B92B196AFFD0B62BFF73FAE8FE22B771" bold="true" box="[151,448,1374,1397]" pageId="9" pageNumber="258">Distribution and ecology.</emphasis>
Known only from the
<typeStatus id="54E47BDAFFD0B62BFD2DFAE8FD15B771" box="[713,759,1374,1397]" pageId="9" pageNumber="258">type</typeStatus>
locality, Wourambouchi, close to Djem-Djem Forest in the
<collectingRegion id="499B0B9AFFD0B62BFF27FA28FE9CB7B1" box="[195,382,1438,1461]" country="Ethiopia" name="Oromiya" pageId="9" pageNumber="258">Oromiya Region</collectingRegion>
, ca.
<quantity id="4CA7689DFFD0B62BFE52FA28FE1FB7B1" box="[438,509,1438,1461]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="258" unit="km" value="60.0">60 km</quantity>
W of Addis-Ababa at approximately
<quantity id="4CA7689DFFD0B62BFEDDFA08FE73B7D1" box="[313,401,1470,1493]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="9" pageNumber="258" unit="m" value="2750.0">2,750 m</quantity>
(
<figureCitation id="1364D9FDFFD0B62BFE44FA08FE00B7D1" box="[416,482,1470,1493]" captionStart="Fig" captionStartId="14.[1132,1165,1128,1147]" captionTargetBox="[152,1098,191,1200]" captionTargetId="figure-437@14.[152,1098,191,1200]" captionTargetPageId="14" captionText="Fig. 5. Afrotropical Helophorus, known distributions.Map by https:// mapswire.com." figureDoi="http://doi.org/10.5281/zenodo.10134103" httpUri="https://zenodo.org/record/10134103/files/figure.png" pageId="9" pageNumber="258">Fig. 5</figureCitation>
).
<taxonomicName id="4C5FBEFBFFD0B62BFE13FA08FD1AB7D1" authorityName="Regimbart" authorityYear="1907" box="[503,760,1470,1493]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="258" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B92B196AFFD0B62BFE13FA08FD1AB7D1" box="[503,760,1470,1493]" italics="true" pageId="9" pageNumber="258">Helophorus aethiopicus</emphasis>
</taxonomicName>
is recorded both north and south of this locality and the precise distribution of, and possible ecological differences between, the two species remain unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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