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<document id="6C77AD9D98C081AA7277B5143432EB82" ID-DOI="10.5281/zenodo.176099" ID-GBIF-Dataset="62c38912-84bc-496c-bccc-cd6b69a322dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="176099" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459827803207" checkinUser="plazi" docAuthor="Lelej, Arkady S., Ullah, Mishkat &amp; Mahmood, Khalid" docDate="2007" docId="03ECBF63A87F5B49FF13B42D04A0F817" docLanguage="en" docName="zt01444p060.pdf" docOrigin="Zootaxa 1444" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ephutomma himalayana Lelej and Ullah, sp. nov." docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="FFD5C71BA87A5B4CFF84B63A0661FFE3" masterDocTitle="Additions to the knowledge of the Mutillidae (Hymenoptera) of Pakistan" masterLastPageNumber="60" masterPageNumber="53" pageNumber="58" updateTime="1698222505284" updateUser="plazi">
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<mods:title id="085B25FA61C8CE2B5111CEB42B3A21A0">Additions to the knowledge of the Mutillidae (Hymenoptera) of Pakistan</mods:title>
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<mods:name id="AC8E5672DA57483DA5CA2AA1DE2A6B0A" type="personal">
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<mods:namePart id="47FDA51C53FC7A96AEFBA845D280BC11">Lelej, Arkady S.</mods:namePart>
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<mods:namePart id="E360418DE2DBF933A0FDF8F9436F9AFF">Ullah, Mishkat</mods:namePart>
</mods:name>
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<mods:namePart id="C765BE7D92F46EF319C90FE61CDC135C">Mahmood, Khalid</mods:namePart>
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<mods:date id="AE776C0E44F68A7D50BAE40895FC359D">2007</mods:date>
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<treatment id="03ECBF63A87F5B49FF13B42D04A0F817" ID-DOI="http://doi.org/10.5281/zenodo.6251659" ID-GBIF-Taxon="119357250" ID-Zenodo-Dep="6251659" LSID="urn:lsid:plazi:treatment:03ECBF63A87F5B49FF13B42D04A0F817" httpUri="http://treatment.plazi.org/id/03ECBF63A87F5B49FF13B42D04A0F817" lastPageNumber="58" pageId="5" pageNumber="58">
<subSubSection id="C35F5DFEA87F5B49FF13B42D074CFDB7" pageId="5" pageNumber="58" type="nomenclature">
<paragraph id="8BFA0E75A87F5B49FF13B42D0499FDD2" blockId="5.[151,760,535,596]" box="[151,760,535,561]" pageId="5" pageNumber="58">
<heading id="D0B2B919A87F5B49FF13B42D0499FDD2" bold="true" box="[151,760,535,561]" fontSize="11" level="1" pageId="5" pageNumber="58" reason="1">
<emphasis id="B931D267A87F5B49FF13B42D0499FDD2" bold="true" box="[151,760,535,561]" pageId="5" pageNumber="58">
<taxonomicName id="4C4575F6A87F5B49FF13B42D04EFFDD2" authority="Lelej and Ullah" authorityName="Lelej and Ullah" box="[151,654,535,561]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="himalayana" status="sp. nov.">
<emphasis id="B931D267A87F5B49FF13B42D07A2FDD2" bold="true" box="[151,451,535,561]" italics="true" pageId="5" pageNumber="58">Ephutomma himalayana</emphasis>
Lelej and Ullah
</taxonomicName>
,
<taxonomicNameLabel id="A2026F1CA87F5B49FD1FB42D0499FDD2" box="[667,760,535,561]" pageId="5" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA0E75A87F5B49FF13B400074CFDB7" blockId="5.[151,760,535,596]" box="[151,301,570,596]" pageId="5" pageNumber="58">
(
<figureCitation id="137E12F0A87F5B49FF1BB4000744FDB7" box="[159,293,570,596]" captionStart="FIGURES 16 25" captionStartId="6.[151,269,1188,1212]" captionTargetBox="[292,1290,199,1163]" captionTargetId="figure@6.[283,1304,194,1164]" captionTargetPageId="6" captionText="FIGURES 16 25. Ephutomma himalayana sp. nov., male, paratype. 16. Forewing; 17. Pedicel and flagellomeres 1 3, ventral view; 18 21. Mandible (18 outer view, 19 inner view, 20 ventral view, 21 dorsal view); 22 25. Genitalia (22 lateral view, 23 ventral view, 24 dorsal view, 25 volsella)." httpUri="https://zenodo.org/record/176102/files/figure.png" pageId="5" pageNumber="58">Figs 1625</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F5DFEA87F5B49FF13B4B00213FCA7" pageId="5" pageNumber="58" type="diagnosis">
<paragraph id="8BFA0E75A87F5B49FF13B4B00213FCA7" blockId="5.[151,1437,650,2036]" pageId="5" pageNumber="58">
<emphasis id="B931D267A87F5B49FF13B4B00776FD47" bold="true" box="[151,279,650,676]" pageId="5" pageNumber="58">Diagnosis.</emphasis>
The male of
<taxonomicName id="4C4575F6A87F5B49FE36B4B004B0FD40" box="[434,721,650,675]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="himalayana">
<emphasis id="B931D267A87F5B49FE36B4B004B0FD40" box="[434,721,650,675]" italics="true" pageId="5" pageNumber="58">Ephutomma himalayana</emphasis>
</taxonomicName>
is most similar to that of
<taxonomicName id="4C4575F6A87F5B49FC78B4B00345FD47" authority="Lelej 1977" authorityName="Lelej" authorityYear="1977" box="[1020,1316,650,676]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="kerzhneri">
<emphasis id="B931D267A87F5B49FC78B4B002EAFD40" box="[1020,1163,650,675]" italics="true" pageId="5" pageNumber="58">E. kerzhneri</emphasis>
(Lelej 1977)
</taxonomicName>
[
<typeStatus id="54FEB0D7A87F5B49FAB6B4B003FDFD47" box="[1330,1436,650,676]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
is examined, deposited in ZIN] in the key to the Palaearctic species (
<bibRefCitation id="EFD47384A87F5B49FC3EB488025FFD2F" author="Lelej" box="[954,1086,690,716]" pageId="5" pageNumber="60" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighboring countries]. Nauka, Leningrad. 268 pp., 64 tables. (In Russian)." type="book" year="1985">Lelej 1985</bibRefCitation>
). It can be distinguished from the latter species in having an additional subapical tooth on the ventral mandibular margin (lacking in
<taxonomicName id="4C4575F6A87F5B49FA07B4E00767FCF8" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="kerzhneri">
<emphasis id="B931D267A87F5B49FA07B4E00767FCF8" italics="true" pageId="5" pageNumber="58">E. kerzhneri</emphasis>
</taxonomicName>
), in having a subbasal ventral mandibular tooth that is unidentate apically (bidentate in
<taxonomicName id="4C4575F6A87F5B49FAABB53806A7FCA0" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="kerzhneri">
<emphasis id="B931D267A87F5B49FAABB53806A7FCA0" italics="true" pageId="5" pageNumber="58">E. kerzhneri</emphasis>
</taxonomicName>
), and in having infuscated wings (hyaline with infuscated apex in
<taxonomicName id="4C4575F6A87F5B49FC56B5100200FCA0" box="[978,1121,810,835]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="kerzhneri">
<emphasis id="B931D267A87F5B49FC56B5100200FCA0" box="[978,1121,810,835]" italics="true" pageId="5" pageNumber="58">E. kerzhneri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F5DFEA87F5B49FF42B56807A7F957" pageId="5" pageNumber="58" type="description">
<paragraph id="8BFA0E75A87F5B49FF42B56805BBFAAF" blockId="5.[151,1437,650,2036]" pageId="5" pageNumber="58">
<emphasis id="B931D267A87F5B49FF42B568073EFC8F" bold="true" box="[198,351,850,876]" pageId="5" pageNumber="58">Description.</emphasis>
MALE. Body length
<quantity id="4CBDA390A87F5B49FDDAB568048AFC8F" box="[606,747,850,876]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.6" metricValueMin="5.4" pageId="5" pageNumber="58" unit="mm" value="7.5" valueMax="9.6" valueMin="5.4">5.49.6 mm</quantity>
(
<typeStatus id="54FEB0D7A87F5B49FD7DB5680500FC8F" box="[761,865,850,876]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
<quantity id="4CBDA390A87F5B49FCFAB56805BDFC8F" box="[894,988,850,876]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="58" unit="mm" value="7.2">7.2 mm</quantity>
). Head width 1.2X head height. Ocelli small, POD: OOD 0.50.6X. Clypeus with anterior shiny depression bordered by arcuate carina which ends anteriorly in two tubercles. Occipital carina weak. Hypostomal carina weakly dilated. Mandible dilated, with inner tooth and additional subapical ventral tooth, bidentate apically, excised beneath, subbasal ventral tooth with one apex (
<figureCitation id="137E12F0A87F5B49FECDB5C807B3FBEF" box="[329,466,1010,1036]" captionStart="FIGURES 16 25" captionStartId="6.[151,269,1188,1212]" captionTargetBox="[292,1290,199,1163]" captionTargetId="figure@6.[283,1304,194,1164]" captionTargetPageId="6" captionText="FIGURES 16 25. Ephutomma himalayana sp. nov., male, paratype. 16. Forewing; 17. Pedicel and flagellomeres 1 3, ventral view; 18 21. Mandible (18 outer view, 19 inner view, 20 ventral view, 21 dorsal view); 22 25. Genitalia (22 lateral view, 23 ventral view, 24 dorsal view, 25 volsella)." httpUri="https://zenodo.org/record/176102/files/figure.png" pageId="5" pageNumber="58">Figs 1821</figureCitation>
). Relation of pedicel and three first flagellomeres 0.5: 1.0: 2.0: 2.0. Frons without process between antennal tubercles. Ocellar area weakly elevated. Frons, vertex, pronotum, mesoscutum, scutellum, and mesopleura densely punctate. Mesoscutum with complete notauli and short parapsidal furrows. Parascutal carinae well developed. Propodeum reticulate with median elongated and two rounded cells on dorsum. Mesopleura beneath without precoxal tubercle. Posterior coxae carinate inside, carina not dentate apically. Tegulae punctate, disc and posterior margin glabrous. Forewing venation as in
<figureCitation id="137E12F0A87F5B49FB3EB2800368FB37" box="[1210,1289,1210,1236]" captionStart="FIGURES 16 25" captionStartId="6.[151,269,1188,1212]" captionTargetBox="[292,1290,199,1163]" captionTargetId="figure@6.[283,1304,194,1164]" captionTargetPageId="6" captionText="FIGURES 16 25. Ephutomma himalayana sp. nov., male, paratype. 16. Forewing; 17. Pedicel and flagellomeres 1 3, ventral view; 18 21. Mandible (18 outer view, 19 inner view, 20 ventral view, 21 dorsal view); 22 25. Genitalia (22 lateral view, 23 ventral view, 24 dorsal view, 25 volsella)." httpUri="https://zenodo.org/record/176102/files/figure.png" pageId="5" pageNumber="58">Fig.16</figureCitation>
. S1 carinate beneath. Metasomal segment 2 densely punctate, punctures sparser on tergal disc. T2 with long lateral felt line. S2 without any trace of felt line. T3T6 densely punctate. T7 densely punctate, apically with median glabrous area. Genitalia laterally, dorsally, and ventrally as in
<figureCitation id="137E12F0A87F5B49FCCCB30805B7FAAF" box="[840,982,1330,1356]" captionStart="FIGURES 16 25" captionStartId="6.[151,269,1188,1212]" captionTargetBox="[292,1290,199,1163]" captionTargetId="figure@6.[283,1304,194,1164]" captionTargetPageId="6" captionText="FIGURES 16 25. Ephutomma himalayana sp. nov., male, paratype. 16. Forewing; 17. Pedicel and flagellomeres 1 3, ventral view; 18 21. Mandible (18 outer view, 19 inner view, 20 ventral view, 21 dorsal view); 22 25. Genitalia (22 lateral view, 23 ventral view, 24 dorsal view, 25 volsella)." httpUri="https://zenodo.org/record/176102/files/figure.png" pageId="5" pageNumber="58">Figs. 2225</figureCitation>
.
</paragraph>
<paragraph id="8BFA0E75A87F5B49FF42B360042CF96F" blockId="5.[151,1437,650,2036]" pageId="5" pageNumber="58">Body, legs, and antennae black; pronotum, mesoscutum, scutellum, tegulae, metanotum, upper part of mesopleura, and upper part of metapleura red. Wings infuscated. Mandibles preapically piceous. Fore tibial spurs yellowish, mid- and hind tibial spurs whitish. Lower frons, genae, antennal tubercles, scape, tibiae, and tarsi covered with dense, recumbent short and scattered, long, erect whitish pubescence. Upper frons, vertex, pronotum, scutellum, tegulae, metanotum medially, propodeum, T1T8, S1S8, and femorae covered with sparse whitish setae. Mesoscutum with sparse black setae. Scutellum and metanotum laterally covered with dense recumbent silvery micropubescence. T1T3 with apical fringe of dense yellowish setae. S2S5 with apical fringes of sparse whitish setae.</paragraph>
<paragraph id="8BFA0E75A87F5B49FF42B0A007A7F957" blockId="5.[151,1437,650,2036]" box="[198,454,1690,1716]" pageId="5" pageNumber="58">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C35F5DFEA87F5B49FF42B0F80599F89F" pageId="5" pageNumber="58" type="materials_examined">
<paragraph id="8BFA0E75A87F5B49FF42B0F80599F89F" blockId="5.[151,1437,650,2036]" pageId="5" pageNumber="58">
<emphasis id="B931D267A87F5B49FF42B0F8071EF93F" bold="true" box="[198,383,1730,1756]" pageId="5" pageNumber="58">
<typeStatus id="54FEB0D7A87F5B49FF42B0F80762F93F" box="[198,259,1730,1756]" pageId="5" pageNumber="58">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54FEB0D7A87F5B49FE1CB0F80464F93F" box="[408,517,1730,1756]" pageId="5" pageNumber="58" type="holotype">Holotype</typeStatus>
♂,
<collectingCountry id="F3524EE5A87F5B49FDC5B0F804EBF93F" box="[577,650,1730,1756]" name="Nepal" pageId="5" pageNumber="58">Nepal</collectingCountry>
, Chitwan National Park, Sauraha,
<geoCoordinate id="EE7168B2A87F5B49FBD4B0F802D5F93F" box="[1104,1204,1730,1756]" direction="north" orientation="latitude" pageId="5" pageNumber="58" precision="925" value="27.566668">27°34'N</geoCoordinate>
,
<geoCoordinate id="EE7168B2A87F5B49FB4EB0F8034BF93F" box="[1226,1322,1730,1756]" direction="east" orientation="longitude" pageId="5" pageNumber="58" precision="925" value="84.48333">84°29'E</geoCoordinate>
,
<quantity id="4CBDA390A87F5B49FAC4B0F803F9F93F" box="[1344,1432,1730,1756]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="58" unit="m" value="200.0">200 m</quantity>
,
<date id="FFFB28B5A87F5B49FF13B0D00773F8E7" box="[151,274,1770,1796]" pageId="5" pageNumber="58" value="1997-05-31">31.V.1997</date>
, M. Hauser [Oberoesterreichisches Landesmuseum Linz,
<collectingCountry id="F3524EE5A87F5B49FC5DB0D00257F8E7" box="[985,1078,1770,1796]" name="Austria" pageId="5" pageNumber="58">Austria</collectingCountry>
].
<typeStatus id="54FEB0D7A87F5B49FBCAB0D002A5F8E7" box="[1102,1220,1770,1796]" pageId="5" pageNumber="58" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F3524EE5A87F5B49FB50B0D0037EF8E7" box="[1236,1311,1770,1796]" name="Nepal" pageId="5" pageNumber="58">Nepal</collectingCountry>
: the same label as
<typeStatus id="54FEB0D7A87F5B49FF7FB1280705F8CF" box="[251,356,1810,1836]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D43C5FCA87F5B49FEF5B12807FAF8C6" box="[369,411,1810,1836]" pageId="5" pageNumber="58" type="generic">1 ♂</specimenCount>
[
<collectionCode id="ED5496B0A87F5B49FE28B1280793F8CF" box="[428,498,1810,1836]" pageId="5" pageNumber="58">IBSS</collectionCode>
]; the same place,
<date id="FFFB28B5A87F5B49FD43B12805E8F8CF" box="[711,905,1810,1836]" pageId="5" pageNumber="58" value="1997-05-31" valueMax="1997-06-04" valueMin="1997-05-31">31.V4.VI.1997</date>
, M. Hauser,
<specimenCount id="9D43C5FCA87F5B49FBA0B128022FF8C6" box="[1060,1102,1810,1836]" pageId="5" pageNumber="58" type="generic">3 ♂</specimenCount>
[col. Schmid-Egger,
<collectionCode id="ED5496B0A87F5B49FAC9B12803F0F8CF" box="[1357,1425,1810,1836]" pageId="5" pageNumber="58">IBSS</collectionCode>
].
<collectingCountry id="F3524EE5A87F5B49FF13B100069EF8B7" box="[151,255,1850,1876]" name="Pakistan" pageId="5" pageNumber="58">Pakistan</collectingCountry>
:
<taxonomicName id="4C4575F6A87F5B49FE8FB1000715F8B7" box="[267,372,1850,1876]" class="Insecta" family="Mutillidae" genus="Smicromyrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="55" phylum="Arthropoda" rank="species" species="margalla" status="sp. nov.">Margalla</taxonomicName>
Hills National Park, 1,
<date id="FFFB28B5A87F5B49FD14B1000565F8B7" box="[656,772,1850,1876]" pageId="5" pageNumber="58" value="1913-06-23">13, 23.VI</date>
, 1,
<date id="FFFB28B5A87F5B49FCABB100051EF8B7" box="[815,895,1850,1876]" pageId="5" pageNumber="58" value="2005-07">12.VII</date>
,
<date id="FFFB28B5A87F5B49FC09B1000267F8B7" box="[909,1030,1850,1876]" pageId="5" pageNumber="58" value="2005-09-08">8.IX.2005</date>
, MT, H. Khurram &amp; M. Fida,
<specimenCount id="9D43C5FCA87F5B49FAF7B10003FDF8AE" box="[1395,1436,1850,1876]" pageId="5" pageNumber="58" type="generic">6 ♂</specimenCount>
[
<collectionCode id="ED5496B0A87F5B49FF19B1580699F89F" box="[157,248,1890,1916]" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="5" pageNumber="58">PMNH</collectionCode>
,
<collectionCode id="ED5496B0A87F5B49FE80B1580726F89F" box="[260,327,1890,1916]" pageId="5" pageNumber="58">IBSS</collectionCode>
];
<date id="FFFB28B5A87F5B49FEDFB1580796F89F" box="[347,503,1890,1916]" pageId="5" pageNumber="58" value="2006-08-25">25.VIII.2006</date>
, MT, H. Khurram &amp; M. Fida,
<specimenCount id="9D43C5FCA87F5B49FCDDB15805E0F896" box="[857,897,1890,1916]" pageId="5" pageNumber="58" type="generic">1 ♂</specimenCount>
[
<collectionCode id="ED5496B0A87F5B49FC14B158058FF89F" box="[912,1006,1890,1916]" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="5" pageNumber="58">PMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="C35F5DFEA87F5B49FF42B1B0044CF847" box="[198,557,1930,1956]" pageId="5" pageNumber="58" type="distribution">
<paragraph id="8BFA0E75A87F5B49FF42B1B0044CF847" blockId="5.[151,1437,650,2036]" box="[198,557,1930,1956]" pageId="5" pageNumber="58">
<emphasis id="B931D267A87F5B49FF42B1B00706F847" bold="true" box="[198,359,1930,1956]" pageId="5" pageNumber="58">Distribution.</emphasis>
<collectingCountry id="F3524EE5A87F5B49FEE9B1B007B5F847" box="[365,468,1930,1956]" name="Pakistan" pageId="5" pageNumber="58">Pakistan</collectingCountry>
,
<collectingCountry id="F3524EE5A87F5B49FE5BB1B00448F847" box="[479,553,1930,1956]" name="Nepal" pageId="5" pageNumber="58">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F5DFEA87F5B49FF42B18804A0F817" pageId="5" pageNumber="58" type="etymology">
<paragraph id="8BFA0E75A87F5B49FF42B18804A0F817" blockId="5.[151,1437,650,2036]" pageId="5" pageNumber="58">
<emphasis id="B931D267A87F5B49FF42B1880732F82F" bold="true" box="[198,339,1970,1996]" pageId="5" pageNumber="58">Etymology.</emphasis>
The specific name is a Latin adjective derived from the Himalaya Mountain range, referring to the region where the species has been found.
</paragraph>
</subSubSection>
</treatment>
</document>