treatments-xml/data/03/E2/CA/03E2CA32FFE3FF8313E2FAB42FCAFDF8.xml
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<document id="E3729C7DEBBEA173E75985A79E479ACB" ID-CLB-Dataset="29292" ID-DOI="10.11646/zootaxa.4452.1.1" ID-GBIF-Dataset="30d459c8-2b2f-4cdf-8637-460394de28ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="1445140" ID-ZooBank="CDDB3757-1416-42B3-950B-4DC6A48239A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538636285412" checkinUser="plazi" docAuthor="Komarek, Albrecht &amp; Hebauer, Franz" docDate="2018" docId="03E2CA32FFE3FF8313E2FAB42FCAFDF8" docLanguage="en" docName="zootaxa.4452.1.1.pdf" docOrigin="Zootaxa 4452 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Agraphydrus orientalis" docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFDBB24AFFA3FFC21375FFD12E3DFFA7" masterDocTitle="Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae)" masterLastPageNumber="101" masterPageNumber="1" pageNumber="65" updateTime="1698564763264" updateUser="ExternalLinkService">
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<mods:title id="838E2405B90D5887D10AA909B0276B07">Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
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<mods:namePart id="4EB7C655EFB2A5960C102EB718048B81">Komarek, Albrecht</mods:namePart>
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<mods:namePart id="D05E4A948BA0EC45A31E46920665F09D">Hebauer, Franz</mods:namePart>
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<treatment id="03E2CA32FFE3FF8313E2FAB42FCAFDF8" ID-DOI="http://doi.org/10.5281/zenodo.5998016" ID-GBIF-Taxon="148462606" ID-Zenodo-Dep="5998016" LSID="urn:lsid:plazi:treatment:03E2CA32FFE3FF8313E2FAB42FCAFDF8" httpUri="http://treatment.plazi.org/id/03E2CA32FFE3FF8313E2FAB42FCAFDF8" lastPageId="65" lastPageNumber="65" pageId="64" pageNumber="65">
<subSubSection id="C35128AFFFE3FF8213E2FAB42C9FFAD8" box="[151,674,1381,1407]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="8BF47B24FFE3FF8213E2FAB42C9FFAD8" blockId="64.[151,674,1381,1407]" box="[151,674,1381,1407]" pageId="64" pageNumber="65">
<heading id="D0BCCC48FFE3FF8213E2FAB42C9FFAD8" bold="true" box="[151,674,1381,1407]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
<taxonomicName id="4C4B00A7FFE3FF8213E2FAB42C9FFAD8" ID-CoL="B3BC" authority="(Orchymont, 1932)" baseAuthorityName="Orchymont" baseAuthorityYear="1932" box="[151,674,1381,1407]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93FA736FFE3FF8213E2FAB42C9FFAD8" bold="true" box="[151,674,1381,1407]" pageId="64" pageNumber="65">
<emphasis id="B93FA736FFE3FF8213E2FAB42F93FAD8" bold="true" box="[151,430,1381,1407]" italics="true" pageId="64" pageNumber="65">Agraphydrus orientalis</emphasis>
(
<bibRefCitation id="EFDA06D5FFE3FF8212C8FAB42CA7FAD8" box="[445,666,1381,1407]" pageId="64" pageNumber="65" refId="ref80214" refString="Orchymont, A. (1932) Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement, Band IX (Tropische Binnengewasser II), 623 - 714, pls. XIV ¯ XVII." type="book chapter">Orchymont, 1932</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35128AFFFE3FF8213E2FA7D2CC7FA46" pageId="64" pageNumber="65" type="reference_group">
<paragraph id="8BF47B24FFE3FF8213E2FA7D2CC7FA65" blockId="64.[151,762,1452,1505]" box="[151,762,1452,1474]" pageId="64" pageNumber="65">
<treatmentCitationGroup id="AB5B5C0AFFE3FF8213E2FA7D2CC7FA65" box="[151,762,1452,1474]" pageId="64" pageNumber="65">
<taxonomicName id="4C4B00A7FFE3FF8213E2FA7D2CCBFA65" ID-CoL="3KGP8" authority="Orchymont 1932: 690" authorityName="Orchymont" authorityPageNumber="690" authorityYear="1932" box="[151,758,1452,1474]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis" subGenus="Agraphydrus">
<emphasis id="B93FA736FFE3FF8213E2FA7D2F36FA65" box="[151,267,1452,1474]" italics="true" pageId="64" pageNumber="65">Helochares</emphasis>
(
<emphasis id="B93FA736FFE3FF82126CFA7D2FA0FA65" box="[281,413,1452,1474]" italics="true" pageId="64" pageNumber="65">Agraphydrus</emphasis>
)
<emphasis id="B93FA736FFE3FF8212D9FA7D2C30FA65" box="[428,525,1452,1474]" italics="true" pageId="64" pageNumber="65">orientalis</emphasis>
<treatmentCitation id="0AEA5D35FFE3FF821161FA7D2CCBFA65" author="" box="[532,758,1452,1474]" page="690" pageId="64" pageNumber="65">
<bibRefCitation id="EFDA06D5FFE3FF821161FA7D2CFEFA65" box="[532,707,1452,1474]" pageId="64" pageNumber="65" refId="ref80214" refString="Orchymont, A. (1932) Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement, Band IX (Tropische Binnengewasser II), 623 - 714, pls. XIV ¯ XVII." type="book chapter">Orchymont 1932</bibRefCitation>
: 690
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF47B24FFE3FF8213E2FA1A2CC7FA46" blockId="64.[151,762,1452,1505]" box="[151,762,1483,1505]" pageId="64" pageNumber="65">
<treatmentCitationGroup id="AB5B5C0AFFE3FF8213E2FA1A2CC7FA46" box="[151,762,1483,1505]" pageId="64" pageNumber="65">
<taxonomicName id="4C4B00A7FFE3FF8213E2FA1A2C72FA46" ID-CoL="B3BC" authority="(Orchymont, 1932)" baseAuthorityName="Orchymont" baseAuthorityYear="1932" box="[151,591,1483,1505]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93FA736FFE3FF8213E2FA1A2FBFFA46" box="[151,386,1483,1505]" italics="true" pageId="64" pageNumber="65">Agraphydrus orientalis</emphasis>
(
<bibRefCitation id="EFDA06D5FFE3FF8212FAFA1A2C75FA46" box="[399,584,1483,1505]" pageId="64" pageNumber="65" refId="ref80214" refString="Orchymont, A. (1932) Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement, Band IX (Tropische Binnengewasser II), 623 - 714, pls. XIV ¯ XVII." type="book chapter">Orchymont, 1932</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="0AEA5D35FFE3FF82112CFA1A2CCBFA46" author="" box="[601,758,1483,1505]" page="128" pageId="64" pageNumber="65">
<bibRefCitation id="EFDA06D5FFE3FF82112CFA1A2CFEFA46" box="[601,707,1483,1505]" pageId="64" pageNumber="65" refId="ref80542" refString="Sato, M. (1965) Some aquatic Coleoptera from Formosa, I. Special Bulletin fo the Lepidopterological Society of Japan, 1, 126 ¯ 129." type="journal article">Satô 1965</bibRefCitation>
: 128
</treatmentCitation>
.
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</paragraph>
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<paragraph id="8BF47B24FFE3FF8213E2F9DC2CE0F981" blockId="64.[151,1437,1549,2007]" box="[151,733,1549,1574]" pageId="64" pageNumber="65">
<materialsCitation id="3B237179FFE3FF8213E2F9DC2CE0F981" ID-GBIF-Occurrence="1914127182" box="[151,733,1549,1574]" country="Indonesia" location="East-Java" pageId="64" pageNumber="65" specimenCount="1" typeStatus="holotype">
<emphasis id="B93FA736FFE3FF8213E2F9DC2F07F981" bold="true" box="[151,314,1549,1574]" pageId="64" pageNumber="65">
<typeStatus id="54F0C586FFE3FF8213E2F9DC2EEFF981" box="[151,210,1549,1574]" pageId="64" pageNumber="65">Type</typeStatus>
Locality
</emphasis>
:
<collectingCountry id="F35C3BB4FFE3FF82123CF9DC2F84F981" box="[329,441,1549,1574]" name="Indonesia" pageId="64" pageNumber="65">Indonesia</collectingCountry>
,
<location id="8E942DFFFFE3FF8212B1F9DF2C0FF982" LSID="urn:lsid:plazi:treatment:03E2CA32FFE3FF8313E2FAB42FCAFDF8:8E942DFFFFE3FF8212B1F9DF2C0FF982" box="[452,562,1550,1573]" country="Indonesia" name="East-Java" pageId="64" pageNumber="65">East-Java</location>
, Ranoe Bedali.
</materialsCitation>
</paragraph>
<paragraph id="8BF47B24FFE3FF8213B2F9E32B17F8E0" blockId="64.[151,1437,1549,2007]" pageId="64" pageNumber="65">
<emphasis id="B93FA736FFE3FF8213B2F9E32C3BF9EC" bold="true" box="[199,518,1586,1611]" pageId="64" pageNumber="65">Type material: Holotype Ƌ</emphasis>
(ISNB): “[Ostjava] Ranoe Bedali \ Badequelle [handwritten] \ Exp. Thienemann \
<date id="FFF55DE4FFE3FF8213E2F9862EC4F9C8" box="[151,249,1623,1647]" pageId="64" pageNumber="65" value="1928-10-10">10 10 28</date>
[handwritten] [mounted on a yellow label: “Coll. R.I.Sc.N.B. / Java”] \ A. dOrchymont det. \
<taxonomicName id="4C4B00A7FFE3FF82166BF9872F01F934" authority="(Agraphydrus)" authorityName="Agraphydrus" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Helochares (Agraphydrus)</taxonomicName>
\
<taxonomicName id="4C4B00A7FFE3FF821220F9AB2F80F934" baseAuthorityName="Orchymont" baseAuthorityYear="1932" box="[341,445,1658,1683]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis">orientalis</taxonomicName>
m. | Type [red label]”.
<emphasis id="B93FA736FFE3FF8211BEF9AB2D7EF934" bold="true" box="[715,835,1658,1683]" pageId="64" pageNumber="65">Paratypes</emphasis>
: INDONESIA: Java: 1 ♂ (ISNB): “Sarangaran ob. \ Telaga Woeroeng Wasserfall Wurung \
<date id="FFF55DE4FFE3FF821128F94E2CF9F910" box="[605,708,1695,1719]" pageId="64" pageNumber="65" value="1928-12-11">11 12 28</date>
[handwritten] \ Exp. Thienemann [mounted on a yellow label: “Coll. R.I.Sc.N.B. \ Java”] \
<taxonomicName id="4C4B00A7FFE3FF8212A6F9132C06F97C" baseAuthorityName="Orchymont" baseAuthorityYear="1932" box="[467,571,1730,1755]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis">orientalis</taxonomicName>
[handwritten] \ Paratype”; 1 ex.: “Sumatra Danaoe Toba \ Balige: Bach,
<quantity id="4CB3D6C1FFE3FF821610F9132E92F959" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="64" pageNumber="65" unit="m" value="1100.0">1100 m</quantity>
,
<date id="FFF55DE4FFE3FF8213B5F9372F11F958" box="[192,300,1766,1791]" pageId="64" pageNumber="65" value="1929-04-05">5 4 1929</date>
[handwritten] \ Exp. Thienemann [mounted on a yellow label: “Coll. R.I.Sc.N.B. / Sumatra”] \
<taxonomicName id="4C4B00A7FFE3FF8213E2F8DB2EC2F884" baseAuthorityName="Orchymont" baseAuthorityYear="1932" box="[151,255,1802,1827]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis">orientalis</taxonomicName>
Paratype [red label]”; 1 ex. (ISNB): “Bali Batoeriti \ Padegombobach
<date id="FFF55DE4FFE3FF82173DF8DB2A98F885" box="[1096,1189,1802,1827]" pageId="64" pageNumber="65" value="1929-06-14">14 6 29</date>
[handwritten] \ Exp. Thienemann [mounted on a yellow label: “Coll. R.I.Sc.N.B. / Indonesia”]
<taxonomicName id="4C4B00A7FFE3FF8210A2F8FF2A02F8E0" baseAuthorityName="Orchymont" baseAuthorityYear="1932" box="[983,1087,1838,1863]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis">orientalis</taxonomicName>
Paratype [red label].
</paragraph>
</subSubSection>
<subSubSection id="C35128AFFFE3FF8313B2F8832FCAFDF8" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="discussion">
<paragraph id="8BF47B24FFE3FF8313B2F8832FCAFDF8" blockId="64.[151,1437,1549,2007]" lastBlockId="65.[151,1437,151,608]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<emphasis id="B93FA736FFE3FF8213B2F8832F09F8CC" bold="true" box="[199,308,1874,1899]" pageId="64" pageNumber="65">Remarks</emphasis>
.
<bibRefCitation id="EFDA06D5FFE3FF821234F8832C2CF8CC" box="[321,529,1874,1899]" pageId="64" pageNumber="65" refId="ref80214" refString="Orchymont, A. (1932) Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement, Band IX (Tropische Binnengewasser II), 623 - 714, pls. XIV ¯ XVII." type="book chapter">Orchymont (1932)</bibRefCitation>
described
<taxonomicName id="4C4B00A7FFE3FF8211FCF8832A1BF8CC" authorityName="Orchymont" authorityYear="1932" box="[649,1062,1874,1899]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="orientalis" subGenus="Agraphydrus">
<emphasis id="B93FA736FFE3FF8211FCF8832D34F8CC" box="[649,777,1874,1899]" italics="true" pageId="64" pageNumber="65">Helochares</emphasis>
(
<emphasis id="B93FA736FFE3FF82106CF8832D96F8CC" box="[793,939,1874,1899]" italics="true" pageId="64" pageNumber="65">Agraphydrus</emphasis>
)
<emphasis id="B93FA736FFE3FF8210CEF8832A1BF8CC" box="[955,1062,1874,1899]" italics="true" pageId="64" pageNumber="65">orientalis</emphasis>
</taxonomicName>
based on several specimens from the Indonesian islands of
<collectingRegion id="498FB5C6FFE3FF8212C3F8A62C2EF828" box="[438,531,1911,1935]" country="Indonesia" name="Sumatera Utara" pageId="64" pageNumber="65">Sumatra</collectingRegion>
and
<collectingRegion id="498FB5C6FFE3FF82113FF8A62C43F829" box="[586,638,1911,1934]" country="Indonesia" name="Jawa Timur" pageId="64" pageNumber="65">Java</collectingRegion>
, and one specimen („Vergleichsmaterial“) from
<collectingRegion id="498FB5C6FFE3FF8217E9F8A62AC8F829" box="[1180,1269,1911,1934]" country="China" name="Yunnan" pageId="64" pageNumber="65">Yunnan</collectingRegion>
. The specimen from
<collectingRegion id="498FB5C6FFE3FF8213A1F84A2F14F815" box="[212,297,1947,1970]" country="China" name="Yunnan" pageId="64" pageNumber="65">Yunnan</collectingRegion>
could not be located in any of the collections listed above, the sex is not indicated, and genitalia were not described or figured. The study of the
<typeStatus id="54F0C586FFE3FF821104F86F2CEEF870" box="[625,723,1982,2007]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
and five
<typeStatus id="54F0C586FFE3FF821048F86E2D95F870" box="[829,936,1983,2007]" pageId="64" pageNumber="65" type="paratype">paratypes</typeStatus>
revealed that two of Orchymonts
<typeStatus id="54F0C586FFE3FF821644F86E2BA1F870" box="[1329,1436,1983,2007]" pageId="64" pageNumber="65" type="paratype">paratypes</typeStatus>
actually belong to a new species (which will be described in a separate part of the revision of
<taxonomicName id="4C4B00A7FFE2FF8317C6FF462B78FF17" authorityName="Komarek &amp; Hebauer" authorityYear="2018" box="[1203,1349,151,176]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B93FA736FFE2FF8317C6FF462B78FF17" box="[1203,1349,151,176]" italics="true" pageId="65" pageNumber="66">Agraphydrus</emphasis>
</taxonomicName>
). In his description
<bibRefCitation id="EFDA06D5FFE2FF83126AFF6D2FC8FF73" box="[287,501,188,213]" pageId="65" pageNumber="66" refId="ref80214" refString="Orchymont, A. (1932) Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement, Band IX (Tropische Binnengewasser II), 623 - 714, pls. XIV ¯ XVII." type="book chapter">Orchymont (1932)</bibRefCitation>
compares
<taxonomicName id="4C4B00A7FFE2FF83110DFF6D2D3AFF72" authorityName="Orchymont" authorityYear="1932" box="[632,775,188,213]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93FA736FFE2FF83110DFF6D2D3AFF72" box="[632,775,188,213]" italics="true" pageId="65" pageNumber="66">A. orientalis</emphasis>
</taxonomicName>
with the few species from the Indian Subcontinent and Africa known at his time. Since that time many specimens of
<taxonomicName id="4C4B00A7FFE2FF831029FF312DD5FF5F" authorityName="Orchymont" authorityYear="1932" box="[860,1000,223,248]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93FA736FFE2FF831029FF312DD5FF5F" box="[860,1000,223,248]" italics="true" pageId="65" pageNumber="66">A. orientalis</emphasis>
</taxonomicName>
were found, exclusively in
<collectingCountry id="F35C3BB4FFE2FF83165DFF0E2BA4FF5F" box="[1320,1433,223,248]" name="Indonesia" pageId="65" pageNumber="66">Indonesia</collectingCountry>
. Material recently collected from twenty localities in
<collectingCountry id="F35C3BB4FFE2FF831078FED52D47FEBA" box="[781,890,260,285]" name="Indonesia" pageId="65" pageNumber="66">Indonesia</collectingCountry>
was studied by the first author.
<emphasis id="B93FA736FFE2FF83167FFED52F3EFEE7" italics="true" pageId="65" pageNumber="66">
Agrapyhdrus
<taxonomicName id="4C4B00A7FFE2FF8313E2FEF62F3EFEE7" authorityName="Orchymont" authorityYear="1932" box="[151,259,295,320]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orientalis">orientalis</taxonomicName>
</emphasis>
belongs to the group of species with almost entirely chagrinated clypeus and unicolored maxillary palpomere 4. These features are also present in the Chinese species
<taxonomicName id="4C4B00A7FFE2FF8310DCFE9D2A25FEC2" box="[937,1048,332,357]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="arduus">
<emphasis id="B93FA736FFE2FF8310DCFE9D2A25FEC2" box="[937,1048,332,357]" italics="true" pageId="65" pageNumber="66">A. arduus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4B00A7FFE2FF83175DFE9D2A8FFEC3" box="[1064,1202,332,356]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="connexus">
<emphasis id="B93FA736FFE2FF83175DFE9D2A8FFEC3" box="[1064,1202,332,356]" italics="true" pageId="65" pageNumber="66">A. connexus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4B00A7FFE2FF8317B6FE9D2B4EFEC2" authorityName="Komarek &amp; Hebauer" authorityYear="2018" box="[1219,1395,332,357]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="gracilipalpis">
<emphasis id="B93FA736FFE2FF8317B6FE9D2B4EFEC2" box="[1219,1395,332,357]" italics="true" pageId="65" pageNumber="66">A. gracilipalpis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4B00A7FFE2FF8316F1FE9D2EDDFE2F" authorityName="Komarek &amp; Hebauer" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="igneus">
<emphasis id="B93FA736FFE2FF8316F1FE9D2EDDFE2F" italics="true" pageId="65" pageNumber="66">A. igneus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4B00A7FFE2FF831387FEA12FABFE2F" authorityName="Komarek &amp; Hebauer" authorityYear="2018" box="[242,406,367,392]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="reticuliceps">
<emphasis id="B93FA736FFE2FF831387FEA12FABFE2F" box="[242,406,367,392]" italics="true" pageId="65" pageNumber="66">A. reticuliceps</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4B00A7FFE2FF8312AEFEA12CACFE2F" authorityName="Komarek &amp; Hebauer" authorityYear="2018" box="[475,657,367,392]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="schoenmanni">
<emphasis id="B93FA736FFE2FF8312AEFEA12CACFE2F" box="[475,657,367,392]" italics="true" pageId="65" pageNumber="66">A. schoenmanni</emphasis>
</taxonomicName>
. Externally,
<taxonomicName id="4C4B00A7FFE2FF831052FEA12D88FE2F" authorityName="Orchymont" authorityYear="1932" box="[807,949,367,392]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93FA736FFE2FF831052FEA12D88FE2F" box="[807,949,367,392]" italics="true" pageId="65" pageNumber="66">A. orientalis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4B00A7FFE2FF8317E2FEA12B1FFE2F" box="[1175,1314,368,392]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="connexus">
<emphasis id="B93FA736FFE2FF8317E2FEA12B1FFE2F" box="[1175,1314,368,392]" italics="true" pageId="65" pageNumber="66">A. connexus</emphasis>
</taxonomicName>
, distinctly differing by features of the aedeagus. The record of
<taxonomicName id="4C4B00A7FFE2FF8311A8FE452D5AFE0A" authorityName="Orchymont" authorityYear="1932" box="[733,871,404,429]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93FA736FFE2FF8311A8FE452D5AFE0A" box="[733,871,404,429]" italics="true" pageId="65" pageNumber="66">A. orientalis</emphasis>
</taxonomicName>
from
<collectingCountry id="F35C3BB4FFE2FF8310D9FE452DC3FE0A" box="[940,1022,404,429]" name="Taiwan" pageId="65" pageNumber="66">Taiwan</collectingCountry>
by
<bibRefCitation id="EFDA06D5FFE2FF83175DFE452A98FE0A" box="[1064,1189,404,429]" pageId="65" pageNumber="66" refId="ref80542" refString="Sato, M. (1965) Some aquatic Coleoptera from Formosa, I. Special Bulletin fo the Lepidopterological Society of Japan, 1, 126 ¯ 129." type="journal article">Satô (1965</bibRefCitation>
; see also
<bibRefCitation id="EFDA06D5FFE2FF83167AFE452EEDFE77" pageId="65" pageNumber="66" refId="ref79100" refString="Gentili, E., Hebauer, F., Jach, M. A., Ji, L. &amp; Schodl, S. (1995) Hydrophilidae: I. Check list of the Hydrophilinae recorded from China (Coleoptera). In: Jach, M. A. &amp; Ji, L. (Eds.), Water Beetles of China. Uol. I. Zoologisch-Botanische Gesellschaft in Osterreich und Wiener Coleopterologenverein, Wien, pp. 207 ¯ 219." type="journal article">
Gentili
<emphasis id="B93FA736FFE2FF831611FE442BA8FE0A" box="[1380,1429,404,429]" italics="true" pageId="65" pageNumber="66">et al</emphasis>
. 1995
</bibRefCitation>
) was based on one single female that could not be examined by the authors of the present study. The fact that a reliable determination of
<taxonomicName id="4C4B00A7FFE2FF8312A4FE0D2C63FE52" authorityName="Orchymont" authorityYear="1932" box="[465,606,476,501]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93FA736FFE2FF8312A4FE0D2C63FE52" box="[465,606,476,501]" italics="true" pageId="65" pageNumber="66">A. orientalis</emphasis>
</taxonomicName>
and its separation from similar species, especially from
<taxonomicName id="4C4B00A7FFE2FF831780FE0D2B42FE53" box="[1269,1407,476,500]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="connexus">
<emphasis id="B93FA736FFE2FF831780FE0D2B42FE53" box="[1269,1407,476,500]" italics="true" pageId="65" pageNumber="66">A. connexus</emphasis>
</taxonomicName>
is only possible by features of the aedeagus leaves the occurrence of this species in
<collectingCountry id="F35C3BB4FFE2FF83174EFE2E2A43FDBF" box="[1083,1150,511,536]" name="China" pageId="65" pageNumber="66">China</collectingCountry>
or
<collectingCountry id="F35C3BB4FFE2FF8317D3FE2E2AC5FDBF" box="[1190,1272,511,536]" name="Taiwan" pageId="65" pageNumber="66">Taiwan</collectingCountry>
very unlikely. The species is therefore excluded from the fauna of
<collectingCountry id="F35C3BB4FFE2FF8311A2FDF52D27FD9A" box="[727,794,548,573]" name="China" pageId="65" pageNumber="66">China</collectingCountry>
and
<collectingCountry id="F35C3BB4FFE2FF83103AFDF52D9CFD9A" box="[847,929,548,573]" name="Taiwan" pageId="65" pageNumber="66">Taiwan</collectingCountry>
and will be described in a future contribution of the revision of
<taxonomicName id="4C4B00A7FFE2FF83122BFD962FCDFDC7" authorityName="Komarek &amp; Hebauer" authorityYear="2018" box="[350,496,583,608]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B93FA736FFE2FF83122BFD962FCDFDC7" box="[350,496,583,608]" italics="true" pageId="65" pageNumber="66">Agraphydrus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>