528 lines
56 KiB
XML
528 lines
56 KiB
XML
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<mods:title id="6C51CD9FCC3CFF06334AF0EEB4B0B80D">Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
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<mods:namePart id="86CC583920AA87241FE4911E63F73D60">Komarek, Albrecht</mods:namePart>
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<mods:namePart id="F15FBD054C380A514B0175138570F162">Hebauer, Franz</mods:namePart>
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<mods:title id="C317A30D8F25106004006F0EAC3D705B">Zootaxa</mods:title>
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<mods:part id="771D2375647FA1895EA8CFC192888735">
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<mods:date id="724F26062D508F96AF4CC1422E1381C9">2018</mods:date>
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<mods:number id="4B76A3337BED0AC529A9BE4F75BD7196">4452</mods:number>
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<treatment id="03E2CA32FF82FFE113E2FD0B2B59FEBA" ID-DOI="http://doi.org/10.5281/zenodo.5998000" ID-GBIF-Taxon="148462615" ID-Zenodo-Dep="5998000" LSID="urn:lsid:plazi:treatment:03E2CA32FF82FFE113E2FD0B2B59FEBA" httpUri="http://treatment.plazi.org/id/03E2CA32FF82FFE113E2FD0B2B59FEBA" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
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<subSubSection id="C35128AFFF82FFE313E2FD0B2FDEFCB1" pageId="33" pageNumber="34" type="nomenclature">
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<paragraph id="8BF47B24FF82FFE313E2FD0B2CA5FD52" blockId="33.[151,664,730,790]" box="[151,664,730,757]" pageId="33" pageNumber="34">
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<heading id="D0BCCC48FF82FFE313E2FD0B2CA5FD52" bold="true" box="[151,664,730,757]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
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<taxonomicName id="4C4B00A7FF82FFE313E2FD0B2CA5FD52" ID-CoL="B39X" authority="(Orchymont, 1927)" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[151,664,730,757]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="coomani">
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<emphasis id="B93FA736FF82FFE313E2FD0B2CA5FD52" bold="true" box="[151,664,730,757]" pageId="33" pageNumber="34">
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<emphasis id="B93FA736FF82FFE313E2FD0B2F99FD53" bold="true" box="[151,420,730,756]" italics="true" pageId="33" pageNumber="34">Agraphydrus coomani</emphasis>
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(
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<bibRefCitation id="EFDA06D5FF82FFE312C6FD0A2CADFD52" box="[435,656,731,757]" pageId="33" pageNumber="34" refId="ref80141" refString="Orchymont, A. (1927) Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums. Journal of the Federated Malay States Museums, 13, 246 ¯ 252." type="journal article">Orchymont, 1927</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BF47B24FF82FFE313E2FD2C2FDEFCB1" blockId="33.[151,664,730,790]" box="[151,483,765,790]" pageId="33" pageNumber="34">(Figs. 17, 18, 19, 52, 97, 139)</paragraph>
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</subSubSection>
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||
<subSubSection id="C35128AFFF82FFE313E2FC942D08FC10" pageId="33" pageNumber="34" type="reference_group">
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<paragraph id="8BF47B24FF82FFE313E2FC942CCDFCFC" blockId="33.[151,821,837,951]" box="[151,752,837,859]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="AB5B5C0AFF82FFE313E2FC942CCDFCFC" box="[151,752,837,859]" pageId="33" pageNumber="34">
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<taxonomicName id="4C4B00A7FF82FFE313E2FC942CD1FCFC" ID-CoL="3KGP6" authority="Orchymont 1927: 248" authorityName="Orchymont" authorityPageNumber="248" authorityYear="1927" box="[151,748,837,859]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="coomani" subGenus="Agraphydrus">
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<emphasis id="B93FA736FF82FFE313E2FC942F36FCFC" box="[151,267,837,859]" italics="true" pageId="33" pageNumber="34">Helochares</emphasis>
|
||
(
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<emphasis id="B93FA736FF82FFE3126CFC942FA0FCFC" box="[281,413,837,859]" italics="true" pageId="33" pageNumber="34">Agraphydrus</emphasis>
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)
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<emphasis id="B93FA736FF82FFE312D9FC972C3EFCFC" box="[428,515,838,859]" italics="true" pageId="33" pageNumber="34">coomani</emphasis>
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<treatmentCitation id="0AEA5D35FF82FFE3117CFC942CD1FCFC" author="" box="[521,748,837,859]" page="248" pageId="33" pageNumber="34">
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<bibRefCitation id="EFDA06D5FF82FFE3117CFC942C87FCFC" box="[521,698,837,859]" pageId="33" pageNumber="34" refId="ref80141" refString="Orchymont, A. (1927) Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums. Journal of the Federated Malay States Museums, 13, 246 ¯ 252." type="journal article">Orchymont 1927</bibRefCitation>
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: 248
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||
</treatmentCitation>
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</taxonomicName>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF47B24FF82FFE313E2FCB52CC6FCDD" blockId="33.[151,821,837,951]" box="[151,763,868,890]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="AB5B5C0AFF82FFE313E2FCB52CC6FCDD" box="[151,763,868,890]" pageId="33" pageNumber="34">
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<taxonomicName id="4C4B00A7FF82FFE313E2FCB52C7BFCDD" ID-CoL="B39X" authority="(Orchymont, 1927)" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[151,582,868,890]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="coomani">
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<emphasis id="B93FA736FF82FFE313E2FCB52F45FCDD" box="[151,376,868,890]" italics="true" pageId="33" pageNumber="34">Agraphydrus coomani</emphasis>
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(
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<bibRefCitation id="EFDA06D5FF82FFE312F3FCB52C03FCDD" box="[390,574,868,890]" pageId="33" pageNumber="34" refId="ref80141" refString="Orchymont, A. (1927) Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums. Journal of the Federated Malay States Museums, 13, 246 ¯ 252." type="journal article">Orchymont, 1927</bibRefCitation>
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)
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</taxonomicName>
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||
;
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<treatmentCitation id="0AEA5D35FF82FFE31125FCB42CCAFCDD" author="" box="[592,759,868,890]" page="115" pageId="33" pageNumber="34">
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<bibRefCitation id="EFDA06D5FF82FFE31125FCB42CF8FCDD" box="[592,709,868,890]" pageId="33" pageNumber="34" refId="ref80704" refString="Watts, C. H. S. (1995) Revision of the Australasian Genera Agraphydrus Regimbart, Chasmogenus Sharp and Helochares Mulsant (Coleoptera, Hydrophilidae). Records of the South Australian Museum, 28 (1), 113 ¯ 130." type="journal article">Watts 1995</bibRefCitation>
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: 115
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BF47B24FF82FFE313E2FC522C43FC3E" blockId="33.[151,821,837,951]" box="[151,638,898,921]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="AB5B5C0AFF82FFE313E2FC522C43FC3E" box="[151,638,898,921]" pageId="33" pageNumber="34">
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<taxonomicName id="4C4B00A7FF82FFE313E2FC522C0CFC3E" ID-CoL="39WGL" authority="Matsui 1994: 217" authorityName="Matsui" authorityPageNumber="217" authorityYear="1994" box="[151,561,899,921]" class="Insecta" family="Hydrophilidae" genus="Enochrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis">
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<emphasis id="B93FA736FF82FFE313E2FC522F4BFC3E" box="[151,374,899,921]" italics="true" pageId="33" pageNumber="34">Enochrus ryukyuensis</emphasis>
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<treatmentCitation id="0AEA5D35FF82FFE31209FC522C0CFC3E" author="" box="[380,561,899,921]" page="217" pageId="33" pageNumber="34">
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<bibRefCitation id="EFDA06D5FF82FFE31209FC522FC3FC3E" box="[380,510,899,921]" pageId="33" pageNumber="34" refId="ref79766" refString="Matsui, E. (1994) Three new species of the genus Enochrus from Japan and Taiwan (Coleoptera: Hydrophilidae). Transactions of the Shikoku entomological Society, 20, 215 ¯ 220." type="journal article">Matsui 1994</bibRefCitation>
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||
: 217
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||
</treatmentCitation>
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</taxonomicName>
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||
.
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<emphasis id="B93FA736FF82FFE3114EFC532C59FC3E" bold="true" box="[571,612,898,921]" pageId="33" pageNumber="34">Syn</emphasis>
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||
.
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||
<emphasis id="B93FA736FF82FFE3111FFC522C45FC3F" bold="true" box="[618,632,899,920]" pageId="33" pageNumber="34">n</emphasis>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BF47B24FF82FFE313E2FC702D08FC10" blockId="33.[151,821,837,951]" box="[151,821,929,951]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="AB5B5C0AFF82FFE313E2FC702D08FC10" box="[151,821,929,951]" pageId="33" pageNumber="34">
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<taxonomicName id="4C4B00A7FF82FFE313E2FC702C05FC10" ID-CoL="39WGL" authority="(Matsui, 1994)" baseAuthorityName="Matsui" baseAuthorityYear="1994" box="[151,568,929,951]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis">
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<emphasis id="B93FA736FF82FFE313E2FC702FA4FC10" box="[151,409,929,951]" italics="true" pageId="33" pageNumber="34">Agraphydrus ryukyuensis</emphasis>
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||
(
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||
<bibRefCitation id="EFDA06D5FF82FFE312D3FC702C0CFC10" box="[422,561,929,951]" pageId="33" pageNumber="34" refId="ref79766" refString="Matsui, E. (1994) Three new species of the genus Enochrus from Japan and Taiwan (Coleoptera: Hydrophilidae). Transactions of the Shikoku entomological Society, 20, 215 ¯ 220." type="journal article">Matsui, 1994</bibRefCitation>
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||
)
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</taxonomicName>
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||
;
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||
<treatmentCitation id="0AEA5D35FF82FFE31136FC702D0FFC10" author="" box="[579,818,929,951]" page="208" pageId="33" pageNumber="34">
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<bibRefCitation id="EFDA06D5FF82FFE31136FC702CC2FC10" box="[579,767,929,951]" pageId="33" pageNumber="34" refId="ref79100" refString="Gentili, E., Hebauer, F., Jach, M. A., Ji, L. & Schodl, S. (1995) Hydrophilidae: I. Check list of the Hydrophilinae recorded from China (Coleoptera). In: Jach, M. A. & Ji, L. (Eds.), Water Beetles of China. Uol. I. Zoologisch-Botanische Gesellschaft in Osterreich und Wiener Coleopterologenverein, Wien, pp. 207 ¯ 219." type="journal article">
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||
Gentili
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<emphasis id="B93FA736FF82FFE311FAFC732C81FC10" box="[655,700,929,951]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
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||
. 1995
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||
</bibRefCitation>
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||
: 208
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||
</treatmentCitation>
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.
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||
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35128AFFF82FFE313E2FC352CEDF80F" pageId="33" pageNumber="34" type="materials_examined">
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<paragraph id="8BF47B24FF82FFE313E2FC352D31FC5A" blockId="33.[151,1436,996,2032]" box="[151,780,996,1021]" pageId="33" pageNumber="34">
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<materialsCitation id="3B237179FF82FFE313E2FC352D31FC5A" ID-GBIF-Occurrence="1914126991" box="[151,780,996,1021]" country="Vietnam" location="Lac Tho" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Hoa Binh Province" typeStatus="holotype">
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<emphasis id="B93FA736FF82FFE313E2FC352F10FC5A" bold="true" box="[151,301,996,1021]" pageId="33" pageNumber="34">
|
||
<typeStatus id="54F0C586FF82FFE313E2FC352EEFFC5A" box="[151,210,996,1021]" pageId="33" pageNumber="34">Type</typeStatus>
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||
locality
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||
</emphasis>
|
||
.
|
||
<collectingCountry id="F35C3BB4FF82FFE31249FC352FA3FC5A" box="[316,414,996,1021]" name="Vietnam" pageId="33" pageNumber="34">Vietnam</collectingCountry>
|
||
,
|
||
<collectingRegion id="498FB5C6FF82FFE312DCFC342CBEFC5A" box="[425,643,996,1021]" country="Vietnam" name="Hoa Binh" pageId="33" pageNumber="34">
|
||
Hoa
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||
<collectingRegion id="498FB5C6FF82FFE312ABFC352CBEFC5A" box="[478,643,996,1021]" inferredBySuffix="true" pageId="33" pageNumber="34">Binh Province</collectingRegion>
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</collectingRegion>
|
||
, “
|
||
<location id="8E942DFFFF82FFE311EDFC352CC0FC5A" LSID="urn:lsid:plazi:treatment:03E2CA32FF82FFE113E2FD0B2B59FEBA:8E942DFFFF82FFE311EDFC352CC0FC5A" box="[664,765,996,1021]" country="Vietnam" name="Lac Tho" pageId="33" pageNumber="34" stateProvince="Hoa Binh Province">Lac Tho</location>
|
||
”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF82FFE313B2FBD92D9DFB16" blockId="33.[151,1436,996,2032]" pageId="33" pageNumber="34">
|
||
<emphasis id="B93FA736FF82FFE313B2FBD92FD1FB86" bold="true" box="[199,492,1032,1057]" pageId="33" pageNumber="34">Type material examined</emphasis>
|
||
:
|
||
<taxonomicName id="4C4B00A7FF82FFE31288FBDB2CBCFB86" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[509,641,1034,1057]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="coomani">
|
||
<emphasis id="B93FA736FF82FFE31288FBDB2CBCFB86" box="[509,641,1034,1057]" italics="true" pageId="33" pageNumber="34">H. coomani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FA736FF82FFE311FEFBD92CC5FB86" bold="true" box="[651,760,1032,1057]" pageId="33" pageNumber="34">Holotype</emphasis>
|
||
<emphasis id="B93FA736FF82FFE31074FBDB2D2BFB86" box="[769,790,1034,1057]" italics="true" pageId="33" pageNumber="34">Ƌ</emphasis>
|
||
(ISNB): “Hoa Binh \ Tonkin \ de Cooman [on a yellow card “Coll. R. I. Sc. N. B. \ Nord Vietnam] | A. d’Orchymont det. \
|
||
<taxonomicName id="4C4B00A7FF82FFE310F0FBFD2A8CFBE2" authority="(Agraphydrus)" authorityName="Agraphydrus" box="[901,1201,1068,1093]" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FA736FF82FFE310F0FBFD2A38FBE2" box="[901,1029,1068,1093]" italics="true" pageId="33" pageNumber="34">Helochares</emphasis>
|
||
(
|
||
<emphasis id="B93FA736FF82FFE31763FBFD2A95FBE2" box="[1046,1192,1068,1093]" italics="true" pageId="33" pageNumber="34">Agraphydrus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
\
|
||
<emphasis id="B93FA736FF82FFE317B2FBFC2B10FBE3" box="[1223,1325,1069,1092]" italics="true" pageId="33" pageNumber="34">Coomani</emphasis>
|
||
m. Type | Type [red label]”. Tonkin is an inofficial name for the northern part of Vietnam. Hòa Bình is the name for a province and it’s capital in Northern Vietnam.
|
||
<bibRefCitation id="EFDA06D5FF82FFE311D9FBA52D42FB2A" box="[684,895,1140,1165]" pageId="33" pageNumber="34" refId="ref80141" refString="Orchymont, A. (1927) Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums. Journal of the Federated Malay States Museums, 13, 246 ¯ 252." type="journal article">Orchymont (1927)</bibRefCitation>
|
||
specifies the type locality as “Lac Tho nr. Hoa Binh”.
|
||
<emphasis id="B93FA736FF82FFE3139DFB492F5DFB16" bold="true" box="[232,352,1176,1201]" pageId="33" pageNumber="34">Paratypes</emphasis>
|
||
:
|
||
<emphasis id="B93FA736FF82FFE3121BFB492FC8FB16" bold="true" box="[366,501,1176,1201]" pageId="33" pageNumber="34">VIETNAM</emphasis>
|
||
: 23 exs. (ISNB): same sampling data.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF82FFE313B2FB6F2F36F9BD" blockId="33.[151,1436,996,2032]" pageId="33" pageNumber="34">
|
||
<taxonomicName id="4C4B00A7FF82FFE313B2FB6F2F56FB71" baseAuthorityName="Matsui" baseAuthorityYear="1994" box="[199,363,1213,1238]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis">
|
||
<emphasis id="B93FA736FF82FFE313B2FB6F2F56FB71" box="[199,363,1213,1238]" italics="true" pageId="33" pageNumber="34">A. ryukyuensis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B93FA736FF82FFE31209FB6C2FD4FB71" bold="true" box="[380,489,1213,1238]" pageId="33" pageNumber="34">Holotype</emphasis>
|
||
<emphasis id="B93FA736FF82FFE31287FB6F2C3AFB72" box="[498,519,1214,1237]" italics="true" pageId="33" pageNumber="34">Ƌ</emphasis>
|
||
(CEM): “[Japan] Kamize-Dam \ Tokunoshima T[own] \ Tokunoshima Is[land] \ Kagoshima Pref. [Ôshima Dist.] \
|
||
<date id="FFF55DE4FF82FFE3116DFB302C97FB5E" box="[536,682,1249,1273]" pageId="33" pageNumber="34" value="1987-08-18">18.VIII.1987</date>
|
||
\ leg. E. Matsui |
|
||
<taxonomicName id="4C4B00A7FF82FFE31006FB302A38FB5D" authorityName="Komarek & Hebauer" authorityYear="2018" box="[883,1029,1249,1274]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FA736FF82FFE31006FB302A38FB5D" box="[883,1029,1249,1274]" italics="true" pageId="33" pageNumber="34">Agraphydrus</emphasis>
|
||
</taxonomicName>
|
||
\
|
||
<taxonomicName id="4C4B00A7FF82FFE3176EFB302B3AFB5D" authority="(Matsui)" authorityName="Matsui" baseAuthorityName="Matsui" baseAuthorityYear="1994" box="[1051,1287,1249,1274]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis">
|
||
<emphasis id="B93FA736FF82FFE3176EFB302AA2FB5D" box="[1051,1183,1249,1274]" italics="true" pageId="33" pageNumber="34">ryukyuensis</emphasis>
|
||
(Matsui)
|
||
</taxonomicName>
|
||
\ det. Matsui E. | Holotype”.
|
||
<emphasis id="B93FA736FF82FFE31238FAD42FF8FAB9" bold="true" box="[333,453,1285,1310]" pageId="33" pageNumber="34">Paratypes</emphasis>
|
||
(CEM):
|
||
<emphasis id="B93FA736FF82FFE31145FAD42CB7FAB9" bold="true" box="[560,650,1285,1310]" pageId="33" pageNumber="34">JAPAN</emphasis>
|
||
: 2 exs. (CEM): same sampling data; 1
|
||
<emphasis id="B93FA736FF82FFE31723FAD72A56FABA" box="[1110,1131,1286,1309]" italics="true" pageId="33" pageNumber="34">Ƌ</emphasis>
|
||
(CEM): Kagoshima Pref., AmamīÔshima Isl., Tatsugou Town, Yairi,
|
||
<date id="FFF55DE4FF82FFE311FFFAF82D1DFAE6" box="[650,800,1321,1345]" pageId="33" pageNumber="34" value="1987-08-20">20.VIII.1987</date>
|
||
, leg. E. Matsui; 1 ex. (CEM): Kagoshima Pref., Amami- Ôshima Isl., Sumiyo Village, Kawauchi River,
|
||
<date id="FFF55DE4FF82FFE311BFFA9F2D62FAC1" box="[714,863,1358,1382]" pageId="33" pageNumber="34" value="1987-08-21">21.VIII.1987</date>
|
||
, E. Matsui; 2 exs. (CEM): Okinawa Prefecture, Kunigami District, Okinawa Island, Ôgimi Village, Henan River,
|
||
<date id="FFF55DE4FF82FFE310EBFAA02A0FFA2D" box="[926,1074,1393,1418]" pageId="33" pageNumber="34" value="1990-08-16">16.VIII.1990</date>
|
||
, leg. E. Matsui; 1 ex. (CEM): Okinawa Prefecture, Ishigaki Island, Sokobaru,
|
||
<date id="FFF55DE4FF82FFE311B7FA472D74FA09" box="[706,841,1430,1454]" pageId="33" pageNumber="34" value="1989-08-07">7.VIII.1989</date>
|
||
, leg. E. Matsui; 1 ex. (CEM): Okinawa Prefecture, Ishigaki Island, Nishihama River,
|
||
<date id="FFF55DE4FF82FFE31162FA682C91FA76" box="[535,684,1465,1489]" pageId="33" pageNumber="34" value="1989-08-12">12.VIII.1989</date>
|
||
, leg. E. Matsui; 1 ex. (CEM): Okinawa Prefecture, Ishigaki Island, Takada,
|
||
<date id="FFF55DE4FF82FFE31380FA0F2FB7FA51" box="[245,394,1502,1526]" pageId="33" pageNumber="34" value="1990-08-12">12.VIII.1990</date>
|
||
, leg. E. Matsui; 2 exs. (CEM): Okinawa Prefecture, Iriomote Island, Komi,
|
||
<date id="FFF55DE4FF82FFE31795FA0F2B5AFA51" box="[1248,1383,1502,1526]" pageId="33" pageNumber="34" value="1990-08-09">9.VIII.1990</date>
|
||
, leg. E. Matsui.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF82FFE313B2F9F72FD3F9C5" blockId="33.[151,1436,996,2032]" pageId="33" pageNumber="34">
|
||
<materialsCitation id="3B237179FF82FFE313B2F9F72FD3F9C5" ID-GBIF-Occurrence="1914126998" collectingDate="1992-12-25" collectorName="E. Matsui" country="Taiwan" location="Daikan River" municipality="Daikei-chin" pageId="33" pageNumber="34" specimenCount="17" typeStatus="paratype">
|
||
We have not seen the
|
||
<specimenCount id="9D4DB0ADFF82FFE312C2F9F72C78F999" box="[439,581,1574,1598]" pageId="33" pageNumber="34" type="generic" typeStatus="paratypes">17 paratypes</specimenCount>
|
||
from
|
||
<collectingCountry id="F35C3BB4FF82FFE311FDF9F42CE7F999" box="[648,730,1573,1598]" name="Taiwan" pageId="33" pageNumber="34">Taiwan</collectingCountry>
|
||
(CEM): “
|
||
<location id="8E942DFFFF82FFE3103DF9F42DDFF999" LSID="urn:lsid:plazi:treatment:03E2CA32FF82FFE113E2FD0B2B59FEBA:8E942DFFFF82FFE3103DF9F42DDFF999" box="[840,994,1573,1598]" country="Taiwan" municipality="Daikei-chin" name="Daikan River" pageId="33" pageNumber="34">Daikan River</location>
|
||
,
|
||
<collectingMunicipality id="6B90E15EFF82FFE31099F9F42A4EF999" box="[1004,1139,1573,1598]" pageId="33" pageNumber="34">Daikei-chin</collectingMunicipality>
|
||
, Tohen Pref.,
|
||
<collectingDate id="EFB1A40CFF82FFE31679F9F42BA5F999" box="[1292,1432,1573,1598]" pageId="33" pageNumber="34" value="1992-12-25">
|
||
<date id="FFF55DE4FF82FFE31679F9F42BA5F999" box="[1292,1432,1573,1598]" pageId="33" pageNumber="34" value="1992-12-25">25.XII.1992</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26BE1EF2FF82FFE313E2F99B2F39F9C5" box="[151,260,1609,1634]" pageId="33" pageNumber="34">E. Matsui</collectorName>
|
||
leg.” (
|
||
<bibRefCitation id="EFDA06D5FF82FFE3123AF9982FDFF9C5" box="[335,482,1609,1634]" pageId="33" pageNumber="34" refId="ref79766" refString="Matsui, E. (1994) Three new species of the genus Enochrus from Japan and Taiwan (Coleoptera: Hydrophilidae). Transactions of the Shikoku entomological Society, 20, 215 ¯ 220." type="journal article">Matsui 1994</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF82FFE313B2F9BF2D11F89C" blockId="33.[151,1436,996,2032]" pageId="33" pageNumber="34">
|
||
<materialsCitation id="3B237179FF82FFE313B2F9BF2A7AF920" ID-GBIF-Occurrence="1914126989" box="[199,1095,1646,1671]" collectionCode="NMW" country="China" location="Fujian" pageId="33" pageNumber="34" specimenCode="CWBS 255" specimenCount="1" stateProvince="Fujian">
|
||
<emphasis id="B93FA736FF82FFE313B2F9BF2C12F920" bold="true" box="[199,559,1646,1671]" pageId="33" pageNumber="34">Additional material examined</emphasis>
|
||
:
|
||
<collectingCountry id="F35C3BB4FF82FFE31135F9BF2CA0F920" box="[576,669,1646,1671]" name="China" pageId="33" pageNumber="34">
|
||
<emphasis id="B93FA736FF82FFE31135F9BF2CA0F920" bold="true" box="[576,669,1646,1671]" pageId="33" pageNumber="34">CHINA</emphasis>
|
||
</collectingCountry>
|
||
:
|
||
<collectingRegion id="498FB5C6FF82FFE311D8F9BF2CC1F920" box="[685,764,1646,1671]" country="China" name="Fujian" pageId="33" pageNumber="34">
|
||
<emphasis id="B93FA736FF82FFE311D8F9BF2CC1F920" bold="true" box="[685,764,1646,1671]" pageId="33" pageNumber="34">Fujian</emphasis>
|
||
</collectingRegion>
|
||
: 2 exs. (
|
||
<collectionCode id="ED5AE3E1FF82FFE31015F9BE2D92F920" box="[864,943,1647,1671]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="33" pageNumber="34">NMW</collectionCode>
|
||
):
|
||
<specimenCode id="DBEDD35FFF82FFE310B7F9BE2A7AF920" box="[962,1095,1646,1671]" collectionCode="CWBS" pageId="33" pageNumber="34">CWBS 255</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B237179FF82FFE31721F9BF2F23F90C" ID-GBIF-Occurrence="1914127012" collectionCode="NMW" country="China" location="Guangdong" pageId="33" pageNumber="34" specimenCode="CWBS 461" specimenCount="1" stateProvince="Guangdong">
|
||
<collectingRegion id="498FB5C6FF82FFE31721F9BF2ADDF920" box="[1108,1248,1646,1671]" country="China" name="Guangdong" pageId="33" pageNumber="34">
|
||
<emphasis id="B93FA736FF82FFE31721F9BF2ADDF920" bold="true" box="[1108,1248,1646,1671]" pageId="33" pageNumber="34">Guangdong</emphasis>
|
||
</collectingRegion>
|
||
: 2
|
||
<emphasis id="B93FA736FF82FFE31672F9BE2B0CF921" box="[1287,1329,1647,1670]" italics="true" pageId="33" pageNumber="34">ƋƋ</emphasis>
|
||
(
|
||
<collectionCode id="ED5AE3E1FF82FFE31637F9BE2BACF920" box="[1346,1425,1647,1671]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="33" pageNumber="34">NMW</collectionCode>
|
||
):
|
||
<specimenCode id="DBEDD35FFF82FFE313E2F9422F23F90C" box="[151,286,1682,1707]" collectionCode="CWBS" pageId="33" pageNumber="34">CWBS 461</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B237179FF82FFE31258F9432CF9F90C" ID-GBIF-Occurrence="1914126986" box="[301,708,1682,1707]" collectionCode="NMW" country="China" location="Hainan" pageId="33" pageNumber="34" specimenCode="CWBS 178" specimenCount="1" stateProvince="Hainan">
|
||
<collectingRegion id="498FB5C6FF82FFE31258F9432FB8F90C" box="[301,389,1682,1707]" country="China" name="Hainan" pageId="33" pageNumber="34">
|
||
<emphasis id="B93FA736FF82FFE31258F9432FB8F90C" bold="true" box="[301,389,1682,1707]" pageId="33" pageNumber="34">Hainan</emphasis>
|
||
</collectingRegion>
|
||
: 1
|
||
<emphasis id="B93FA736FF82FFE312C4F9422FFBF90D" box="[433,454,1683,1706]" italics="true" pageId="33" pageNumber="34">Ƌ</emphasis>
|
||
(
|
||
<collectionCode id="ED5AE3E1FF82FFE312ADF9422C1AF90C" box="[472,551,1683,1707]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="33" pageNumber="34">NMW</collectionCode>
|
||
):
|
||
<specimenCode id="DBEDD35FFF82FFE31148F9422CF9F90C" box="[573,708,1683,1707]" collectionCode="CWBS" pageId="33" pageNumber="34">CWBS 178</specimenCode>
|
||
</materialsCitation>
|
||
; 1
|
||
<emphasis id="B93FA736FF82FFE31199F9422D3CF90D" box="[748,769,1683,1706]" italics="true" pageId="33" pageNumber="34">Ƌ</emphasis>
|
||
(NMW): CWBS 187; 64 exs. (NMW): CWBS 192; 1
|
||
<emphasis id="B93FA736FF82FFE316F2F9422BA1F90D" box="[1415,1436,1683,1706]" italics="true" pageId="33" pageNumber="34">Ƌ</emphasis>
|
||
(NMW): CWBS 197; 11 exs. (NMW): CWBS 198; 3 exs. (NMW): CWBS 199; 2 exs. (NMW): CWBS 201; 54 exs. (NMW): CWBS 202; 2 exs. (NMW): CWBS 203; 15 exs. (NMW): CWBS 204; 5 exs. (NMW): CWBS 207; 3 exs. (NMW): CWBS 210; 31 exs. (NMW): CWBS 211; 3 exs. (NMW): CWBS 212; 7 exs. (NMW): CWBS 214; 50 exs. (NMW): CWBS 215; 12 exs. (NMW): CWBS 216.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF82FFE313B2F8972CEDF80F" blockId="33.[151,1436,996,2032]" pageId="33" pageNumber="34">
|
||
<collectingCountry id="F35C3BB4FF82FFE313B2F8972F0BF8F8" box="[199,310,1862,1887]" name="Taiwan" pageId="33" pageNumber="34">
|
||
<emphasis id="B93FA736FF82FFE313B2F8972F0BF8F8" bold="true" box="[199,310,1862,1887]" pageId="33" pageNumber="34">TAIWAN</emphasis>
|
||
</collectingCountry>
|
||
: 2 exs. (CEM): “Tahhan [= Dahan] River \
|
||
<collectingRegion id="498FB5C6FF82FFE31058F8992DEBF8F8" box="[813,982,1862,1888]" country="Taiwan" name="Taoyuan" pageId="33" pageNumber="34">Taoyuan Hsien</collectingRegion>
|
||
\
|
||
<collectingCountry id="F35C3BB4FF82FFE3109AF8962A66F8F8" box="[1007,1115,1863,1887]" name="Taiwan" pageId="33" pageNumber="34">TAIWAN</collectingCountry>
|
||
\
|
||
<date id="FFF55DE4FF82FFE31701F8972AC0F8F8" box="[1140,1277,1862,1887]" pageId="33" pageNumber="34" value="1992-12-25">25.XII.1992</date>
|
||
\ leg. Matsui, E.”. These specimens are almost certainly from the same locality as the other Taiwanese
|
||
<typeStatus id="54F0C586FF82FFE31791F8BD2B6DF823" box="[1252,1360,1900,1924]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B00A7FF82FFE316F1F8BD2F26F800" baseAuthorityName="Matsui" baseAuthorityYear="1994" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ryukyuensis">
|
||
<emphasis id="B93FA736FF82FFE316F1F8BD2F26F800" italics="true" pageId="33" pageNumber="34">A. ryukyuensis</emphasis>
|
||
</taxonomicName>
|
||
, but they are not labelled as
|
||
<typeStatus id="54F0C586FF82FFE31128F8412CF1F80F" box="[605,716,1936,1960]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35128AFFF82FFE013B2F8622ACCFEE7" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="diagnosis">
|
||
<paragraph id="8BF47B24FF82FFE313B2F8622BA1F857" blockId="33.[151,1436,996,2032]" pageId="33" pageNumber="34">
|
||
<emphasis id="B93FA736FF82FFE313B2F8622FF1F86B" bold="true" box="[199,460,1971,1996]" pageId="33" pageNumber="34">Differential diagnosis</emphasis>
|
||
. This species belongs to the group of species with apically infuscated maxillary palpomere 4 and absence of clypeal microsculpture, together with
|
||
<taxonomicName id="4C4B00A7FF82FFE310F1F8092DD4F848" box="[900,1001,2008,2031]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="comes">
|
||
<emphasis id="B93FA736FF82FFE310F1F8092DD4F848" box="[900,1001,2008,2031]" italics="true" pageId="33" pageNumber="34">A. comes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF82FFE31082F8092A44F857" box="[1015,1145,2008,2032]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="B93FA736FF82FFE31082F8092A44F857" box="[1015,1145,2008,2032]" italics="true" pageId="33" pageNumber="34">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF82FFE317F2F8092B31F857" baseAuthorityName="Matsui" baseAuthorityYear="1994" box="[1159,1292,2007,2032]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="B93FA736FF82FFE317F2F8092B31F857" box="[1159,1292,2007,2032]" italics="true" pageId="33" pageNumber="34">A. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF82FFE3166FF8092BA1F857" authorityName="Komarek & Hebauer" authorityYear="2018" box="[1306,1436,2007,2032]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="jilanzhui">
|
||
<emphasis id="B93FA736FF82FFE3166FF8092BA1F857" box="[1306,1436,2007,2032]" italics="true" pageId="33" pageNumber="34">A. jilanzhui</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013E2FF462ACCFEE7" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
and
|
||
<taxonomicName id="4C4B00A7FF81FFE013BDFF492F75FF17" authorityName="Komarek & Hebauer" authorityYear="2018" box="[200,328,151,176]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="B93FA736FF81FFE013BDFF492F75FF17" box="[200,328,151,176]" italics="true" pageId="34" pageNumber="35">A. robustus</emphasis>
|
||
</taxonomicName>
|
||
(see also Remarks under
|
||
<taxonomicName id="4C4B00A7FF81FFE01107FF492CCEFF17" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[626,755,152,176]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="coomani">
|
||
<emphasis id="B93FA736FF81FFE01107FF492CCEFF17" box="[626,755,152,176]" italics="true" pageId="34" pageNumber="35">A. coomani</emphasis>
|
||
</taxonomicName>
|
||
). Specimens from
|
||
<collectingCountry id="F35C3BB4FF81FFE010BBFF462A2CFF17" box="[974,1041,151,176]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
|
||
differ from
|
||
<taxonomicName id="4C4B00A7FF81FFE017EAFF492B1FFF17" authorityName="Komarek & Hebauer" authorityYear="2018" box="[1183,1314,151,176]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="jilanzhui">
|
||
<emphasis id="B93FA736FF81FFE017EAFF492B1FFF17" box="[1183,1314,151,176]" italics="true" pageId="34" pageNumber="35">A. jilanzhui</emphasis>
|
||
</taxonomicName>
|
||
by yellow pronotum with mesal infuscation (pronotum black in
|
||
<taxonomicName id="4C4B00A7FF81FFE01184FF6D2D4EFF72" authorityName="Komarek & Hebauer" authorityYear="2018" box="[753,883,188,213]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="jilanzhui">
|
||
<emphasis id="B93FA736FF81FFE01184FF6D2D4EFF72" box="[753,883,188,213]" italics="true" pageId="34" pageNumber="35">A. jilanzhui</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<taxonomicName id="4C4B00A7FF81FFE010BCFF6D2A75FF72" authorityName="Komarek & Hebauer" authorityYear="2018" box="[969,1096,188,213]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="B93FA736FF81FFE010BCFF6D2A75FF72" box="[969,1096,188,213]" italics="true" pageId="34" pageNumber="35">A. robustus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF81FFE01722FF6D2AE6FF72" baseAuthorityName="Matsui" baseAuthorityYear="1994" box="[1111,1243,188,213]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="B93FA736FF81FFE01722FF6D2AE6FF72" box="[1111,1243,188,213]" italics="true" pageId="34" pageNumber="35">A. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4B00A7FF81FFE0166FFF6D2BA1FF72" authorityName="Komarek & Hebauer" authorityYear="2018" box="[1306,1436,188,213]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="jilanzhui">
|
||
<emphasis id="B93FA736FF81FFE0166FFF6D2BA1FF72" box="[1306,1436,188,213]" italics="true" pageId="34" pageNumber="35">A. jilanzhui</emphasis>
|
||
</taxonomicName>
|
||
by absence of apical extension of parameres, and from all species by shape of parameres (apex slightly bending mesad) and by finer elytral ground punctation. Similar aedeagus is present in
|
||
<taxonomicName id="4C4B00A7FF81FFE01774FED52AA0FEBB" authorityName="Komarek & Hebauer" authorityYear="2018" box="[1025,1181,260,284]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="wangmiaoi">
|
||
<emphasis id="B93FA736FF81FFE01774FED52AA0FEBB" box="[1025,1181,260,284]" italics="true" pageId="34" pageNumber="35">A. wangmiaoi</emphasis>
|
||
</taxonomicName>
|
||
, a species with evenly oval body shape (versus widening posteriorly in
|
||
<taxonomicName id="4C4B00A7FF81FFE011CDFEF92D0AFEE7" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[696,823,296,320]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="coomani">
|
||
<emphasis id="B93FA736FF81FFE011CDFEF92D0AFEE7" box="[696,823,296,320]" italics="true" pageId="34" pageNumber="35">A. coomani</emphasis>
|
||
</taxonomicName>
|
||
) and unicolored maxillary palpomeres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35128AFFF81FFE013B2FE9D2C7AFAFF" pageId="34" pageNumber="35" type="description">
|
||
<paragraph id="8BF47B24FF81FFE013B2FE9D2B47FE2E" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FE9D2F6CFEC2" bold="true" box="[199,337,332,357]" pageId="34" pageNumber="35">Description</emphasis>
|
||
. Total length: 1.8̄
|
||
<quantity id="4CB3D6C1FF81FFE01153FE9C2CB8FEC3" box="[550,645,333,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="34" pageNumber="35" unit="mm" value="2.2">2.2 mm</quantity>
|
||
; elytral width: 0.8̄
|
||
<quantity id="4CB3D6C1FF81FFE01017FE9C2D83FEC3" box="[866,958,333,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="35" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; E.I.: 1.4, P.I.: 1.9̄2.2, elytra 3.0̄3.2× as long as pronotum. Habitus slender, elytra slightly widening posterior to midlength, moderately convex dorsally.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013B2FE452DBEFDC5" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FE452F77FE0A" box="[199,330,404,429]" italics="true" pageId="34" pageNumber="35">Coloration.</emphasis>
|
||
Labrum, clypeus and frons black; clypeus with distinct, delimited, triangular preocular patches, about as wide as eye or larger, in few cases of very dark colored individuals preocular patches smaller than eye; maxillary palpi yellow, palpomere 4 with infuscation apically, distinct in most cases, rarely indistinct, very rarely absent; pronotum yellow, unicolored, or with undefined dark brown to black central patch with variable extension, or unicolored dark brown to black with narrow yellow lateral rim; elytra yellow, unicolored or with variably sized undefined infuscations; ventrites dark brown or black; legs yellow.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013B2FDBC2C23FC9D" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FDBC2F39FD21" box="[199,260,621,646]" italics="true" pageId="34" pageNumber="35">Head</emphasis>
|
||
(Fig. 52). Clypeus with distinctly, evenly excised anterior margin; C.I.: 3.3, lateral length ratio clypeus/ eyes = 1.5̄1.6; microsculpture absent; ground punctures fine, interspaces 2̄3× as wide as punctures; systematic punctures moderately distinct. Eyes large, very slightly protruding. Antennae with nine antennomeres. Maxillary palpi slender, 1.1̄1.2× as long as pronotum in midline, 0.9̄1.0× as long as maximum width of clypeus; length ratio palpomere 4:3 = 1.2̄1.4, palpomere 4 slightly asymmetrical. Mentum with very fine, very widely spaced punctures, without microsculpture.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013B2FC972BA0FC00" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FC972F1DFCF8" box="[199,288,838,863]" italics="true" pageId="34" pageNumber="35">Thorax.</emphasis>
|
||
Pronotal ground punctation as on head; systematic punctures moderately distinct. Elytral ground punctation as on head and pronotum or slightly finer; systematic punctures indistinct, arranged in four rows, mesal rows 1̄3 with strongly reduced number of punctures, not reaching anterior margin. Mesoventrite with mesal bulge.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013B2FC652BA0FC48" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FC652EC1FC6C" box="[199,252,948,971]" italics="true" pageId="34" pageNumber="35">Legs</emphasis>
|
||
(Fig. 97). Pubescence present on proximal half of profemur or slightly more extended, on proximal 2/3 of meso- and metafemur; hairline oblique on profemur, slightly oblique on mesofemur, almost straight on metafemur.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013B2FC2A2A12FBB3" blockId="34.[151,1437,151,2018]" box="[199,1071,1019,1044]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FC2A2F0CFBB3" box="[199,305,1019,1044]" italics="true" pageId="34" pageNumber="35">Abdomen</emphasis>
|
||
. Ventrite 5 with shallow to almost semicircular apical emargination.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013B2FBCE2C7AFAFF" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FBCE2F09FB9F" box="[199,308,1055,1080]" italics="true" pageId="34" pageNumber="35">Aedeagus</emphasis>
|
||
(Fig. 139). Length:
|
||
<quantity id="4CB3D6C1FF81FFE01169FBF12CB9FB90" box="[540,644,1056,1080]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="34" pageNumber="35" unit="mm" value="0.33">0.33 mm</quantity>
|
||
. Phallobase shorter than parameres, about as long as wide, or slightly longer than wide, bending with obtuse angle towards narrowly rounded proximal apex; delimited manubrium absent; border between pigmented and unpigmented portion of ventral face indistinct, almost reaching proximal apex in midline. Parameres slender; margins weakly sigmoid, tapering towards blunt, asymmetrical apex, slightly inclining mesad; base slightly extending into phallobase, ventral face indistinct, slightly wider than dorsal face. Median lobe about as wide as one paramere, finger-shaped; apex bluntly rounded, not reaching apex of parameres; ventral face slightly wider and distinctly shorter than dorsal face; corona in subapical position; pair of styli present, arising from base of median lobe, not reaching its apex; basal apophyses moderately long, slightly inclining laterad, extending to distal third of phallobase.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35128AFFF81FFE013B2FAB52CA8F845" pageId="34" pageNumber="35" type="discussion">
|
||
<paragraph id="8BF47B24FF81FFE013B2FAB52D4CF942" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<emphasis id="B93FA736FF81FFE013B2FAB52F09FADA" bold="true" box="[199,308,1380,1405]" pageId="34" pageNumber="35">Remarks</emphasis>
|
||
. Specimens from
|
||
<collectingCountry id="F35C3BB4FF81FFE01288FAB42C7DFADB" box="[509,576,1381,1404]" name="Japan" pageId="34" pageNumber="35">Japan</collectingCountry>
|
||
, originally described as
|
||
<taxonomicName id="4C4B00A7FF81FFE01038FAB42DD2FADA" baseAuthorityName="Matsui" baseAuthorityYear="1994" box="[845,1007,1380,1405]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="ryukyuensis">
|
||
<emphasis id="B93FA736FF81FFE01038FAB42DD2FADA" box="[845,1007,1380,1405]" italics="true" pageId="34" pageNumber="35">A. ryukyuensis</emphasis>
|
||
</taxonomicName>
|
||
, do not differ from other specimens of
|
||
<taxonomicName id="4C4B00A7FF81FFE013E2FA582F21FA07" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[151,284,1417,1440]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="coomani">
|
||
<emphasis id="B93FA736FF81FFE013E2FA582F21FA07" box="[151,284,1417,1440]" italics="true" pageId="34" pageNumber="35">A. coomani</emphasis>
|
||
</taxonomicName>
|
||
in the external and male genital morphology that lies within the spectrum of variability of this widespread species.
|
||
<taxonomicName id="4C4B00A7FF81FFE0120EFA7D2C49FA63" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[379,628,1452,1477]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="coomani">
|
||
<emphasis id="B93FA736FF81FFE0120EFA7D2C49FA63" box="[379,628,1452,1477]" italics="true" pageId="34" pageNumber="35">Agraphydrus coomani</emphasis>
|
||
</taxonomicName>
|
||
varies in the pronotal coloration (between unicolored yellow and almost completely black) and slightly in the shape of the parameres (between more broadly and more narrowly rounded). The character states show a certain regional distribution pattern, but mixed populations occur: In
|
||
<collectingCountry id="F35C3BB4FF81FFE0167AFA242B6FF9AB" box="[1295,1362,1525,1548]" name="Japan" pageId="34" pageNumber="35">Japan</collectingCountry>
|
||
, some regions of
|
||
<collectingCountry id="F35C3BB4FF81FFE01264F9C92FBCF996" box="[273,385,1560,1585]" name="Indonesia" pageId="34" pageNumber="35">Indonesia</collectingCountry>
|
||
, the
|
||
<collectingCountry id="F35C3BB4FF81FFE012C2F9C92C05F996" box="[439,568,1560,1585]" name="Philippines" pageId="34" pageNumber="35">Philippines</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F35C3BB4FF81FFE01100F9C82D72F996" box="[629,847,1560,1585]" name="Papua New Guinea" pageId="34" pageNumber="35">
|
||
<collectingRegion id="498FB5C6FF81FFE01100F9C82C84F997" box="[629,697,1561,1584]" country="Indonesia" name="Papua Barat" pageId="34" pageNumber="35">Papua</collectingRegion>
|
||
New Guinea
|
||
</collectingCountry>
|
||
the pronotum of most individuals is almost entirely dark brown or black with narrow yellow lateral rim; in
|
||
<collectingCountry id="F35C3BB4FF81FFE01073F9ED2D71F9F2" box="[774,844,1596,1621]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE0102DF9ED2D90F9F2" box="[856,941,1596,1621]" name="Taiwan" pageId="34" pageNumber="35">Taiwan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE010CDF9EC2DCCF9F3" box="[952,1009,1597,1620]" name="Laos" pageId="34" pageNumber="35">Laos</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE01088F9ED2A5EF9F2" box="[1021,1123,1596,1621]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F35C3BB4FF81FFE017EAF9ED2AC3F9F2" box="[1183,1278,1596,1621]" name="Vietnam" pageId="34" pageNumber="35">Vietnam</collectingCountry>
|
||
the pronotum is mostly unicolored yellow or with small, indistinct central infuscation, very rarely the central patch has a wider extension. The parameres are more broadly rounded in many individuals from
|
||
<collectingCountry id="F35C3BB4FF81FFE01752F9552A50F93A" box="[1063,1133,1668,1693]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE0170EF9542A89F93B" box="[1147,1204,1669,1692]" name="Laos" pageId="34" pageNumber="35">Laos</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE017B7F9552B15F93A" box="[1218,1320,1668,1693]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE01643F9552BA5F93A" box="[1334,1432,1668,1693]" name="Vietnam" pageId="34" pageNumber="35">Vietnam</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE013E2F9782EE7F967" box="[151,218,1705,1728]" name="Japan" pageId="34" pageNumber="35">Japan</collectingCountry>
|
||
, some Indonesian islands,
|
||
<collectingCountry id="F35C3BB4FF81FFE01177F9792C56F966" box="[514,619,1704,1729]" name="Malaysia" pageId="34" pageNumber="35">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF81FFE01103F9792CDCF966" box="[630,737,1704,1729]" name="Australia" pageId="34" pageNumber="35">Australia</collectingCountry>
|
||
; they are more narrowly rounded in some islands of
|
||
<collectingCountry id="F35C3BB4FF81FFE0165DF9792BA5F966" box="[1320,1432,1704,1729]" name="Indonesia" pageId="34" pageNumber="35">Indonesia</collectingCountry>
|
||
, and mixed populations occur in New
|
||
<collectingCountry id="F35C3BB4FF81FFE0114EF91D2CB1F942" box="[571,652,1740,1765]" name="Guinea-Bissau" pageId="34" pageNumber="35">Guinea</collectingCountry>
|
||
and the
|
||
<collectingCountry id="F35C3BB4FF81FFE0119EF91D2D50F942" box="[747,877,1740,1765]" name="Philippines" pageId="34" pageNumber="35">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF47B24FF81FFE013B2F9202CA8F845" blockId="34.[151,1437,151,2018]" pageId="34" pageNumber="35">
|
||
<taxonomicName id="4C4B00A7FF81FFE013B2F9202F9AF8AE" box="[199,423,1777,1802]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">
|
||
<emphasis id="B93FA736FF81FFE013B2F9202F9AF8AE" box="[199,423,1777,1802]" italics="true" pageId="34" pageNumber="35">Agraphydrus comes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF81FFE012C0F9232C0AF8AE" box="[437,567,1777,1801]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="B93FA736FF81FFE012C0F9232C0AF8AE" box="[437,567,1777,1801]" italics="true" pageId="34" pageNumber="35">A. confusus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF81FFE01133F9232CFBF8AE" baseAuthorityName="Orchymont" baseAuthorityYear="1927" box="[582,710,1778,1801]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="coomani">
|
||
<emphasis id="B93FA736FF81FFE01133F9232CFBF8AE" box="[582,710,1778,1801]" italics="true" pageId="34" pageNumber="35">A. coomani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF81FFE011A0F9232D64F8AD" baseAuthorityName="Matsui" baseAuthorityYear="1994" box="[725,857,1777,1802]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="ishiharai">
|
||
<emphasis id="B93FA736FF81FFE011A0F9232D64F8AD" box="[725,857,1777,1802]" italics="true" pageId="34" pageNumber="35">A. ishiharai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4B00A7FF81FFE0101DF9232DD7F8AD" authorityName="Komarek & Hebauer" authorityYear="2018" box="[872,1002,1777,1802]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="jilanzhui">
|
||
<emphasis id="B93FA736FF81FFE0101DF9232DD7F8AD" box="[872,1002,1777,1802]" italics="true" pageId="34" pageNumber="35">A. jilanzhui</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4B00A7FF81FFE0175CF9232A95F8AD" authorityName="Komarek & Hebauer" authorityYear="2018" box="[1065,1192,1777,1802]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="B93FA736FF81FFE0175CF9232A95F8AD" box="[1065,1192,1777,1802]" italics="true" pageId="34" pageNumber="35">A. robustus</emphasis>
|
||
</taxonomicName>
|
||
form a group of very similar species, sharing the following features: habitus slender (E.I.: 1.4̄1.6; P.I.: 1.9̄2.0); elytra widening posterior to midlength (in contrast to all other Oriental species of
|
||
<taxonomicName id="4C4B00A7FF81FFE01001F8E82A3BF8F5" authorityName="Komarek & Hebauer" authorityYear="2018" box="[884,1030,1849,1874]" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FA736FF81FFE01001F8E82A3BF8F5" box="[884,1030,1849,1874]" italics="true" pageId="34" pageNumber="35">Agraphydrus</emphasis>
|
||
</taxonomicName>
|
||
); distinct preocular patches present; eyes large (lateral margin of clypeus less than 2.0× as long as lateral margin of eyes); maxillary palpi slender, palpomere 4 infuscated apically; C.I.: 2.9̄3.3; number of nine antennomeres; clypeal microsculpture absent; metafemoral pubescence present on proximal 2/3–3/4; apical emargination present on abdominal ventrite 5; aedeagus slender with undefined manubrium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35128AFFF80FFE113B2FF492DFFFF5F" pageId="35" pageNumber="36" type="biology_ecology">
|
||
<paragraph id="8BF47B24FF80FFE113B2FF492DFFFF5F" blockId="35.[151,1436,152,285]" pageId="35" pageNumber="36">
|
||
<emphasis id="B93FA736FF80FFE113B2FF492F19FF16" bold="true" box="[199,292,152,177]" pageId="35" pageNumber="36">Ecology</emphasis>
|
||
. Collected in
|
||
<collectingCountry id="F35C3BB4FF80FFE112B8FF492C2DFF16" box="[461,528,152,177]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
|
||
between sea level and
|
||
<quantity id="4CB3D6C1FF80FFE11052FF482D4CFF17" box="[807,881,153,177]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="m" value="800.0">800 m</quantity>
|
||
a.s.l. in rivers, including potamal sections, and streams, flowing through degraded forests, pastures, and cultivated land, on banks with shrubs, grass, sand, or mud; in various kinds of pools with aquatic vegetation; also in brackish water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35128AFFF80FFE113B2FED52B59FEBA" box="[199,1380,260,285]" pageId="35" pageNumber="36" type="distribution">
|
||
<paragraph id="8BF47B24FF80FFE113B2FED52B59FEBA" blockId="35.[151,1436,152,285]" box="[199,1380,260,285]" pageId="35" pageNumber="36">
|
||
<emphasis id="B93FA736FF80FFE113B2FED52F64FEBA" bold="true" box="[199,345,260,285]" pageId="35" pageNumber="36">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F35C3BB4FF80FFE11212FED72F97FEBA" box="[359,426,262,285]" name="Japan" pageId="35" pageNumber="36">Japan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C3BB4FF80FFE112C0FED52FC5FEBA" box="[437,504,260,285]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="498FB5C6FF80FFE11173FED52C6FFEBA" box="[518,594,260,285]" country="China" name="Fujian" pageId="35" pageNumber="36">Fujian</collectingRegion>
|
||
,
|
||
<collectingRegion id="498FB5C6FF80FFE11128FED52CDFFEBA" box="[605,738,260,285]" country="China" name="Guangdong" pageId="35" pageNumber="36">Guangdong</collectingRegion>
|
||
,
|
||
<collectingRegion id="498FB5C6FF80FFE1119BFED52D7CFEBA" box="[750,833,260,285]" country="China" name="Hainan" pageId="35" pageNumber="36">Hainan</collectingRegion>
|
||
) and
|
||
<collectingCountry id="F35C3BB4FF80FFE1100AFED52DECFEBA" box="[895,977,260,285]" name="Taiwan" pageId="35" pageNumber="36">Taiwan</collectingCountry>
|
||
to
|
||
<collectingCountry id="F35C3BB4FF80FFE11080FED52A62FEBA" box="[1013,1119,260,285]" name="Australia" pageId="35" pageNumber="36">Australia</collectingCountry>
|
||
. First record for
|
||
<collectingCountry id="F35C3BB4FF80FFE1166FFED52B5CFEBA" box="[1306,1377,260,285]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |