242 lines
30 KiB
XML
242 lines
30 KiB
XML
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<mods:title id="F6B91888B967195D732F59F60AC38A39">First record and a new species of the fossil dragonfly genus Proinogomphus (Odonata: Liassogomphidae) from the Early Jurassic of Bascharage in the Grand Duchy of Luxembourg</mods:title>
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<mods:namePart id="C8B7EB647593010B25A62E28402C4FD1">Bechly, Günter</mods:namePart>
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<mods:title id="FBF209AF46C0CFF54BE4839402D61FAD">Zootaxa</mods:title>
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<mods:date id="10523C1BFA1529E9771B49534196EA7B">2018</mods:date>
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<treatment id="03E2C3245E27FFC211C8F9B24DABBEBA" ID-DOI="http://doi.org/10.5281/zenodo.5946719" ID-GBIF-Taxon="148462278" ID-Zenodo-Dep="5946719" LSID="urn:lsid:plazi:treatment:03E2C3245E27FFC211C8F9B24DABBEBA" httpUri="http://treatment.plazi.org/id/03E2C3245E27FFC211C8F9B24DABBEBA" lastPageId="1" lastPageNumber="109" pageId="0" pageNumber="108">
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<subSubSection id="C35121B95E27FFC311C8F9B248E4BA75" box="[151,471,1614,1639]" pageId="0" pageNumber="108" type="nomenclature">
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<paragraph id="8BF472325E27FFC311C8F9B248E4BA75" blockId="0.[151,1436,1614,1855]" box="[151,471,1614,1639]" pageId="0" pageNumber="108">
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<heading id="D0BCC55E5E27FFC311C8F9B248E4BA75" bold="true" box="[151,471,1614,1639]" fontSize="10" level="2" pageId="0" pageNumber="108" reason="2">
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<emphasis id="B93FAE205E27FFC311C8F9B248E4BA75" bold="true" box="[151,471,1614,1639]" pageId="0" pageNumber="108">
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<taxonomicName id="4C4B09B15E27FFC311C8F9B248E1BA75" authority="Bode, 1953" authorityName="Bode" authorityYear="1953" box="[151,466,1614,1639]" class="Insecta" family="Liassogomphidae" genus="Elattogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC311C8F9B24874BA75" bold="true" box="[151,327,1614,1639]" italics="true" pageId="0" pageNumber="108">Elattogomphus</emphasis>
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<bibRefCitation id="EFDA0FC35E27FFC31012F9B248E1BA75" box="[333,466,1614,1639]" pageId="0" pageNumber="108" refId="ref2697" refString="Bode, A. (1953) Die Insektenfauna des Ostniedersachsischen Oberen Lias. Palaeontographica, Abteilung A, 103, 1 - 375, 15 pls." type="journal article">Bode, 1953</bibRefCitation>
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</taxonomicName>
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.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35121B95E27FFC311C8F98F4AD4BABD" pageId="0" pageNumber="108" type="reference_group">
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<paragraph id="8BF472325E27FFC311C8F98F4AD4BABD" blockId="0.[151,1436,1614,1855]" pageId="0" pageNumber="108">
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<taxonomicName id="4C4B09B15E27FFC311C8F98F48C8BA99" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[151,507,1650,1675]" class="Insecta" family="Liassogomphidae" genus="Heterothemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC311C8F98F481EBA99" box="[151,301,1651,1675]" italics="true" pageId="0" pageNumber="108">Heterothemis</emphasis>
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Handlirsch, 1906
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</taxonomicName>
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(=
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<taxonomicName id="4C4B09B15E27FFC31377F98F4A41BA99" authority="Handlirsch, 1920" authorityName="Handlirsch" authorityYear="1920" box="[552,882,1650,1675]" class="Insecta" family="Liassogomphidae" genus="Gomphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC31377F98F4B92BA99" box="[552,673,1651,1675]" italics="true" pageId="0" pageNumber="108">Gomphites</emphasis>
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Handlirsch, 1920
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</taxonomicName>
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,
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<taxonomicName id="4C4B09B15E27FFC312DFF98F4DEABA99" authority="Cowley, 1934" authorityName="Cowley" authorityYear="1934" box="[896,1241,1650,1675]" class="Insecta" family="Liassogomphidae" genus="Liassogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC312DFF98F4D1DBA99" box="[896,1070,1651,1675]" italics="true" pageId="0" pageNumber="108">Liassogomphus</emphasis>
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Cowley, 1934
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</taxonomicName>
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,
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<taxonomicName id="4C4B09B15E27FFC315B8F98F486ABABD" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" class="Insecta" family="Liassogomphidae" genus="Palaeogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC315B8F98F4CAFBA99" box="[1255,1436,1651,1675]" italics="true" pageId="0" pageNumber="108">Palaeogomphus</emphasis>
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<bibRefCitation id="EFDA0FC35E27FFC311C8F96A486ABABD" box="[151,345,1686,1711]" pageId="0" pageNumber="108" refId="ref3083" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten. II. Teil. Annalen des Naturhistorischen Museums in Wien, 49 (1938), 1 - 240, 16 pls." type="journal article">Handlirsch, 1939</bibRefCitation>
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</taxonomicName>
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) with the single type species
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<taxonomicName id="4C4B09B15E27FFC313FCF96B4AD7BABD" authority="(Buckman, 1843)" authorityName="Buckman" authorityYear="1843" baseAuthorityName="Buckman" baseAuthorityYear="1843" box="[675,996,1686,1711]" class="Insecta" family="Liassogomphidae" genus="Heterothemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="species" species="brodiei">
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<emphasis id="B93FAE205E27FFC313FCF96B4A26BABC" box="[675,789,1686,1710]" italics="true" pageId="0" pageNumber="108">H. brodiei</emphasis>
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(Buckman, 1843)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C35121B95E27FFC311C8F94749DABAE5" pageId="0" pageNumber="108" type="discussion">
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<paragraph id="8BF472325E27FFC311C8F94749DABAE5" blockId="0.[151,1436,1614,1855]" pageId="0" pageNumber="108">
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Comment: The synonymy of
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<taxonomicName id="4C4B09B15E27FFC310AEF9474BACBAC1" authorityName="Cowley" authorityYear="1934" box="[497,671,1723,1747]" class="Insecta" family="Liassogomphidae" genus="Liassogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC310AEF9474BACBAC1" box="[497,671,1723,1747]" italics="true" pageId="0" pageNumber="108">Liassogomphus</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="4C4B09B15E27FFC313B8F9474D7EBAC1" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[743,1101,1722,1747]" class="Insecta" family="Liassogomphidae" genus="Heterothemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC313B8F9474A4EBAC1" box="[743,893,1723,1747]" italics="true" pageId="0" pageNumber="108">Heterothemis</emphasis>
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Handlirsch, 1906
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</taxonomicName>
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was established by
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<bibRefCitation id="EFDA0FC35E27FFC31461F94749D6BAE5" pageId="0" pageNumber="108" refId="ref2534" refString="Ansorge, A. (2004) Insekten aus Liasgeoden der Ahrensburger Geschiebesippe - mit einem Ausblick auf lokale Anreicherungen von Liasgeoden in Mecklenburg-Vorpommern. Archiv fur Geschiebekunde, 3, 779 - 784." type="journal article">Ansorge (2004)</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35121B95E27FFC211C8F8FF4851BD0F" lastPageId="1" lastPageNumber="109" pageId="0" pageNumber="108" type="reference_group">
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||
<paragraph id="8BF472325E27FFC311C8F8FF4ABDBB09" blockId="0.[151,1436,1614,1855]" box="[151,910,1794,1819]" pageId="0" pageNumber="108">
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<taxonomicName id="4C4B09B15E27FFC311C8F8FF48C2BB09" authority="Cowley, 1942" authorityName="Cowley" authorityYear="1942" box="[151,497,1794,1819]" class="Insecta" family="Liassogomphidae" genus="Phthitogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC311C8F8FF487CBB09" box="[151,335,1795,1819]" italics="true" pageId="0" pageNumber="108">Phthitogomphus</emphasis>
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<bibRefCitation id="EFDA0FC35E27FFC3100AF8FE48C2BB09" box="[341,497,1794,1819]" pageId="0" pageNumber="108" refId="ref2789" refString="Cowley, J. (1942) Descriptions of some genera of fossil Odonata. Proceedings of the Royal Entomological Society of London (B), 11 (5), 63 - 78. https: // doi. org / 10.1111 / j. 1365 - 3113.1942. tb 00725. x" type="journal article">Cowley, 1942</bibRefCitation>
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</taxonomicName>
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(=
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<taxonomicName id="4C4B09B15E27FFC31347F8FF4AB1BB09" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[536,898,1794,1819]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC31347F8FF4B86BB09" box="[536,693,1795,1819]" italics="true" pageId="0" pageNumber="108">Paragomphus</emphasis>
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<bibRefCitation id="EFDA0FC35E27FFC313E3F8FE4AB1BB09" box="[700,898,1794,1819]" pageId="0" pageNumber="108" refId="ref3083" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten. II. Teil. Annalen des Naturhistorischen Museums in Wien, 49 (1938), 1 - 240, 16 pls." type="journal article">Handlirsch, 1939</bibRefCitation>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BF472325E27FFC311C8F8DA4D76BB2D" blockId="0.[151,1436,1614,1855]" box="[151,1093,1830,1855]" pageId="0" pageNumber="108">
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<taxonomicName id="4C4B09B15E27FFC311C8F8DA48D8BB2D" authority="Cowley, 1942" authorityName="Cowley" authorityYear="1942" box="[151,491,1830,1855]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FAE205E27FFC311C8F8DA487ABB2C" box="[151,329,1830,1854]" italics="true" pageId="0" pageNumber="108">Proinogomphus</emphasis>
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||
<bibRefCitation id="EFDA0FC35E27FFC3100FF8DA48D8BB2D" box="[336,491,1830,1855]" pageId="0" pageNumber="108" refId="ref2789" refString="Cowley, J. (1942) Descriptions of some genera of fossil Odonata. Proceedings of the Royal Entomological Society of London (B), 11 (5), 63 - 78. https: // doi. org / 10.1111 / j. 1365 - 3113.1942. tb 00725. x" type="journal article">Cowley, 1942</bibRefCitation>
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||
</taxonomicName>
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||
(=
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<taxonomicName id="4C4B09B15E27FFC3134DF8DA4ABFBB2D" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[530,908,1830,1855]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B93FAE205E27FFC3134DF8DA4BF7BB2C" box="[530,708,1830,1854]" italics="true" pageId="0" pageNumber="108">Proinogomphus</emphasis>
|
||
<bibRefCitation id="EFDA0FC35E27FFC31394F8DA4ABFBB2D" box="[715,908,1830,1855]" pageId="0" pageNumber="108" refId="ref3083" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten. II. Teil. Annalen des Naturhistorischen Museums in Wien, 49 (1938), 1 - 240, 16 pls." type="journal article">Handlirsch, 1939</bibRefCitation>
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||
</taxonomicName>
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||
numen nudum).
|
||
</paragraph>
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||
<paragraph id="8BF472325E27FFC21198F8934851BD0F" blockId="0.[151,1436,1902,2035]" lastBlockId="1.[151,1437,151,680]" lastPageId="1" lastPageNumber="109" pageId="0" pageNumber="108">
|
||
A key was provided by
|
||
<bibRefCitation id="EFDA0FC35E27FFC3108DF8924B4EBB95" box="[466,637,1902,1927]" pageId="0" pageNumber="108" refId="ref2789" refString="Cowley, J. (1942) Descriptions of some genera of fossil Odonata. Proceedings of the Royal Entomological Society of London (B), 11 (5), 63 - 78. https: // doi. org / 10.1111 / j. 1365 - 3113.1942. tb 00725. x" type="journal article">Cowley (1942)</bibRefCitation>
|
||
, and the family was first revised by
|
||
<bibRefCitation id="EFDA0FC35E27FFC3154CF8924DFEBB95" box="[1043,1229,1902,1927]" pageId="0" pageNumber="108" refId="ref3420" refString="Nel, A., Martinez-Delclos, X., Paichler, J. C. & Henrotay, M. (1993) Les ' Anisozygptera' fossiles. Phylogenie et classification (Odonata). Martinia, Numero Hors Serie 3, 1 - 311." type="journal article">
|
||
Nel
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||
<emphasis id="B93FAE205E27FFC3151DF8934D49BB94" box="[1090,1146,1902,1926]" italics="true" pageId="0" pageNumber="108">et al.</emphasis>
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||
(1993)
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||
</bibRefCitation>
|
||
, who removed the genera
|
||
<taxonomicName id="4C4B09B15E27FFC311B6F86F4B71BBB9" authority="Bode, 1953" authorityName="Bode" authorityYear="1953" box="[233,578,1938,1963]" class="Insecta" family="Myopophlebiidae" genus="Strongylogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B93FAE205E27FFC311B6F86F488ABBB9" box="[233,441,1939,1963]" italics="true" pageId="0" pageNumber="108">Strongylogomphus</emphasis>
|
||
<bibRefCitation id="EFDA0FC35E27FFC3109EF86E4B71BBB9" box="[449,578,1938,1963]" pageId="0" pageNumber="108" refId="ref2697" refString="Bode, A. (1953) Die Insektenfauna des Ostniedersachsischen Oberen Lias. Palaeontographica, Abteilung A, 103, 1 - 375, 15 pls." type="journal article">Bode, 1953</bibRefCitation>
|
||
</taxonomicName>
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||
and
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||
<taxonomicName id="4C4B09B15E27FFC31325F86F4A88BBB9" authority="Cowley, 1942" authorityName="Cowley" authorityYear="1942" box="[634,955,1938,1963]" class="Insecta" family="Liassogomphidae" genus="Necygomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B93FAE205E27FFC31325F86F4A24BBB9" box="[634,791,1939,1963]" italics="true" pageId="0" pageNumber="108">Necygomphus</emphasis>
|
||
<bibRefCitation id="EFDA0FC35E27FFC31240F86E4A88BBB9" box="[799,955,1938,1963]" pageId="0" pageNumber="108" refId="ref2789" refString="Cowley, J. (1942) Descriptions of some genera of fossil Odonata. Proceedings of the Royal Entomological Society of London (B), 11 (5), 63 - 78. https: // doi. org / 10.1111 / j. 1365 - 3113.1942. tb 00725. x" type="journal article">Cowley, 1942</bibRefCitation>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C4B09B15E27FFC312BCF86F4C65BBB9" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[995,1366,1938,1963]" class="Insecta" family="Liassogomphidae" genus="Necrogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FAE205E27FFC312BCF86F4DBFBBB9" box="[995,1164,1939,1963]" italics="true" pageId="0" pageNumber="108">Necrogomphus</emphasis>
|
||
<bibRefCitation id="EFDA0FC35E27FFC315CCF86E4C65BBB9" box="[1171,1366,1938,1963]" pageId="0" pageNumber="108" refId="ref3083" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten. II. Teil. Annalen des Naturhistorischen Museums in Wien, 49 (1938), 1 - 240, 16 pls." type="journal article">Handlirsch, 1939</bibRefCitation>
|
||
</taxonomicName>
|
||
) from
|
||
<taxonomicName id="4C4B09B15E27FFC311C8F84A486FBBDD" authorityName="Tillyard" authorityYear="1935" box="[151,348,1974,1999]" class="Insecta" family="Liassogomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="family">Liassogomphidae</taxonomicName>
|
||
and transferred them to
|
||
<taxonomicName id="4C4B09B15E27FFC3132CF84A4A0BBBDD" authorityName="Bode" authorityYear="1953" box="[627,824,1974,1999]" class="Insecta" family="Myopophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="108" phylum="Arthropoda" rank="family">Myopophlebiidae</taxonomicName>
|
||
within Heterophlebioidea. A second revision by
|
||
<bibRefCitation id="EFDA0FC35E27FFC31439F84B4879BBE1" pageId="0" pageNumber="108" refId="ref2956" refString="Etter, W. & Kuhn, O. (2000) An articulated dragonfly (Insecta, Odonata) from the Upper Liassic Posidonia Shale of northern Switzerland. Palaeontology, 43 (5), 967 - 977. https: // doi. org / 10.1111 / 1475 - 4983.00157" type="journal article">Etter & Kuhn (2000)</bibRefCitation>
|
||
retained these remaining four genera, but reduced the number of species from 9 to 4 based on assumed wing venational variability similar to extant
|
||
<taxonomicName id="4C4B09B15E26FFC213A7FF6B4A41BCA2" ID-CoL="63Y" box="[760,882,151,176]" class="Insecta" family="Aeshnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
|
||
. An additional genus
|
||
<taxonomicName id="4C4B09B15E26FFC21536FF644CAFBCA2" authority="Ren, 1994" authorityName="Ren" authorityYear="1994" box="[1129,1436,152,176]" class="Insecta" family="Liassogomphidae" genus="Chrysogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FAE205E26FFC21536FF644C2CBCA2" box="[1129,1311,152,176]" italics="true" pageId="1" pageNumber="109">Chrysogomphus</emphasis>
|
||
<bibRefCitation id="EFDA0FC35E26FFC21477FF644CAFBCA2" box="[1320,1436,152,176]" pageId="1" pageNumber="109" refId="ref3467" refString="Ren, D. (1994) A new genus and species of Liassogomphidae (Insecta, Odonata) from Late Jurassic of Liaoning province. Geoscience, 8 (3), 254 - 258." type="journal article">Ren, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
was described from the Early Cretaceous of China, but later identified as a basal Aeshnoptera (
|
||
<bibRefCitation id="EFDA0FC35E26FFC2159EFF404CA2BCC6" box="[1217,1425,188,212]" pageId="1" pageNumber="109" refId="ref3314" refString="Huang, D. - Y., Nel, A. & Lin, Q. - B. (2003) A new genus and species of aeshnopteran dragonfly from the Lower Cretaceous of China. Cretaceous Research, 24, 141 - 147. https: // doi. org / 10.1016 / S 0195 - 6671 (03) 00027 - 2" type="journal article">
|
||
Huang
|
||
<emphasis id="B93FAE205E26FFC2144BFF414C7EBCC6" box="[1300,1357,188,212]" italics="true" pageId="1" pageNumber="109">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
). According to the cladistic analysis by
|
||
<bibRefCitation id="EFDA0FC35E26FFC21310FF234A15BCEA" box="[591,806,223,248]" pageId="1" pageNumber="109" refId="ref3009" refString="Fleck, G., Bechly, G., Martinez-Delclos, X., Jarzembowski, E., Coram, R. & Nel, A. (2003) Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Societe Entomologique de France, New Series, 39 (1), 55 - 93. https: // doi. org / 10.1080 / 00379271.2003.10697363" type="journal article">
|
||
Fleck
|
||
<emphasis id="B93FAE205E26FFC213CAFF1D4BE3BCEA" box="[661,720,224,248]" italics="true" pageId="1" pageNumber="109">et al.</emphasis>
|
||
(2003)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C4B09B15E26FFC2126CFF234ACBBCEA" authorityName="Tillyard" authorityYear="1935" box="[819,1016,223,248]" class="Insecta" family="Liassogomphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="family">Liassogomphidae</taxonomicName>
|
||
represents the sister group of crown group Anisoptera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35121B95E26FFC21198FEDB4DABBEBA" pageId="1" pageNumber="109" type="description">
|
||
<paragraph id="8BF472325E26FFC21198FEDB4DABBEBA" blockId="1.[151,1437,151,680]" pageId="1" pageNumber="109">
|
||
The present study describes a new species of the liassogomphid genus
|
||
<taxonomicName id="4C4B09B15E26FFC21558FED44D8ABD52" authorityName="Handlirsch" authorityYear="1939" box="[1031,1209,296,320]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FAE205E26FFC21558FED44D8ABD52" box="[1031,1209,296,320]" italics="true" pageId="1" pageNumber="109">Proinogomphus</emphasis>
|
||
</taxonomicName>
|
||
. Its
|
||
<typeStatus id="54F0CC905E26FFC215AEFED44C12BD52" box="[1265,1313,296,320]" pageId="1" pageNumber="109">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C4B09B15E26FFC214D8FED549ECBD76" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="species" species="brodei">
|
||
<emphasis id="B93FAE205E26FFC214D8FED549ECBD76" italics="true" pageId="1" pageNumber="109">P. brodei</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EFDA0FC35E26FFC210E8FEB04BBBBD76" box="[439,648,332,357]" pageId="1" pageNumber="109" refId="ref3083" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten. II. Teil. Annalen des Naturhistorischen Museums in Wien, 49 (1938), 1 - 240, 16 pls." type="journal article">Handlirsch (1939)</bibRefCitation>
|
||
, based on a figure of
|
||
<bibRefCitation id="EFDA0FC35E26FFC212D9FEB04D24BD76" box="[902,1047,332,357]" pageId="1" pageNumber="109" refId="ref2662" refString="Bode, A. (1907) Orthoptera und Neuroptera aus dem oberen Lias von Braunschweig. Jahrbuch der Preussischen geologischen Landesanstalt, 25, 218 - 245, pls. 6 - 7." type="journal article">Bode (1907)</bibRefCitation>
|
||
, but his introduction of the genus
|
||
<taxonomicName id="4C4B09B15E26FFC211C8FE8C487ABD9A" authorityName="Handlirsch" authorityYear="1939" box="[151,329,368,392]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FAE205E26FFC211C8FE8C487ABD9A" box="[151,329,368,392]" italics="true" pageId="1" pageNumber="109">Proinogomphus</emphasis>
|
||
</taxonomicName>
|
||
was considered a nomen nudum by
|
||
<bibRefCitation id="EFDA0FC35E26FFC213ABFE934AACBD9A" box="[756,927,367,392]" pageId="1" pageNumber="109" refId="ref2789" refString="Cowley, J. (1942) Descriptions of some genera of fossil Odonata. Proceedings of the Royal Entomological Society of London (B), 11 (5), 63 - 78. https: // doi. org / 10.1111 / j. 1365 - 3113.1942. tb 00725. x" type="journal article">Cowley (1942)</bibRefCitation>
|
||
because “Handlirsch has not complied with Art. 25c 1–3 of the International Code, so that Cowley redescribed this genus. Cowley’s view was ignored by
|
||
<bibRefCitation id="EFDA0FC35E26FFC211C8FE444868BDC2" box="[151,347,440,464]" pageId="1" pageNumber="109" refId="ref2725" refString="Carpenter F. M. (1992) Superclass Hexapoda. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vol. 3. The Geological Society of America, Boulder, CO and The University of Kansas, Lawrence, KS, pp. 80 - 81." type="book chapter">Carpenter (1992)</bibRefCitation>
|
||
and the revision of
|
||
<bibRefCitation id="EFDA0FC35E26FFC2130FFE4B4A26BDC2" box="[592,789,439,464]" pageId="1" pageNumber="109" refId="ref3420" refString="Nel, A., Martinez-Delclos, X., Paichler, J. C. & Henrotay, M. (1993) Les ' Anisozygptera' fossiles. Phylogenie et classification (Odonata). Martinia, Numero Hors Serie 3, 1 - 311." type="journal article">
|
||
Nel
|
||
<emphasis id="B93FAE205E26FFC213DCFE454BF3BDC2" box="[643,704,440,464]" italics="true" pageId="1" pageNumber="109">et al.</emphasis>
|
||
(1993)
|
||
</bibRefCitation>
|
||
(adopted in the Paleobiology Database PaleoDB taxon number
|
||
<date id="FFF554F25E26FFC211AAFE20487EBDE7" box="[245,333,476,501]" pageId="1" pageNumber="109" value="39-06-18">180639</date>
|
||
), but accepted in the latest revision by
|
||
<bibRefCitation id="EFDA0FC35E26FFC2124EFE214D37BDE6" box="[785,1028,476,501]" pageId="1" pageNumber="109" refId="ref2956" refString="Etter, W. & Kuhn, O. (2000) An articulated dragonfly (Insecta, Odonata) from the Upper Liassic Posidonia Shale of northern Switzerland. Palaeontology, 43 (5), 967 - 977. https: // doi. org / 10.1111 / 1475 - 4983.00157" type="journal article">Etter & Kuhn (2000)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EFDA0FC35E26FFC2154EFE204DACBDE7" box="[1041,1183,476,501]" pageId="1" pageNumber="109" refId="ref2697" refString="Bode, A. (1953) Die Insektenfauna des Ostniedersachsischen Oberen Lias. Palaeontographica, Abteilung A, 103, 1 - 375, 15 pls." type="journal article">Bode (1953)</bibRefCitation>
|
||
described two further species
|
||
<taxonomicName id="4C4B09B15E26FFC211B0FDFD4884BE0A" authorityName="Bode" authorityYear="1953" box="[239,439,512,536]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="species" species="aequalicellatus">
|
||
<emphasis id="B93FAE205E26FFC211B0FDFD4884BE0A" box="[239,439,512,536]" italics="true" pageId="1" pageNumber="109">P. aequalicellatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B09B15E26FFC210B0FDFD4B4BBE0A" authorityName="Bode" authorityYear="1953" box="[495,632,512,536]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="species" species="bicellatus">
|
||
<emphasis id="B93FAE205E26FFC210B0FDFD4B4BBE0A" box="[495,632,512,536]" italics="true" pageId="1" pageNumber="109">P. bicellatus</emphasis>
|
||
</taxonomicName>
|
||
, which have been synonymized with the
|
||
<typeStatus id="54F0CC905E26FFC21511FDFC4D4DBE0A" box="[1102,1150,512,536]" pageId="1" pageNumber="109">type</typeStatus>
|
||
species by
|
||
<bibRefCitation id="EFDA0FC35E26FFC2145DFDFC49D7BE2E" pageId="1" pageNumber="109" refId="ref2956" refString="Etter, W. & Kuhn, O. (2000) An articulated dragonfly (Insecta, Odonata) from the Upper Liassic Posidonia Shale of northern Switzerland. Palaeontology, 43 (5), 967 - 977. https: // doi. org / 10.1111 / 1475 - 4983.00157" type="journal article">Etter & Kuhn (2000)</bibRefCitation>
|
||
, because they shall not differ in any major character from the
|
||
<typeStatus id="54F0CC905E26FFC212C5FDD94AF9BE2F" box="[922,970,549,573]" pageId="1" pageNumber="109">type</typeStatus>
|
||
species, which appears to be at least a bit dubious concerning
|
||
<taxonomicName id="4C4B09B15E26FFC21022FDB54B75BE72" authorityName="Bode" authorityYear="1953" box="[381,582,584,608]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="species" species="aequalicellatus">
|
||
<emphasis id="B93FAE205E26FFC21022FDB54B75BE72" box="[381,582,584,608]" italics="true" pageId="1" pageNumber="109">P. aequalicellatus</emphasis>
|
||
</taxonomicName>
|
||
because this species differs from
|
||
<taxonomicName id="4C4B09B15E26FFC21293FDB54D01BE72" box="[972,1074,584,608]" class="Insecta" family="Liassogomphidae" genus="Proinogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="109" phylum="Arthropoda" rank="species" species="brodei">
|
||
<emphasis id="B93FAE205E26FFC21293FDB54D01BE72" box="[972,1074,584,608]" italics="true" pageId="1" pageNumber="109">P. brodei</emphasis>
|
||
</taxonomicName>
|
||
in the fact that the triangle and subtriangle seem to be free in as in the new species described here. Nevertheless, I herein tentatively retain this synonymy, as there are sufficient other diagnostic characters to distinguish the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |