167 lines
19 KiB
XML
167 lines
19 KiB
XML
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<mods:titleInfo id="6BC29639D75D7EB10460A3D79C026AB9">
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<mods:title id="89FD5F7585C3AC997F2FF25929266507">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:name id="DE27BBF8FA3545CB2B7B9A2225FDB426" type="personal">
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<mods:namePart id="432B157C8163593AD02D0DDEAFAE0006">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="30F69CF248B826A2D5D7C2495CA8BE22" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8BDCAFA9961C5053574166392EBB2601">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="5842EBF9DAC4BDF4AA7A8A265D01C75E" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FB83A3FD8809EE692A633B31AE7448D1">Edgecombe, Gregory D.</mods:namePart>
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<mods:nameIdentifier id="17D5AA249807645DF1DDAD473A66B222" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:title id="874435790270CB9846A6D0BD0DFE83D5">Zootaxa</mods:title>
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<mods:part id="2662003D4074C0F39656BAC25A2532CA">
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<mods:date id="1FD2F29BD23BFC92DDC35260478B911B">2020</mods:date>
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<mods:number id="4B14982AA8027ABCDEFD40B65E2A01C1">2020-08-10</mods:number>
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<mods:number id="3D489D052A7E55506FD3D79BB7266699">4825</mods:number>
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<treatment id="03DE092DFFD5D722FF13F8E82974DE9B" ID-DOI="http://doi.org/10.5281/zenodo.4402206" ID-GBIF-Taxon="177020737" ID-Zenodo-Dep="4402206" LSID="urn:lsid:plazi:treatment:03DE092DFFD5D722FF13F8E82974DE9B" httpUri="http://treatment.plazi.org/id/03DE092DFFD5D722FF13F8E82974DE9B" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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<subSubSection id="C36DEBB0FFD5D72DFF13F8E82CA6DB3C" box="[151,483,1801,1828]" pageId="39" pageNumber="40" type="nomenclature">
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<paragraph id="8BC8B83BFFD5D72DFF13F8E82CA6DB3C" blockId="39.[151,483,1801,1828]" box="[151,483,1801,1828]" pageId="39" pageNumber="40">
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<heading id="D0800F57FFD5D72DFF13F8E82CA6DB3C" bold="true" box="[151,483,1801,1828]" fontSize="11" level="2" pageId="39" pageNumber="40" reason="4">
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<taxonomicName id="4C77C3B8FFD5D72DFF13F8E82CA6DB3C" ID-CoL="33KD" authority="Wurmli, 1972" authorityName="Wurmli" authorityYear="1972" box="[151,483,1801,1828]" class="Chilopoda" family="Scolopendridae" genus="Asanadopsis" kingdom="Animalia" order="Scolopendromorpha" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFD5D72DFF13F8E82CA6DB3C" bold="true" box="[151,483,1801,1828]" pageId="39" pageNumber="40">
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<emphasis id="B9036429FFD5D72DFF13F8E82C6ADB3B" bold="true" box="[151,303,1801,1828]" italics="true" pageId="39" pageNumber="40">Asanadopsis</emphasis>
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<bibRefCitation id="EFE6C5CAFFD5D72DFEB2F8E82CA6DB3C" author="Wurmli, M." box="[310,483,1801,1827]" pageId="39" pageNumber="40" pagination="88 - 104" refId="ref51745" refString="Wurmli, M. (1972) Chilopoda von Sumba und Flores I. Scolopendromorpha. Verhandlungen der Naturforschenden Gesellschaft in Basel, 82 (1), 88 - 104." type="journal article" year="1972">Würmli, 1972</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFD5D72DFF13F8AF2EDFDB79" box="[151,922,1870,1894]" pageId="39" pageNumber="40" type="type_taxon">
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<paragraph id="8BC8B83BFFD5D72DFF13F8AF2EDFDB79" blockId="39.[151,922,1870,1894]" box="[151,922,1870,1894]" pageId="39" pageNumber="40">
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<emphasis id="B9036429FFD5D72DFF13F8AF2C66DB79" bold="true" box="[151,291,1870,1894]" pageId="39" pageNumber="40">
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<typeStatus id="54CC0699FFD5D72DFF13F8AF2D89DB79" box="[151,204,1870,1894]" pageId="39" pageNumber="40">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C77C3B8FFD5D72DFEAEF8AF2FD7DB79" ID-CoL="5VRVN" authority="Wurmli, 1972" authorityName="Wurmli" authorityYear="1972" box="[298,658,1870,1894]" class="Chilopoda" family="Scolopendridae" genus="Asanadopsis" kingdom="Animalia" order="Scolopendromorpha" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="nueschi">
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<emphasis id="B9036429FFD5D72DFEAEF8AF2CB8DB79" box="[298,509,1870,1894]" italics="true" pageId="39" pageNumber="40">Asanadopsis nueschi</emphasis>
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<bibRefCitation id="EFE6C5CAFFD5D72DFD87F8AF2FD7DB79" author="Wurmli, M." box="[515,658,1870,1894]" pageId="39" pageNumber="40" pagination="88 - 104" refId="ref51745" refString="Wurmli, M. (1972) Chilopoda von Sumba und Flores I. Scolopendromorpha. Verhandlungen der Naturforschenden Gesellschaft in Basel, 82 (1), 88 - 104." type="journal article" year="1972">Würmli, 1972</bibRefCitation>
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</taxonomicName>
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(by original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFD5D722FF13F8752F71DCCB" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="diagnosis">
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<paragraph id="8BC8B83BFFD5D72DFF13F8752CB1DBE9" blockId="39.[151,1436,1940,2038]" pageId="39" pageNumber="40">
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<emphasis id="B9036429FFD5D72DFF13F8752C57DBB1" bold="true" box="[151,274,1940,1966]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
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Antennae very short, not reaching the anterior margin of tergite 1 when reflexed, cephalic plate with very short and wide apart paired sutures in the middle (fig. 5 of
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<bibRefCitation id="EFE6C5CAFFD5D72DFCE9F859294DDBCD" author="Wurmli, M." box="[877,1032,1976,2002]" pageId="39" pageNumber="40" pagination="88 - 104" refId="ref51745" refString="Wurmli, M. (1972) Chilopoda von Sumba und Flores I. Scolopendromorpha. Verhandlungen der Naturforschenden Gesellschaft in Basel, 82 (1), 88 - 104." type="journal article" year="1972">Würmli 1972</bibRefCitation>
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). Legs without tarsal spur. Ultimate sternite without median sulcus.
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</paragraph>
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<paragraph id="8BC8B83BFFDAD722FF43FF772C80DCAF" blockId="40.[151,1437,150,645]" box="[199,453,150,176]" pageId="40" pageNumber="41">
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<emphasis id="B9036429FFDAD722FF43FF772CECDCAF" bold="true" box="[199,425,150,176]" pageId="40" pageNumber="41">Number of species.</emphasis>
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1.
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</paragraph>
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<paragraph id="8BC8B83BFFDAD722FF43FF5B2F71DCCB" blockId="40.[151,1437,150,645]" box="[199,564,186,212]" pageId="40" pageNumber="41">
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<emphasis id="B9036429FFDAD722FF43FF5B2CF2DCCB" bold="true" box="[199,439,186,212]" pageId="40" pageNumber="41">Sexual dimorphism.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFDAD722FF43FF3F2805DE23" pageId="40" pageNumber="41" type="discussion">
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<paragraph id="8BC8B83BFFDAD722FF43FF3F2805DE23" blockId="40.[151,1437,150,645]" pageId="40" pageNumber="41">
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<emphasis id="B9036429FFDAD722FF43FF3F2C7EDCE7" bold="true" box="[199,315,222,248]" pageId="40" pageNumber="41">Remarks.</emphasis>
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Treated as a genus in
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<bibRefCitation id="EFE6C5CAFFDAD722FDC0FF3F2E91DCE7" author="Edgecombe, G. D. & Bonato, L." box="[580,980,222,249]" pageId="40" pageNumber="41" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 400)</bibRefCitation>
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,
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<bibRefCitation id="EFE6C5CAFFDAD722FC66FF3E29B9DCE7" author="Bonato, L. & Orlando, M. & Zapparoli, M. & Fusco, G. & Bortolin, F." box="[994,1276,222,249]" pageId="40" pageNumber="41" pagination="1 - 23" refId="ref45789" refString="Bonato, L., Orlando, M., Zapparoli, M., Fusco, G. & Bortolin, F. (2017) New insights into Plutonium, one of the largest and least known European centipedes (Chilopoda): distribution, evolution and morphology. Zoological Journal of the Linnean Society, 20, 1 - 23. https: // doi. org / 10.1093 / zoolinnean / zlw 026" type="journal article" year="2017">
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Bonato
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<emphasis id="B9036429FFDAD722FBB9FF3E293DDCE7" box="[1085,1144,222,248]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
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(2017: 19)
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</bibRefCitation>
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. There are no available data on this genus except for the original description. The only real difference between this doubtful monotypic genus (known from the
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<typeStatus id="54CC0699FFDAD722FE60FEC72F03DD5F" box="[484,582,294,320]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
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only) and the close relative
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<taxonomicName id="4C77C3B8FFDAD722FC02FEC72EADDD5F" authorityName="Meinert" authorityYear="1886" box="[902,1000,294,320]" class="Chilopoda" family="Scolopendridae" genus="Asanada" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFDAD722FC02FEC72EADDD5F" box="[902,1000,294,320]" italics="true" pageId="40" pageNumber="41">Asanada</emphasis>
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</taxonomicName>
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seems to be a presence of short paired cephalic sutures in the former. The very uncertain and minor differences in the length of antennae, shape/length of a dorsal brush of maxillae 2, and the shape of lateral margins[!] of tergites 1–2 etc. mentioned by
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<bibRefCitation id="EFE6C5CAFFDAD722FB7EFE8F2C34DDB3" author="Edgecombe, G. D. & Bonato, L." pageId="40" pageNumber="41" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 400)</bibRefCitation>
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are at most of intrageneric level and do not support generic status of
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<taxonomicName id="4C77C3B8FFDAD722FBF1FE732846DDB3" authorityName="Wurmli" authorityYear="1972" box="[1141,1283,402,428]" class="Chilopoda" family="Scolopendridae" genus="Asanadopsis" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFDAD722FBF1FE732846DDB3" box="[1141,1283,402,428]" italics="true" pageId="40" pageNumber="41">Asanadopsis</emphasis>
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</taxonomicName>
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. For example two adult specimens of
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<taxonomicName id="4C77C3B8FFDAD722FE1BFE572F08DDCF" authorityName="Jangi & Dass" authorityYear="1984" box="[415,589,438,464]" class="Chilopoda" family="Scolopendridae" genus="Asanada" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="indica">
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<emphasis id="B9036429FFDAD722FE1BFE572F08DDCF" box="[415,589,438,464]" italics="true" pageId="40" pageNumber="41">Asanada indica</emphasis>
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</taxonomicName>
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mentioned above (Rc 7484) have the antenna barely reaching tergite 1 (
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<figureCitation id="134CA4BEFFDAD722FAEBFE572DF1DDEB" captionStart="FIGURES 78–83" captionStartId="42.[151,264,1708,1732]" captionTargetBox="[168,1421,193,1665]" captionTargetId="figure-19@42.[151,1436,181,1684]" captionTargetPageId="42" captionText="FIGURES 78–83. Asanada sokotrana Pocock, 1899; BMNH 1934.2.15.151–154 78 LBS 21 ventrally; Asanada indica Jangi & Dass, 1984; Smaller ad. Rc 7484 79 head + LBS 1 dorsally; Asanada brevicornis Meinert, 1886; BMNH 1889.7.15.21–23 80 apical part of maxilla 2 dorso-laterally; Asanada sokotrana Pocock, 1899; BMNH 1899.7.4.85–86 81 LBS 20–21 and ultimate legs dorsally; Otostigmus (O.) multidens Haase, 1887; Ad. Rc 7510 82 right pleuron of LBS 3 laterally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6747 83 left side of ultimate LBS + right ultimate leg medially; (an)—antenna, (cx)—coxopleuron, (f)—ultimate femur, (im)—intersclerite membrane, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pts)—non enlarged ultimate pretarsus, (s)—open spiracle with atrium, (su)—spines of ultimate prefemur, (T1)—tergite 1, (t1)—tarsus 1, (t2)—tarsus 2, (ti)—tibia, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402181" httpUri="https://zenodo.org/record/4402181/files/figure.png" pageId="40" pageNumber="41">Fig. 79</figureCitation>
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) and the dorsal brush of maxillae 2 very short. Also
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<bibRefCitation id="EFE6C5CAFFDAD722FC80FE3B2E9DDDEA" author="Wurmli, M." box="[772,984,474,501]" pageId="40" pageNumber="41" pagination="88 - 104" refId="ref51745" refString="Wurmli, M. (1972) Chilopoda von Sumba und Flores I. Scolopendromorpha. Verhandlungen der Naturforschenden Gesellschaft in Basel, 82 (1), 88 - 104." type="journal article" year="1972">Würmli (1972: 95)</bibRefCitation>
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wrote that his
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<taxonomicName id="4C77C3B8FFDAD722FB04FE3B282EDDEB" authorityName="Wurmli" authorityYear="1972" box="[1152,1387,474,500]" class="Chilopoda" family="Scolopendridae" genus="Asanadopsis" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="nueschi">
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<emphasis id="B9036429FFDAD722FB04FE3B282EDDEB" box="[1152,1387,474,500]" italics="true" pageId="40" pageNumber="41">Asanadopsis nueschi</emphasis>
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</taxonomicName>
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“im ganzen steht sie aber der
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<emphasis id="B9036429FFDAD722FE38FE1F2FDFDE07" box="[444,666,510,536]" italics="true" pageId="40" pageNumber="41">Asanada-philippina</emphasis>
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-gruppe sehr nahe”. Thus, in our opinion, this form seems to be no more than a species of
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<taxonomicName id="4C77C3B8FFDAD722FE1DFDC32CBEDE23" authorityName="Meinert" authorityYear="1886" box="[409,507,546,572]" class="Chilopoda" family="Scolopendridae" genus="Asanada" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFDAD722FE1DFDC32CBEDE23" box="[409,507,546,572]" italics="true" pageId="40" pageNumber="41">Asanada</emphasis>
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</taxonomicName>
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, but we prefer to retain it until the type series is found and re-investigated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFDAD722FF43FDA72974DE9B" pageId="40" pageNumber="41" type="reference_group">
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<paragraph id="8BC8B83BFFDAD722FF43FDA72974DE9B" blockId="40.[151,1437,150,645]" pageId="40" pageNumber="41">
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<treatmentCitationGroup id="AB679F15FFDAD722FF43FDA72974DE9B" pageId="40" pageNumber="41">
|
||
<taxonomicName id="4C77C3B8FFDAD722FF43FDA72CF4DE7F" ID-CoL="5VRVN" authorityName="Wurmli" authorityYear="1972" box="[199,433,582,608]" class="Chilopoda" family="Scolopendridae" genus="Asanadopsis" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="nueschi">
|
||
<emphasis id="B9036429FFDAD722FF43FDA72CF4DE7F" box="[199,433,582,608]" italics="true" pageId="40" pageNumber="41">Asanadopsis nueschi</emphasis>
|
||
</taxonomicName>
|
||
was described from “Mau Marru, Ostsumba” (East Sumba,
|
||
<collectingCountry id="F360F8ABFFDAD722FBCAFDA729FADE7E" box="[1102,1215,582,609]" name="Indonesia" pageId="40" pageNumber="41">Indonesia</collectingCountry>
|
||
), but
|
||
<treatmentCitation id="0AD69E2AFFDAD722FB78FDA72C37DE9B" author="Edgecombe, G. D. & Bonato, L." page="400" pageId="40" pageNumber="41" year="2011">
|
||
<bibRefCitation id="EFE6C5CAFFDAD722FB78FDA72C37DE9B" author="Edgecombe, G. D. & Bonato, L." pageId="40" pageNumber="41" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 400)</bibRefCitation>
|
||
</treatmentCitation>
|
||
erroneously stated the range of
|
||
<taxonomicName id="4C77C3B8FFDAD722FD5DFD8B2E22DE9B" ID-CoL="33KD" authorityName="Wurmli" authorityYear="1972" box="[729,871,618,644]" class="Chilopoda" family="Scolopendridae" genus="Asanadopsis" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDAD722FD5DFD8B2E22DE9B" box="[729,871,618,644]" italics="true" pageId="40" pageNumber="41">Asanadopsis</emphasis>
|
||
</taxonomicName>
|
||
to be “
|
||
<collectingRegion id="49B376D9FFDAD722FC32FD8B2966DE9B" box="[950,1059,618,644]" country="Indonesia" name="Sulawesi Utara" pageId="40" pageNumber="41">Sulawesi</collectingRegion>
|
||
”.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |