treatments-xml/data/03/C6/87/03C687F7FF914E59FF512E5F126A8E8E.xml

<document id="BE12DC1503B34D4F1CB1C0E7C3B6B568" ID-CLB-Dataset="71624" ID-DOI="10.11646/phytotaxa.550.1.4" ID-GBIF-Dataset="5e426e82-dcd9-445e-a608-445b63c44164" ID-ISSN="1179-3163" ID-Zenodo-Dep="6630634" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654850953123" checkinUser="plazi" docAuthor="Büneker, Henrique Mallmann &amp; Mariath, Jorge Ernesto De Araujo" docDate="2022" docId="03C687F7FF914E59FF512E5F126A8E8E" docLanguage="en" docName="phytotaxa.550.1.4.pdf" docOrigin="Phytotaxa 550 (1)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Dyckia pseudodelicata Buneker &amp; Mariath 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="FFFFFF8FFF904E51FFD9294B175C8C5A" masterDocTitle="Dyckia pseudodelicata, a new species of the D. selloa complex (Bromeliaceae, Pitcairnioideae) from Rio Grande do Sul, Brazil" masterLastPageNumber="70" masterPageNumber="59" pageNumber="60" updateTime="1699339256572" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B20F258E4AB9512D6FBE631E91FA1650">Dyckia pseudodelicata, a new species of the D. selloa complex (Bromeliaceae, Pitcairnioideae) from Rio Grande do Sul, Brazil</mods:title>
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<mods:name id="74DB1F058531C57346B3EF835A2DC709" type="personal">
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<mods:namePart id="8D00CB348802987B6C6195F0FFDEF2AD">Büneker, Henrique Mallmann</mods:namePart>
<mods:affiliation id="2E421A51178AAC4D97331541DE673D13">Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; Programa de Pós-graduação em Botânica, Instituto de Biociências, UFRGS, Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:namePart id="7EB93EA57B0DDF2424CF260166854BB7">Mariath, Jorge Ernesto De Araujo</mods:namePart>
<mods:nameIdentifier id="27406AA0E1EEF09A89CE1F7C18B51E27" type="ORCID">0000-0002-2776-6781</mods:nameIdentifier>
<mods:affiliation id="4F55B4FB9B67D5580FF8720F19CE7C9C">Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; jorge. mariath @ ufrgs. br; https: // orcid. org / 0000 - 0002 - 2776 - 6781</mods:affiliation>
<mods:nameIdentifier id="1BB2BDD26C51462098DF6AD2A04A4A12" type="email">jorge.mariath@ufrgs.br</mods:nameIdentifier>
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<mods:title id="365CAE820BE1B5F82F1459D55FF45AA0">Phytotaxa</mods:title>
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<mods:date id="63C11C4D4497A9877C87C008E6D24F44">2022</mods:date>
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<mods:number id="E0EF156DA9F4776914CE2DB0AB0A26D1">2022-06-10</mods:number>
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<mods:number id="EE66C1F8B271D220E78FA09FEAACBD23">550</mods:number>
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<paragraph id="8BD036E1FF914E50FF512E5F158A8B74" blockId="1.[136,1082,1812,1839]" box="[136,726,1812,1839]" pageId="1" pageNumber="60">
<heading id="D098818DFF914E50FF512E5F158A8B74" box="[136,726,1812,1839]" fontSize="11" level="2" pageId="1" pageNumber="60" reason="3">
<taxonomicName id="4C6F4D62FF914E50FF512E5F152D8B74" authority="Buneker &amp; Mariath" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[136,625,1812,1839]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="60" phylum="Tracheophyta" rank="species" species="pseudodelicata" status="sp. nov.">
<emphasis id="B91BEAF3FF914E50FF512E5F16D78B75" bold="true" box="[136,395,1812,1839]" italics="true" pageId="1" pageNumber="60">Dyckia pseudodelicata</emphasis>
Büneker &amp; Mariath
</taxonomicName>
, 
<emphasis id="B91BEAF3FF914E50FDA22E5E158A8B74" box="[635,726,1813,1838]" italics="true" pageId="1" pageNumber="60">
<taxonomicNameLabel id="A2285788FF914E50FDA22E5E158F8B74" box="[635,723,1813,1838]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
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</paragraph>
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<subSubSection id="C375656AFF914E50FD072E5F13668B74" box="[734,1082,1812,1838]" pageId="1" pageNumber="60" type="description">
<paragraph id="8BD036E1FF914E50FD072E5F13668B74" blockId="1.[136,1082,1812,1839]" box="[734,1082,1812,1838]" pageId="1" pageNumber="60">
<heading id="D098818DFF914E50FD072E5F13668B74" box="[734,1082,1812,1838]" fontSize="11" level="2" pageId="1" pageNumber="60" reason="3">
<figureCitation id="13542A64FF914E50FD072E5F14288B74" box="[734,884,1812,1838]" captionStart="FIGURE 1" captionStartId="2.[136,229,1799,1821]" captionTargetBox="[151,1436,187,1772]" captionTargetId="figure-25@2.[151,1436,187,1772]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Dyckia pseudodelicata (A H.M. Büneker 635 et al.; B–D Büneker 913 et al.) in habitat. A. Vegetative habit. B. Leaves detail. C. Fertile habit. D. Detail of apical portion of the inflorescence." figureDoi="http://doi.org/10.5281/zenodo.6630636" httpUri="https://zenodo.org/record/6630636/files/figure.png" pageId="1" pageNumber="60">Figure 1A–D</figureCitation>
, 
<figureCitation id="13542A64FF914E50FC592E5F14E28B74" box="[896,958,1812,1838]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="1" pageNumber="60">2A–J</figureCitation>
, and 
<figureCitation id="13542A64FF914E50FC202E5F136A8B74" box="[1017,1078,1812,1838]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="1" pageNumber="60">3A–I</figureCitation>
.
</heading>
</paragraph>
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<subSubSection id="C375656AFF914E50FF512E2B13758473" pageId="1" pageNumber="60" type="diagnosis">
<paragraph id="8BD036E1FF914E50FF512E2B13758473" blockId="1.[136,1452,1888,2089]" pageId="1" pageNumber="60">
<emphasis id="B91BEAF3FF914E50FF512E2B13758473" italics="true" pageId="1" pageNumber="60">
Tantam habet haec species cum Dyckia delicata similitudinem ut primo intuitu foliis eandem diceret, nisi differt ramis inflorescentiae tenuibus (
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diam. vs. crassis, 
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diam.) et pluribus (usque 18 ramis vs. usque 7 ramis), floribus apicum ramorum subverticillatis (vs. laxe dispositis, numquam subverticillatis), floribus minoribus (
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vs. 
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longi), marginibus petalorum cum trichomibus triformibus (flabelliformia, uniseriata-capitata et dendritica vs. cum trichomibus uniformibus: filiformia aut unicellularia, vel sine trichomibus), lobis stigmatis minoribus in anthesem (
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vs. 
<quantity id="4C979B04FF914E50FBFF2EBB13D8845F" box="[1062,1156,2032,2053]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="60" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
longi), et seminibus rudimentis suberectis, in duas series per loculum dispositis (vs. patentibus, in 2–4 series per loculum).
</emphasis>
</paragraph>
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<paragraph id="8BD036E1FF924E53FF512E4C14728B1B" blockId="2.[136,1451,1799,1857]" pageId="2" pageNumber="61">
<emphasis id="B91BEAF3FF924E53FF512E4C17AB8B46" bold="true" box="[136,247,1799,1821]" pageId="2" pageNumber="61">FIGURE 1</emphasis>
. 
<emphasis id="B91BEAF3FF924E53FEDA2E4C166B8B46" bold="true" box="[259,311,1799,1820]" pageId="2" pageNumber="61">A–D.</emphasis>
<taxonomicName id="4C6F4D62FF924E53FEE72E4C15458B46" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[318,537,1799,1820]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="61" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF924E53FEE72E4C15458B46" box="[318,537,1799,1820]" italics="true" pageId="2" pageNumber="61">Dyckia pseudodelicata</emphasis>
</taxonomicName>
(
<emphasis id="B91BEAF3FF924E53FDFE2E4C15648B46" bold="true" box="[551,568,1799,1820]" pageId="2" pageNumber="61">A</emphasis>
<emphasis id="B91BEAF3FF924E53FDE72E43147A8B46" box="[574,806,1799,1821]" italics="true" pageId="2" pageNumber="61">H.M. Büneker 635 et al.</emphasis>
; 
<emphasis id="B91BEAF3FF924E53FCE82E4C14028B46" bold="true" box="[817,862,1799,1820]" pageId="2" pageNumber="61">B–D</emphasis>
<emphasis id="B91BEAF3FF924E53FCBD2E4C13498B46" box="[868,1045,1799,1821]" italics="true" pageId="2" pageNumber="61">Büneker 913 et al.</emphasis>
) in habitat. 
<emphasis id="B91BEAF3FF924E53FB512E4C13C38B46" bold="true" box="[1160,1183,1799,1820]" pageId="2" pageNumber="61">A.</emphasis>
Vegetative habit. 
<emphasis id="B91BEAF3FF924E53FA922E4C123E8B46" bold="true" box="[1355,1378,1799,1820]" pageId="2" pageNumber="61">B.</emphasis>
Leaves detail. 
<emphasis id="B91BEAF3FF924E53FF102E6017BD8B1B" bold="true" box="[201,225,1835,1857]" pageId="2" pageNumber="61">C.</emphasis>
Fertile habit. 
<emphasis id="B91BEAF3FF924E53FEBE2E6016238B1A" bold="true" box="[359,383,1835,1856]" pageId="2" pageNumber="61">D.</emphasis>
Detail of apical portion of the inflorescence.
</paragraph>
</caption>
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<paragraph id="8BD036E1FF924E53FF512E3814538BF7" blockId="2.[136,1452,1904,1965]" pageId="2" pageNumber="61">
<emphasis id="B91BEAF3FF924E53FF512E38179E8BD2" bold="true" box="[136,194,1907,1928]" pageId="2" pageNumber="61">
<collectingRegion id="49ABF803FF924E53FF512E3817E28BD2" box="[136,190,1907,1928]" pageId="2" pageNumber="61">Type</collectingRegion>
:
</emphasis>
— 
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<collectingCountry id="F3787671FF924E53FF072E3816648BD2" box="[222,312,1907,1928]" name="Brazil" pageId="2" pageNumber="61">BRAZIL</collectingCountry>
. 
<collectingRegion id="49ABF803FF924E53FE9B2E3816A18BD3" box="[322,509,1907,1929]" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="61">Rio Grande do Sul</collectingRegion>
: 
<emphasis id="B91BEAF3FF924E53FDD02E38156E8BD3" bold="true" box="[521,562,1907,1929]" pageId="2" pageNumber="61">Jari</emphasis>
, encosta rochosa próxima ao arroio 
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<collectingDate id="EF95E9C9FF924E53FBFE2E3813978BD2" box="[1063,1227,1907,1929]" pageId="2" pageNumber="61" value="2020-01-17">17 January 2020</collectingDate>
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. 
<collectionCode id="ED7EAE24FF924E53FB342E3F12588BD3" box="[1261,1284,1908,1929]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="61" type="Herbarium">M</collectionCode>
. 
<location id="8EB0603AFF924E53FAD62E3812038BD2" LSID="urn:lsid:plazi:treatment:03C687F7FF914E59FF512E5F126A8E8E:8EB0603AFF924E53FAD62E3812038BD2" box="[1295,1375,1907,1928]" country="Brazil" name="Buneker" pageId="2" pageNumber="61" stateProvince="Jari">Büneker</location>
913, 
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. 
<location id="8EB0603AFF924E53FF642EDC17A18BF6" LSID="urn:lsid:plazi:treatment:03C687F7FF914E59FF512E5F126A8E8E:8EB0603AFF924E53FF642EDC17A18BF6" box="[189,253,1943,1964]" country="Brazil" name="Witeck" pageId="2" pageNumber="61" stateProvince="Jari">Witeck</location>
&amp; 
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. 
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. 
<location id="8EB0603AFF924E53FE902EDC16258BF6" LSID="urn:lsid:plazi:treatment:03C687F7FF914E59FF512E5F126A8E8E:8EB0603AFF924E53FE902EDC16258BF6" box="[329,377,1943,1964]" country="Brazil" name="Silva" pageId="2" pageNumber="61" stateProvince="Jari">Silva</location>
</emphasis>
(
<typeStatus id="54D48843FF924E53FE5C2EDC16828BF7" box="[389,478,1943,1965]" pageId="2" pageNumber="61" type="holotype">holotype</typeStatus>
: 
<collectionCode id="ED7EAE24FF924E53FE3E2EDC154B8BF6" box="[487,535,1943,1964]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="2" pageNumber="61">ICN</collectionCode>
!, isotypes: 
<collectionCode id="ED7EAE24FF924E53FD592EDC15978BF6" box="[640,715,1943,1964]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14337" lsid="urn:lsid:biocol.org:col:14337" name="Universidade Federal de Santa Maria" pageId="2" pageNumber="61" type="Herbarium">SMDB</collectionCode>
!, 
<collectionCode id="ED7EAE24FF924E53FD012EDC15A38BF7" box="[728,767,1943,1965]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="2" pageNumber="61" type="Herbarium">RB</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C375656AFF924E52FF512E90148288AA" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61" type="description">
<paragraph id="8BD036E1FF924E52FF512E90148288AA" blockId="2.[136,1451,2011,2074]" lastBlockId="3.[136,1452,158,2056]" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61">
<emphasis id="B91BEAF3FF924E53FF512E90179F8BAC" bold="true" box="[136,195,2011,2038]" italics="true" pageId="2" pageNumber="61">Herb</emphasis>
saxicolous, rhizomatous, 
<quantity id="4C979B04FF924E53FE302E97152B8BAC" box="[489,631,2012,2038]" metricMagnitude="-1" metricUnit="m" metricValue="8.299999999999999" metricValueMax="11.899999999999999" metricValueMin="4.699999999999999" pageId="2" pageNumber="61" unit="m" value="0.83" valueMax="1.19" valueMin="0.47">0.47–1.19 m</quantity>
tall when flowering; 
<emphasis id="B91BEAF3FF924E53FCB22E9714EA8BAF" box="[875,950,2012,2037]" italics="true" pageId="2" pageNumber="61">rosette</emphasis>
<quantity id="4C979B04FF924E53FC672E9013728BAC" box="[958,1070,2011,2038]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.9" metricValueMin="1.5" pageId="2" pageNumber="61" unit="cm" value="22.0" valueMax="29.0" valueMin="15.0">15–29 cm</quantity>
in diameter, symmetrical. 
<emphasis id="B91BEAF3FF924E53FA852E9712F78BAC" bold="true" box="[1372,1451,2012,2038]" italics="true" pageId="2" pageNumber="61">Leaves</emphasis>
50–108, the inner ones suberect, the outer ones patent to reflexed; 
<emphasis id="B91BEAF3FF924E53FC822EB414C58443" box="[859,921,2047,2073]" italics="true" pageId="2" pageNumber="61">blade</emphasis>
7–18 × 
<quantity id="4C979B04FF924E53FC29214B13338440" box="[1008,1135,2048,2074]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="2" pageNumber="61" unit="cm" value="1.5" valueMax="1.9" valueMin="1.1">1.1–1.9 cm</quantity>
, arched, narrowly triangular, flexible, adaxial surface convex, purplish-cinereous, densely lepidote, abaxial face convex, longitudinally veinedstriate, cinereous, lepidote between the veins, margins densely spinose-serrate, apex terminating in a rigid, pungent spine; 
<emphasis id="B91BEAF3FF934E52FF0D29AC16468D5A" box="[212,282,231,256]" italics="true" pageId="3" pageNumber="62">spines</emphasis>
conspicuous, those of the blades margins narrowly triangular, patent or subpatent-retrorse, sometimes hooked at the apex, flexible, lepidote, pinkish-white at the base, whitish at the median portion and at the apex, apex not pungent. 
<emphasis id="B91BEAF3FF934E52FEC7286516E78D13" bold="true" box="[286,443,302,329]" italics="true" pageId="3" pageNumber="62">Inflorescence</emphasis>
axillar, erect or suberect; 
<emphasis id="B91BEAF3FF934E52FD3C286514168D12" box="[741,842,302,328]" italics="true" pageId="3" pageNumber="62">peduncle</emphasis>
<quantity id="4C979B04FF934E52FC8A2865149F8D13" box="[851,963,302,329]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" metricValueMax="5.3" metricValueMin="1.9" pageId="3" pageNumber="62" unit="cm" value="36.0" valueMax="53.0" valueMin="19.0">19–53 cm</quantity>
long, green to purple, devoid of trichomes and pruinose; 
<emphasis id="B91BEAF3FF934E52FEFC2819168A8D36" box="[293,470,338,364]" italics="true" pageId="3" pageNumber="62">peduncle bracts</emphasis>
polystichous, those of the median portion 1.5–2.6 × 
<quantity id="4C979B04FF934E52FBC7281913D48D37" box="[1054,1160,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="3" pageNumber="62" unit="cm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 cm</quantity>
, stramineous, shorter than the internodes; 
<emphasis id="B91BEAF3FF934E52FEE0283D15C88DCA" box="[313,660,374,400]" italics="true" pageId="3" pageNumber="62">fertile portion of inflorescence</emphasis>
composed, once-branched, 
<quantity id="4C979B04FF934E52FC0D283D13198DCB" box="[980,1093,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="3.9" metricValueMax="5.0" metricValueMin="2.8" pageId="3" pageNumber="62" unit="cm" value="39.0" valueMax="50.0" valueMin="28.0">28–50 cm</quantity>
long; 
<emphasis id="B91BEAF3FF934E52FB4B283D12088DCA" box="[1170,1364,374,400]" italics="true" pageId="3" pageNumber="62">first order bracts</emphasis>
similar to peduncle bracts, 
<quantity id="4C979B04FF934E52FEBE28D116BA8DEF" box="[359,486,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="3" pageNumber="62" unit="cm" value="0.9" valueMax="1.3" valueMin="0.5">0.5–1.3 cm</quantity>
long; 
<emphasis id="B91BEAF3FF934E52FDE828D115C48DEE" box="[561,664,410,436]" italics="true" pageId="3" pageNumber="62">branches</emphasis>
<quantity id="4C979B04FF934E52FD7928D115AB8DEE" box="[672,759,410,436]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="4.572" metricValueMin="1.016" pageId="3" pageNumber="62" unit="in" value="11.0" valueMax="18.0" valueMin="4.0">4–18 in</quantity>
number, suberect to patent, 
<quantity id="4C979B04FF934E52FBE528D113C38DEF" box="[1084,1183,410,437]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.7" metricValueMin="0.9" pageId="3" pageNumber="62" unit="cm" value="13.0" valueMax="17.0" valueMin="9.0">9–17 cm</quantity>
long, 
<quantity id="4C979B04FF934E52FB3028D112018DEF" box="[1257,1373,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.7" metricValueMin="2.0" pageId="3" pageNumber="62" unit="mm" value="2.85" valueMax="3.7" valueMin="2.0">2–3.7 mm</quantity>
diam., greenish to purple, devoid of trichomes, pruinose; 
<emphasis id="B91BEAF3FF934E52FD6128F5141F8D82" box="[696,835,446,472]" italics="true" pageId="3" pageNumber="62">floral bracts</emphasis>
1.5–3 × 
<quantity id="4C979B04FF934E52FC7C28F513448D83" box="[933,1048,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="62" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
, oval-triangular, base green, median and apical region soon dry, yellowish, the apex brown, attenuate-acuminate, ecarinate, devoid of trichomes, pruinose, shorter than the sepals. 
<emphasis id="B91BEAF3FF934E52FE4E2B4D16A88E7B" bold="true" box="[407,500,518,545]" italics="true" pageId="3" pageNumber="62">Flowers</emphasis>
<quantity id="4C979B04FF934E52FE242B4D153B8E7B" box="[509,615,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="3" pageNumber="62" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 cm</quantity>
long, inconspicuously pedicellate, patent, polystichous or subverticillate; 
<emphasis id="B91BEAF3FF934E52FF512B6117B88E1E" box="[136,228,554,580]" italics="true" pageId="3" pageNumber="62">pedicels</emphasis>
ca. 
<quantity id="4C979B04FF934E52FECB2B61163B8E1F" box="[274,359,554,581]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="62" unit="mm" value="0.7">0.7 mm</quantity>
long, green, pruinose; 
<emphasis id="B91BEAF3FF934E52FDB02B6115F38E1E" box="[617,687,554,580]" italics="true" pageId="3" pageNumber="62">sepals</emphasis>
2.1–2.5 × 
<quantity id="4C979B04FF934E52FCF12B6114C08E1F" box="[808,924,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="3" pageNumber="62" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
, suborbicular, apex obtuse to rounded, green at the base, yellow at the apex, slightly carinate, devoid of trichomes and pruinose on the abaxial surface, margins with very sparse dendritic trichomes or glabrous; 
<emphasis id="B91BEAF3FF934E52FD5E2B3915968ED6" box="[647,714,626,652]" italics="true" pageId="3" pageNumber="62">petals</emphasis>
8–9 × 
<quantity id="4C979B04FF934E52FCC72B3914C88ED7" box="[798,916,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="3" pageNumber="62" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 mm</quantity>
, not unguiculate, erect to incurving toward the apex, oblanceolate, yellow, glabrous on both surfaces, proximal margins exhibit sparse dendritic trichomes, median margins plicate, exhibit densely arranged uniseriate-capitate or dendritic trichomes, distally margins exhibit narrowtriangular multicellular emergences giving the margins a torn appearance and also exhibit white-hyaline trichomes densely arranged, usually alternating between uniseriate-capitate and flabelliform, apex round-cucullate; 
<emphasis id="B91BEAF3FF934E52FA9F2A4917AA8F1A" italics="true" pageId="3" pageNumber="62">dendritic trichomes</emphasis>
, those of the petal proximal margins with foot with 1–2 living cells, stalk uniseriate with 3–6 cells usually dead or living basal cells, little elongated, region of branching little expanded, with 2–7 branches formed by strongly elongate, dead cells; those of the petal median margins with foot with 1–2 living cells, stalk uniseriate with 2–6 slightly elongate dead cells, branching region expanded with (2–)3–6 anticlinally arranged cells, followed by 3–9 branches formed by elongate dead cells; 
<emphasis id="B91BEAF3FF934E52FE3F2AFC14738F8A" box="[486,815,950,976]" italics="true" pageId="3" pageNumber="62">uniseriate-capitate trichomes</emphasis>
, those of the petal median margins with 3–6 living cells; those distal petal margins with 3–8 living cells; 
<emphasis id="B91BEAF3FF934E52FD6D2A9114F38FAE" box="[692,943,986,1012]" italics="true" pageId="3" pageNumber="62">flabelliform trichomes</emphasis>
of the petal distal margins with foot of 1–2 living cells, stalk uniseriate with 1–2(–3) hardly elongate dead cells, or live basal cells, expanded region with 3–7 anticlinally arranged dead cells, followed by (4–)6–15 branches formed by elongate dead cells; 
<emphasis id="B91BEAF3FF934E52FB7E2D68125D8866" box="[1191,1281,1059,1084]" italics="true" pageId="3" pageNumber="62">stamens</emphasis>
included in the corolla during opening of the thecae, later becoming exserted; 
<emphasis id="B91BEAF3FF934E52FC9D2D0D14FC883A" box="[836,928,1094,1120]" italics="true" pageId="3" pageNumber="62">filament</emphasis>
<quantity id="4C979B04FF934E52FC7F2D0D135F883B" box="[934,1027,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="3" pageNumber="62" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
long, free above the short hypantium, the antepetalous ones adnate to the petals for ca. 
<quantity id="4C979B04FF934E52FD7A2D2115A788DF" box="[675,763,1130,1157]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="62" unit="mm" value="0.3">0.3 mm</quantity>
; 
<emphasis id="B91BEAF3FF934E52FCDC2D21141288DE" box="[773,846,1130,1156]" italics="true" pageId="3" pageNumber="62">anther</emphasis>
<quantity id="4C979B04FF934E52FC8D2D21149988DF" box="[852,965,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="62" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long, not rostrate; 
<emphasis id="B91BEAF3FF934E52FB4C2D20125B88DE" box="[1173,1287,1131,1156]" italics="true" pageId="3" pageNumber="62">sporangia</emphasis>
unequal; 
<emphasis id="B91BEAF3FF934E52FAAA2D2112F788DE" box="[1395,1451,1130,1156]" italics="true" pageId="3" pageNumber="62">pistil</emphasis>
<quantity id="4C979B04FF934E52FF512DC517AD88F3" box="[136,241,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="3" pageNumber="62" unit="cm" value="0.95" valueMax="1.0" valueMin="0.9">0.9–1 cm</quantity>
long; 
<emphasis id="B91BEAF3FF934E52FEE02DC4162588F2" box="[313,377,1167,1192]" italics="true" pageId="3" pageNumber="62">ovary</emphasis>
1.5–2 × 
<quantity id="4C979B04FF934E52FE062DC5153488F3" box="[479,616,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="3" pageNumber="62" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
, green; 
<emphasis id="B91BEAF3FF934E52FD1B2DC5144D88F2" box="[706,785,1166,1192]" italics="true" pageId="3" pageNumber="62">locules</emphasis>
pauciovulate; 
<emphasis id="B91BEAF3FF934E52FC6E2DC514A388F2" box="[951,1023,1166,1192]" italics="true" pageId="3" pageNumber="62">ovules</emphasis>
suberectly positioned, inserted in two rows per locule; 
<emphasis id="B91BEAF3FF934E52FE9B2DF916298896" box="[322,373,1202,1228]" italics="true" pageId="3" pageNumber="62">style</emphasis>
<quantity id="4C979B04FF934E52FEA22DF916848897" box="[379,472,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="62" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
long, yellowish-white; 
<emphasis id="B91BEAF3FF934E52FD062DF814768896" box="[735,810,1203,1228]" italics="true" pageId="3" pageNumber="62">stigma</emphasis>
conduplicate, stigmatic lobes 
<quantity id="4C979B04FF934E52FBA22DF913B18897" box="[1147,1261,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="3" pageNumber="62" unit="mm" value="1.2" valueMax="1.4" valueMin="1.0">1–1.4 mm</quantity>
long, suberect to erect, yellow. 
<emphasis id="B91BEAF3FF934E52FEFC2D9D16DE88AB" bold="true" box="[293,386,1238,1265]" italics="true" pageId="3" pageNumber="62">Capsule</emphasis>
ovoid, dark brown when mature. 
<emphasis id="B91BEAF3FF934E52FD272D9D146388AB" bold="true" box="[766,831,1238,1265]" italics="true" pageId="3" pageNumber="62">Seeds</emphasis>
oblanceoloid.
</paragraph>
</subSubSection>
<subSubSection id="C375656AFF934E52FF642DB1155689DB" pageId="3" pageNumber="62" type="etymology">
<paragraph id="8BD036E1FF934E52FF642DB1155689DB" blockId="3.[136,1452,158,2056]" pageId="3" pageNumber="62">
<emphasis id="B91BEAF3FF934E52FF642DB11614894E" bold="true" box="[189,328,1274,1300]" pageId="3" pageNumber="62">Etymology:</emphasis>
—The specific epithet “pseudodelicata” refers to the vegetative similarity between the new species and 
<taxonomicName id="4C6F4D62FF934E52FF6E2C54166F8962" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[183,307,1310,1336]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF934E52FF6E2C54166F8962" box="[183,307,1310,1336]" italics="true" pageId="3" pageNumber="62">D. delicata</emphasis>
</taxonomicName>
. The new species was not flowering when first collected and was believed to be 
<taxonomicName id="4C6F4D62FF934E52FB6C2C54126D8962" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[1205,1329,1310,1336]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF934E52FB6C2C54126D8962" box="[1205,1329,1310,1336]" italics="true" pageId="3" pageNumber="62">D. delicata</emphasis>
</taxonomicName>
. However, significant morphological differences found after checking its inflorescences revealed it to be a new species, which we decided to call “false” 
<taxonomicName id="4C6F4D62FF934E52FE5E2C2C155F89DA" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[391,515,1382,1408]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF934E52FE5E2C2C155F89DA" box="[391,515,1382,1408]" italics="true" pageId="3" pageNumber="62">D. delicata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C375656AFF934E52FF642CC1149C8AFB" pageId="3" pageNumber="62" type="distribution">
<paragraph id="8BD036E1FF934E52FF642CC1149C8AFB" blockId="3.[136,1452,158,2056]" pageId="3" pageNumber="62">
<emphasis id="B91BEAF3FF934E52FF642CC116AD89FE" bold="true" box="[189,497,1418,1444]" pageId="3" pageNumber="62">Distribution and Habitat:</emphasis>
—The new species occurs as a saxicolous heliophyte on a rocky basaltic escarpment facing north, near the Toropi-mirim stream. Only one population of the species was located in the municipality of Jari, state of 
<collectingRegion id="49ABF803FF934E52FF392C9916EE89B6" box="[224,434,1490,1516]" country="Brazil" name="Rio Grande do Sul" pageId="3" pageNumber="62">Rio Grande do Sul</collectingRegion>
, southern 
<collectingCountry id="F3787671FF934E52FDFD2C99153089B6" box="[548,620,1490,1516]" name="Brazil" pageId="3" pageNumber="62">Brazil</collectingCountry>
. Several similar environments occur in the region, and so it is possible that there are further populations of the species. Some similar environments were found within a ca. 
<quantity id="4C979B04FF934E52FB162CBD12558A4A" box="[1231,1289,1526,1552]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="62" unit="km" value="4.0">4 km</quantity>
radius around the known population of 
<taxonomicName id="4C6F4D62FF934E52FE782F5015318A6E" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[417,621,1562,1588]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF934E52FE782F5015318A6E" box="[417,621,1562,1588]" italics="true" pageId="3" pageNumber="62">D. pseudodelicata</emphasis>
</taxonomicName>
, but only populations of 
<taxonomicName id="4C6F4D62FF934E52FC5C2F5014B68A6E" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[901,1002,1562,1588]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF934E52FC5C2F5014B68A6E" box="[901,1002,1562,1588]" italics="true" pageId="3" pageNumber="62">D. selloa</emphasis>
</taxonomicName>
(on a rocky vertical escarpment) and 
<taxonomicName id="4C6F4D62FF934E52FA492F5016D18A03" authority="Smith (1932: 6)" authorityName="Smith" authorityPageNumber="6" authorityYear="1932" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="dusenii">
<emphasis id="B91BEAF3FF934E52FA492F5017858A02" italics="true" pageId="3" pageNumber="62">D. dusenii</emphasis>
<bibRefCitation id="EFFE4B10FF934E52FF072F7516D18A03" author="Smith, L. B." box="[222,397,1598,1625]" pageId="3" pageNumber="62" pagination="3 - 42" refId="ref8928" refString="Smith, L. B. (1932) Studies in the Bromeliaceae - III. Contributions from the Gray Herbarium of Harvard University 98: 3 - 42. https: // doi. org / 10.5962 / p. 336146" type="journal article" year="1932">Smith (1932: 6)</bibRefCitation>
</taxonomicName>
(on a hillside with shallow litholic soil) were found. Two individuals observed in the population of 
<taxonomicName id="4C6F4D62FF934E52FF7E2F28162F8A26" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[167,371,1634,1660]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF934E52FF7E2F28162F8A26" box="[167,371,1634,1660]" italics="true" pageId="3" pageNumber="62">D. pseudodelicata</emphasis>
</taxonomicName>
were probably natural hybrids between 
<taxonomicName id="4C6F4D62FF934E52FCE02F2813598A26" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[825,1029,1634,1660]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF934E52FCE02F2813598A26" box="[825,1029,1634,1660]" italics="true" pageId="3" pageNumber="62">D. pseudodelicata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6F4D62FF934E52FBE52F2813EC8A26" authorityName="Smith" authorityYear="1932" box="[1084,1200,1634,1660]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="dusenii">
<emphasis id="B91BEAF3FF934E52FBE52F2813EC8A26" box="[1084,1200,1634,1660]" italics="true" pageId="3" pageNumber="62">D. dusenii</emphasis>
</taxonomicName>
, and probable hybrids of 
<taxonomicName id="4C6F4D62FF934E52FF7F2FCC16508AFA" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[166,268,1670,1696]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF934E52FF7F2FCC16508AFA" box="[166,268,1670,1696]" italics="true" pageId="3" pageNumber="62">D. selloa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6F4D62FF934E52FE9B2FCC15528AFA" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[322,526,1670,1696]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF934E52FE9B2FCC15528AFA" box="[322,526,1670,1696]" italics="true" pageId="3" pageNumber="62">D. pseudodelicata</emphasis>
</taxonomicName>
were observed in vicinity to 
<taxonomicName id="4C6F4D62FF934E52FC8D2FCC14E58AFA" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[852,953,1670,1696]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF934E52FC8D2FCC14E58AFA" box="[852,953,1670,1696]" italics="true" pageId="3" pageNumber="62">D. selloa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C375656AFF934E59FF642FE0157B8ED6" lastPageId="8" lastPageNumber="67" pageId="3" pageNumber="62" type="discussion">
<paragraph id="8BD036E1FF934E52FF642FE0151C8452" blockId="3.[136,1452,158,2056]" pageId="3" pageNumber="62">
We estimate that the population of 
<taxonomicName id="4C6F4D62FF934E52FD8E2FE0147B8A9E" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[599,807,1706,1732]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF934E52FD8E2FE0147B8A9E" box="[599,807,1706,1732]" italics="true" pageId="3" pageNumber="62">D. pseudodelicata</emphasis>
</taxonomicName>
has about 
<specimenCount id="9D69FD68FF934E52FC712FE113488A9E" box="[936,1044,1706,1732]" pageId="3" pageNumber="62" type="adult">200 adult</specimenCount>
individuals. The population size is being reduced due to habitat loss, as pioneer tree species, mainly 
<taxonomicName id="4C6F4D62FF934E52FCB42F8512FB8AB2" authority="Bentham (1882: 67)" authorityName="Bentham" authorityPageNumber="67" authorityYear="1882" box="[877,1447,1742,1769]" class="Magnoliopsida" family="Rutaceae" genus="Helietta" kingdom="Plantae" order="Sapindales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="apiculata">
<emphasis id="B91BEAF3FF934E52FCB42F8513658AB2" box="[877,1081,1742,1768]" italics="true" pageId="3" pageNumber="62">Helietta apiculata</emphasis>
<bibRefCitation id="EFFE4B10FF934E52FB992F85127F8AB3" author="Bentham, G." box="[1088,1315,1742,1769]" pageId="3" pageNumber="62" pagination="66 - 67" refId="ref6185" refString="Bentham, G. (1882) Plate 1385, Helietta parvifolia. Icones plantarum 14: 66 - 67." type="journal article" year="1882">Bentham (1882: 67)</bibRefCitation>
(Rutaceae)
</taxonomicName>
, are settling in the site and decreasing the availability of light. The natural reduction of the habitat of the species, and of the relictual xerophyte community to which it is associated, is worrying as some rare species of 
<taxonomicName id="4C6F4D62FF934E52FB272E5D12338B6A" box="[1278,1391,1814,1840]" class="Magnoliopsida" family="Cactaceae" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="family">Cactaceae</taxonomicName>
(
<emphasis id="B91BEAF3FF934E52FA5B2E5C12F78B6A" box="[1410,1451,1815,1840]" italics="true" pageId="3" pageNumber="62">i.e.,</emphasis>
<taxonomicName id="4C6F4D62FF934E52FF512E7114408B0F" authority="(Ritter 1966: 50) Guiggi (2012: 6)" authorityName="Guiggi" authorityPageNumber="6" authorityYear="2012" baseAuthorityName="Ritter" baseAuthorityPageNumber="50" baseAuthorityYear="1966" box="[136,796,1850,1877]" class="Magnoliopsida" family="Cactaceae" genus="Eriocephala" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis id="B91BEAF3FF934E52FF512E7116D78B0E" box="[136,395,1850,1876]" italics="true" pageId="3" pageNumber="62">Eriocephala magnifica</emphasis>
(
<bibRefCitation id="EFFE4B10FF934E52FE422E7115168B0F" author="Ritter, F." box="[411,586,1850,1877]" pageId="3" pageNumber="62" pagination="50 - 53" refId="ref8687" refString="Ritter, F. (1966) Eriocactus magnificus Ritter spec. nova. Succulenta 45: 50 - 53." type="journal article" year="1966">Ritter 1966: 50</bibRefCitation>
) 
<bibRefCitation id="EFFE4B10FF934E52FD822E7114408B0F" author="Guiggi, A." box="[603,796,1850,1877]" pageId="3" pageNumber="62" pagination="5 - 9" refId="ref7450" refString="Guiggi, A. (2012) New genera and combinations in the family of Cactaceae Jussieu (Magnoliopsida-Cactales). Cactology 3: 5 - 9." type="journal article" year="2012">Guiggi (2012: 6)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C6F4D62FF934E52FCF32E70165C8B23" authority="(Ritter 1970: 109) Herm (1993: 64)" authorityName="Herm" authorityPageNumber="64" authorityYear="1993" baseAuthorityName="Ritter" baseAuthorityPageNumber="109" baseAuthorityYear="1970" class="Magnoliopsida" family="Cactaceae" form="purpureus" genus="Notocactus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="form" species="horstii">
<emphasis id="B91BEAF3FF934E52FCF32E7014A68B0E" box="[810,1018,1850,1876]" italics="true" pageId="3" pageNumber="62">Notocactus horstii</emphasis>
f. 
<emphasis id="B91BEAF3FF934E52FBC22E7013D18B0E" box="[1051,1165,1851,1876]" italics="true" pageId="3" pageNumber="62">purpureus</emphasis>
(
<bibRefCitation id="EFFE4B10FF934E52FB442E7112078B0E" author="Ritter, F." box="[1181,1371,1850,1876]" pageId="3" pageNumber="62" pagination="108 - 109" refId="ref8708" refString="Ritter, F. (1970) Nieuwe cactussen uit Zuid-Amerika I. Succulenta 49: 108 - 109." type="journal article" year="1970">Ritter 1970: 109</bibRefCitation>
) 
<bibRefCitation id="EFFE4B10FF934E52FAB52E70165C8B23" author="Herm, K." pageId="3" pageNumber="62" pagination="59 - 64" refId="ref7523" refString="Herm, K. (1993) N. horstii (Ritter) forma purpureus (Ritter) Herm forma nova. Internoto 14: 59 - 64." type="journal article" year="1993">Herm (1993: 64)</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C6F4D62FF934E52FEE02E14132F8B22" authority="(Ritter 1979: 169) Gerloff (1993: 93))" authorityName="Gerloff" authorityPageNumber="93" authorityYear="1993" baseAuthorityName="Ritter" baseAuthorityPageNumber="169" baseAuthorityYear="1979" box="[313,1139,1886,1913]" class="Magnoliopsida" family="Cactaceae" form="securituberculatus" genus="Notocactus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="form" species="oxycostatus">
<emphasis id="B91BEAF3FF934E52FEE02E1416808B22" box="[313,476,1887,1912]" italics="true" pageId="3" pageNumber="62">N. oxycostatus</emphasis>
f. 
<emphasis id="B91BEAF3FF934E52FE212E15159B8B22" box="[504,711,1886,1912]" italics="true" pageId="3" pageNumber="62">securituberculatus</emphasis>
(
<bibRefCitation id="EFFE4B10FF934E52FD0C2E1514D28B23" author="Ritter, F." box="[725,910,1886,1913]" pageId="3" pageNumber="62" refId="ref8728" refString="Ritter, F. (1979) Kakteen in Sudamerika. Band 1. Brasilien / Uruguay / Paraguay. Friedrich Ritter Selbstverlag, Spangenberg, 374 pp." type="book" year="1979">Ritter 1979: 169</bibRefCitation>
) 
<bibRefCitation id="EFFE4B10FF934E52FC442E15132F8B22" author="Gerloff, N." box="[925,1139,1886,1912]" pageId="3" pageNumber="62" pagination="83 - 95" refId="ref7293" refString="Gerloff, N. (1993) Notocactus oxycostatus Buining &amp; Brederoo fa. securituberculatus (Ritter) Gerloff comb. et stat. nov. Internoto 14: 83 - 95." type="journal article" year="1993">Gerloff (1993: 93))</bibRefCitation>
</taxonomicName>
occur in association with it, in addition to other species characteristic of this environment (
<emphasis id="B91BEAF3FF934E52FCE12EC8143C8BC6" box="[824,864,1923,1948]" italics="true" pageId="3" pageNumber="62">e.g.</emphasis>
<taxonomicName id="4C6F4D62FF934E52FCBF2EC817F38B9A" authority="(Martius 1829: 36) Wiehler (1978: 72)" authorityName="Wiehler" authorityPageNumber="72" authorityYear="1978" baseAuthorityName="Martius" baseAuthorityPageNumber="36" baseAuthorityYear="1829" class="Magnoliopsida" family="Gesneriaceae" genus="Sinningia" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="sellovii">
<emphasis id="B91BEAF3FF934E52FCBF2EC813748BC6" box="[870,1064,1922,1949]" italics="true" pageId="3" pageNumber="62">Sinningia sellovii</emphasis>
(
<bibRefCitation id="EFFE4B10FF934E52FBEC2EC913AA8BC7" author="Martius, C. F. P." box="[1077,1270,1922,1949]" pageId="3" pageNumber="62" pagination="1 - 198" refId="ref8245" refString="Martius, C. F. P. (1829) Nova Genera et Species Plantarum Brasiliensium. Nova Genera et Species Plantarum 3: 1 - 198." type="journal article" year="1829">Martius 1829: 36</bibRefCitation>
) 
<bibRefCitation id="EFFE4B10FF934E52FADD2EC917F38B9A" author="Wiehler, H." pageId="3" pageNumber="62" pagination="61 - 93" refId="ref9413" refString="Wiehler, H. (1978) Miscellaneous transfers and new species of Neotropical Gesneriaceae. Selbyana 5: 61 - 93." type="journal article" year="1978">Wiehler (1978: 72)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C6F4D62FF934E52FF672EEC16248B9A" box="[190,376,1958,1985]" class="Magnoliopsida" family="Gesneriaceae" genus="Sinningia" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="macrostachya">
<emphasis id="B91BEAF3FF934E52FF672EEC16248B9A" box="[190,376,1958,1985]" italics="true" pageId="3" pageNumber="62">S. macrostachya</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFE4B10FF934E52FE512EED157A8B9A" author="Lindley, J" box="[392,550,1958,1984]" pageId="3" pageNumber="62" pagination="1202" refId="ref8076" refString="Lindley, J (1828) Gesneria macrostachya. Long-spiked Gesneria. Botanical Register 14: t. 1202." type="journal article" year="1828">Lindley 1828</bibRefCitation>
: t. 1202) 
<bibRefCitation id="EFFE4B10FF934E52FD4F2EED14C88B9B" author="Chautems, A. P." box="[662,916,1958,1985]" pageId="3" pageNumber="62" pagination="381 - 388" refId="ref6907" refString="Chautems, A. P. (1990) Taxonomic revision of Sinningia Nees: nomenclatural changes and new synonymies. Candollea 45: 381 - 388." type="journal article" year="1990">Chautems (1990: 386)</bibRefCitation>
(
<taxonomicName id="4C6F4D62FF934E52FC7C2EED131D8B9B" box="[933,1089,1958,1985]" class="Magnoliopsida" family="Gesneriaceae" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="family">Gesneriaceae</taxonomicName>
), 
<taxonomicName id="4C6F4D62FF934E52FB8C2EED156A8BBF" authority="Herbert (1837: 161)" authorityName="Herbert" authorityPageNumber="161" authorityYear="1837" class="Liliopsida" family="Amaryllidaceae" genus="Habranthus" isUncertain="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="pedunculosus">
<emphasis id="B91BEAF3FF934E52FB8C2EED13878B9A" box="[1109,1243,1958,1984]" italics="true" pageId="3" pageNumber="62">Habranthus</emphasis>
aff. 
<emphasis id="B91BEAF3FF934E52FACA2EED12F78B9A" box="[1299,1451,1958,1984]" italics="true" pageId="3" pageNumber="62">pedunculosus</emphasis>
<bibRefCitation id="EFFE4B10FF934E52FF512E8116348BBF" author="Herbert, W." box="[136,360,1994,2021]" pageId="3" pageNumber="62" refId="ref7504" refString="Herbert, W. (1837) Amaryllidaceae. James Ridway &amp; Sons, London, 428 pp." type="book" year="1837">Herbert (1837: 161)</bibRefCitation>
(Amaryllidaceae)
</taxonomicName>
, and 
<taxonomicName id="4C6F4D62FF934E52FDA92E81148E8BBE" box="[624,978,1994,2020]" class="Magnoliopsida" family="Asteraceae" genus="Hatschbachiella" isUncertain="true" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="species" species="tweedieana">
<emphasis id="B91BEAF3FF934E52FDA92E8114748BBE" box="[624,808,1994,2020]" italics="true" pageId="3" pageNumber="62">Hatschbachiella</emphasis>
cf. 
<emphasis id="B91BEAF3FF934E52FC8B2E81148E8BBE" box="[850,978,1994,2020]" italics="true" pageId="3" pageNumber="62">tweedieana</emphasis>
</taxonomicName>
(Hooker &amp; Arnott 1835[1836]: 242) 
<bibRefCitation id="EFFE4B10FF934E52FAAA2E8116C78452" author="King, R. M. &amp; Robinson, H. E." pageId="3" pageNumber="62" pagination="393 - 394" refId="ref7652" refString="King, R. M. &amp; Robinson, H. E. (1972) Studies in the Eupatorieae (Asteraceae). LXXXI. A new genus, Hatschbachiella. Phytologia 23: 393 - 394. https: // doi. org / 10.5962 / bhl. part. 19873" type="journal article" year="1972">King &amp; Robinson (1972: 394)</bibRefCitation>
(
<taxonomicName id="4C6F4D62FF934E52FE702EA515768452" box="[425,554,2030,2056]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="62" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
)).
</paragraph>
<caption id="DF106669FF944E55FF512FA713998B35" ID-DOI="http://doi.org/10.5281/zenodo.6630638" ID-Zenodo-Dep="6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="4" pageNumber="63" startId="4.[136,229,1772,1794]" targetBox="[170,1416,190,1748]" targetPageId="4">
<paragraph id="8BD036E1FF944E55FF512FA713998B35" blockId="4.[136,1451,1772,1903]" pageId="4" pageNumber="63">
<emphasis id="B91BEAF3FF944E55FF512FA7166D8B5B" bold="true" box="[136,305,1772,1794]" pageId="4" pageNumber="63">FIGURE 2. A–J.</emphasis>
<taxonomicName id="4C6F4D62FF944E55FEEE2FA6154D8B58" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[311,529,1773,1794]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="4" pageNumber="63" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF944E55FEEE2FA6154D8B58" box="[311,529,1773,1794]" italics="true" pageId="4" pageNumber="63">Dyckia pseudodelicata</emphasis>
</taxonomicName>
(
<emphasis id="B91BEAF3FF944E55FDC72FA615918B58" box="[542,717,1773,1794]" italics="true" pageId="4" pageNumber="63">Büneker 913 et al.</emphasis>
), 
<emphasis id="B91BEAF3FF944E55FD062FA6144E8B58" bold="true" box="[735,786,1773,1794]" pageId="4" pageNumber="63">K–T.</emphasis>
<taxonomicName id="4C6F4D62FF944E55FCC12FA614F28B58" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[792,942,1773,1794]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="4" pageNumber="63" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF944E55FCC12FA614F28B58" box="[792,942,1773,1794]" italics="true" pageId="4" pageNumber="63">Dyckia delicata</emphasis>
</taxonomicName>
(
<emphasis id="B91BEAF3FF944E55FC622FA613368B58" box="[955,1130,1773,1794]" italics="true" pageId="4" pageNumber="63">Büneker 395 et al.</emphasis>
). 
<emphasis id="B91BEAF3FF944E55FBA52FA713848B58" bold="true" box="[1148,1240,1772,1794]" pageId="4" pageNumber="63">A and K.</emphasis>
Median region of leaf adaxial face. 
<emphasis id="B91BEAF3FF944E55FED02E5A163F8B7C" bold="true" box="[265,355,1809,1830]" pageId="4" pageNumber="63">B and L.</emphasis>
Median region of leaf abaxial face. 
<emphasis id="B91BEAF3FF944E55FD1D2E5B14798B7C" bold="true" box="[708,805,1808,1830]" pageId="4" pageNumber="63">C and M.</emphasis>
Detail of adaxial face leaf margin with spines. 
<emphasis id="B91BEAF3FF944E55FB2B2E5A120C8B7C" bold="true" box="[1266,1360,1809,1830]" pageId="4" pageNumber="63">D and N.</emphasis>
Detail of abaxial face leaf margin with spines. 
<emphasis id="B91BEAF3FF944E55FE282E7E15118B10" bold="true" box="[497,589,1844,1866]" pageId="4" pageNumber="63">E and O.</emphasis>
Details of trichomes on the surface of the leaf adaxial surface. 
<emphasis id="B91BEAF3FF944E55FB6A2E7E125A8B10" bold="true" box="[1203,1286,1845,1866]" pageId="4" pageNumber="63">F and P.</emphasis>
Main axis of the inflorescence. 
<emphasis id="B91BEAF3FF944E55FECD2E13162E8B34" bold="true" box="[276,370,1880,1902]" pageId="4" pageNumber="63">G and Q.</emphasis>
Detail of flower in anthesis. 
<emphasis id="B91BEAF3FF944E55FD502E1215BA8B34" bold="true" box="[649,742,1881,1902]" pageId="4" pageNumber="63">H and R.</emphasis>
Detail of buds. 
<emphasis id="B91BEAF3FF944E55FC592E1214938B34" bold="true" box="[896,975,1880,1902]" pageId="4" pageNumber="63">I and S.</emphasis>
Anther. 
<emphasis id="B91BEAF3FF944E55FBF82E12132F8B34" bold="true" box="[1057,1139,1881,1902]" pageId="4" pageNumber="63">J and T.</emphasis>
Stigma.
</paragraph>
</caption>
<paragraph id="8BD036E1FF944E54FF642ED613AF8D5A" blockId="4.[136,1452,1949,2084]" lastBlockId="5.[136,1452,158,256]" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63">
The vegetational physiognomy in which the xerophytic relict is inserted, and where the new species occurs, is a transitional region between the Pampa grasslands and the seasonal forest of the Atlantic Forest domain. This transition is located at the edge of the Southern Brazilian Plateau, in the central region of the state of 
<collectingRegion id="49ABF803FF944E55FB522EAE123C8BA5" box="[1163,1376,2021,2047]" country="Brazil" name="Rio Grande do Sul" pageId="4" pageNumber="63">Rio Grande do Sul</collectingRegion>
, in the hydrographic basin of the Toropi-mirim stream, a tributary of the Toropi river. Taxonomic studies of plants in this region have revealed a significant number of endemic and rare (
<bibRefCitation id="EFFE4B10FF954E54FC8A29D5131B8CE2" author="Marchiori, J. N. C. &amp; Canto-Dorow, T. S. &amp; Buneker, H. M. &amp; Essi, L. &amp; Breier, T. B. &amp; Pontes, R. C." box="[851,1095,158,184]" pageId="5" pageNumber="64" pagination="1 - 16" refId="ref8159" refString="Marchiori, J. N. C., Canto-Dorow, T. S., Buneker, H. M., Essi, L., Breier, T. B. &amp; Pontes, R. C. (2014) Campos e florestas no curso medio do Rio Toropi, Rio Grande do Sul (Brasil). Retrato de um admiravel patrimonio ameacado. Balduinia 45: 1 - 16. https: // doi. org / 10.5902 / 2358198014685" type="journal article" year="2014">
Marchiori 
<emphasis id="B91BEAF3FF954E54FC1529D413588CE2" box="[972,1028,158,184]" italics="true" pageId="5" pageNumber="64">et al.</emphasis>
2014
</bibRefCitation>
; 
<bibRefCitation id="EFFE4B10FF954E54FB8829D512F88CE3" author="Buneker, H. M. &amp; Witeck-Neto, L." box="[1105,1444,158,185]" pageId="5" pageNumber="64" pagination="1 - 14" refId="ref6392" refString="Buneker, H. M. &amp; Witeck-Neto, L. (2016) Levantamento de Bromeliaceae na regiao do curso medio do rio Toropi, Rio Grande do Sul, Brasil. Balduinia 52: 1 - 14. https: // doi. org / 10.5902 / 2358198022371" type="journal article" year="2016">Büneker &amp; Witeck-Neto 2016</bibRefCitation>
) or new species over the last two decades (
<bibRefCitation id="EFFE4B10FF954E54FDBD298914518C86" author="Menezes, L. C." box="[612,781,194,220]" pageId="5" pageNumber="64" pagination="58 - 63" refId="ref8312" refString="Menezes, L. C. (2009) Uma nova especie do Rio Grande do Sul: Cyrtopodium witeckii. Boletim CAOB 75: 58 - 63." type="journal article" year="2009">Menezes 2009</bibRefCitation>
; 
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; 
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Büneker 
<emphasis id="B91BEAF3FF954E54FB48298813968C86" box="[1169,1226,194,220]" italics="true" pageId="5" pageNumber="64">et al.</emphasis>
2013
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, 
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; Deble &amp; Alvez 2013; 
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Silveira 
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2019
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).
</paragraph>
<caption id="DF106669FF954E54FF512E5017E08453" ID-DOI="http://doi.org/10.5281/zenodo.6630640" ID-Zenodo-Dep="6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="5" pageNumber="64" startId="5.[136,229,1819,1841]" targetBox="[183,1404,281,1794]" targetPageId="5">
<paragraph id="8BD036E1FF954E54FF512E5017E08453" blockId="5.[136,1453,1819,2057]" pageId="5" pageNumber="64">
<emphasis id="B91BEAF3FF954E54FF512E5017A48B6A" bold="true" box="[136,248,1819,1841]" pageId="5" pageNumber="64">FIGURE 3</emphasis>
. 
<emphasis id="B91BEAF3FF954E54FEDC2E50166D8B6A" bold="true" box="[261,305,1819,1840]" pageId="5" pageNumber="64">A–I.</emphasis>
<taxonomicName id="4C6F4D62FF954E54FEE02E5015488B6A" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[313,532,1819,1840]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="5" pageNumber="64" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF954E54FEE02E5015488B6A" box="[313,532,1819,1840]" italics="true" pageId="5" pageNumber="64">Dyckia pseudodelicata</emphasis>
</taxonomicName>
(
<emphasis id="B91BEAF3FF954E54FDFA2E50158B8B6A" box="[547,727,1819,1841]" italics="true" pageId="5" pageNumber="64">Büneker 913 et al.</emphasis>
), 
<emphasis id="B91BEAF3FF954E54FD322E5014478B6B" bold="true" box="[747,795,1819,1841]" pageId="5" pageNumber="64">J–O.</emphasis>
<taxonomicName id="4C6F4D62FF954E54FCFB2E5014E68B6A" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[802,954,1819,1840]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="5" pageNumber="64" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF954E54FCFB2E5014E68B6A" box="[802,954,1819,1840]" italics="true" pageId="5" pageNumber="64">Dyckia delicata</emphasis>
</taxonomicName>
(
<emphasis id="B91BEAF3FF954E54FC102E5013208B6A" box="[969,1148,1819,1841]" italics="true" pageId="5" pageNumber="64">Büneker 395 et al.</emphasis>
). 
<emphasis id="B91BEAF3FF954E54FB492E5012788B6A" bold="true" box="[1168,1316,1819,1840]" pageId="5" pageNumber="64">A–F and J–M.</emphasis>
Petal margin. 
<emphasis id="B91BEAF3FF954E54FF512E7417FC8B0E" bold="true" box="[136,160,1855,1876]" pageId="5" pageNumber="64">A.</emphasis>
Trichomes of the proximal margin. 
<emphasis id="B91BEAF3FF954E54FE232E74154C8B0E" bold="true" box="[506,528,1855,1876]" pageId="5" pageNumber="64">B.</emphasis>
Trichomes of median margin. 
<emphasis id="B91BEAF3FF954E54FCEF2E7414128B0F" bold="true" box="[822,846,1855,1877]" pageId="5" pageNumber="64">C.</emphasis>
Trichomes of the distal margin. 
<emphasis id="B91BEAF3FF954E54FB5C2E7413C18B0E" bold="true" box="[1157,1181,1855,1876]" pageId="5" pageNumber="64">D.</emphasis>
Dendritic trichome on DIC. 
<emphasis id="B91BEAF3FF954E54FF512E2817C28B22" bold="true" box="[136,158,1891,1912]" pageId="5" pageNumber="64">E.</emphasis>
Flabelliform trichome on DIC. 
<emphasis id="B91BEAF3FF954E54FE0E2E2816B68B22" bold="true" box="[471,490,1891,1912]" pageId="5" pageNumber="64">F.</emphasis>
Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. 
<emphasis id="B91BEAF3FF954E54FB042E2813B38B22" bold="true" box="[1245,1263,1891,1912]" pageId="5" pageNumber="64">J.</emphasis>
Emergences of the distal margin. 
<emphasis id="B91BEAF3FF954E54FECD2ECC16718BC6" bold="true" box="[276,301,1927,1948]" pageId="5" pageNumber="64">K.</emphasis>
Unicellular trichomes of the apical region. 
<emphasis id="B91BEAF3FF954E54FD0E2ECC15B28BC6" bold="true" box="[727,750,1927,1948]" pageId="5" pageNumber="64">L.</emphasis>
Emergences on DIC. 
<emphasis id="B91BEAF3FF954E54FC1F2ECC14BF8BC6" bold="true" box="[966,995,1927,1948]" pageId="5" pageNumber="64">M.</emphasis>
Unicellular filiform trichomes on DIC. 
<emphasis id="B91BEAF3FF954E54FAB22ECC17FC8B9A" bold="true" pageId="5" pageNumber="64">G and N.</emphasis>
Transverse section of the median region of ovary. 
<emphasis id="B91BEAF3FF954E54FD5C2EE015BF8B9B" bold="true" box="[645,739,1963,1985]" pageId="5" pageNumber="64">H and O.</emphasis>
Detail of transverse section of median region of ovary, where two rows of ovules can be seen in 
<emphasis id="B91BEAF3FF954E54FE852E8416338BBE" bold="true" box="[348,367,1999,2020]" pageId="5" pageNumber="64">H</emphasis>
and four in 
<emphasis id="B91BEAF3FF954E54FE3F2E8416A58BBF" bold="true" box="[486,505,1999,2021]" pageId="5" pageNumber="64">O</emphasis>
. 
<emphasis id="B91BEAF3FF954E54FDDC2E84150D8BBE" bold="true" box="[517,593,1999,2020]" pageId="5" pageNumber="64">I and P</emphasis>
. Seed. 
<emphasis id="B91BEAF3FF954E54FD412E8414CC8BBF" bold="true" box="[664,912,1999,2021]" pageId="5" pageNumber="64">E, G, H, L, M, N and O.</emphasis>
Stained with Safranin. 
<emphasis id="B91BEAF3FF954E54FBAA2E8413968BBE" bold="true" box="[1139,1226,1999,2020]" pageId="5" pageNumber="64">D and F.</emphasis>
Stained with Toluidine Blue.
</paragraph>
</caption>
<paragraph id="8BD036E1FF964E57FF6429D513B88DCA" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="65">
<emphasis id="B91BEAF3FF964E57FF6429D515678CE2" bold="true" box="[189,571,158,184]" pageId="6" pageNumber="65">Observations and discussion:—</emphasis>
The new species is morphologically similar to 
<taxonomicName id="4C6F4D62FF964E57FB8029D4138B8CE2" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[1113,1239,158,184]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FB8029D4138B8CE2" box="[1113,1239,158,184]" italics="true" pageId="6" pageNumber="65">D. delicata</emphasis>
</taxonomicName>
, mainly due to its vegetative aspect (
<figureCitation id="13542A64FF964E57FE8E298916868C86" box="[343,474,194,220]" captionStart="FIGURE 1" captionStartId="2.[136,229,1799,1821]" captionTargetBox="[151,1436,187,1772]" captionTargetId="figure-25@2.[151,1436,187,1772]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Dyckia pseudodelicata (A H.M. Büneker 635 et al.; B–D Büneker 913 et al.) in habitat. A. Vegetative habit. B. Leaves detail. C. Fertile habit. D. Detail of apical portion of the inflorescence." figureDoi="http://doi.org/10.5281/zenodo.6630636" httpUri="https://zenodo.org/record/6630636/files/figure.png" pageId="6" pageNumber="65">Figs. 1A–B</figureCitation>
, and 
<figureCitation id="13542A64FF964E57FDCC298915C38C86" box="[533,671,194,220]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">2A–E, K–O</figureCitation>
), which is unique among the species of the 
<taxonomicName id="4C6F4D62FF964E57FB5B298813BB8C86" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[1154,1255,194,220]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FB5B298813BB8C86" box="[1154,1255,194,220]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex, as both species have leaf blades that resemble chilopods of the genus 
<taxonomicName id="4C6F4D62FF964E57FC9829AD13978D5B" authority="Linnaeus (1758: 637)" authorityName="Linnaeus" authorityPageNumber="637" authorityYear="1758" box="[833,1227,230,257]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAF3FF964E57FC9829AD14938D5A" box="[833,975,230,256]" italics="true" pageId="6" pageNumber="65">Scolopendra</emphasis>
<bibRefCitation id="EFFE4B10FF964E57FC0F29AD13978D5B" author="Linnaeus, C." box="[982,1227,230,257]" pageId="6" pageNumber="65" refId="ref8096" refString="Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata, vol. 1. Laurentius Salvius, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus (1758: 637)</bibRefCitation>
</taxonomicName>
. The leaf blades are succulent, arched, reflexed, both faces are convex, and also possess dense conspicuous trichomes (
<figureCitation id="13542A64FF964E57FB0C284112288D7E" box="[1237,1396,266,292]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2E and O</figureCitation>
) and densely spiny margins with highly developed, although flexible and not pungent, spines (
<figureCitation id="13542A64FF964E57FB54286512088D13" box="[1165,1364,302,329]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2A–D, K–N</figureCitation>
). These vegetative characteristics have made 
<taxonomicName id="4C6F4D62FF964E57FDF4281815F78D36" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[557,683,338,364]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FDF4281815F78D36" box="[557,683,338,364]" italics="true" pageId="6" pageNumber="65">D. delicata</emphasis>
</taxonomicName>
highly appreciated and widespread among gardeners and collectors of ornamental succulent plants (
<bibRefCitation id="EFFE4B10FF964E57FE37283D152C8DCA" author="Sachs, H." box="[494,624,374,400]" pageId="6" pageNumber="65" pagination="118 - 121" refId="ref8757" refString="Sachs, H. (2011) Searching for Dyckia delicata. Die Bromelie 2011: 118 - 121." type="journal article" year="2011">Sachs 2011</bibRefCitation>
) and bestow 
<taxonomicName id="4C6F4D62FF964E57FCDC283C148D8DCA" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[773,977,374,400]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF964E57FCDC283C148D8DCA" box="[773,977,374,400]" italics="true" pageId="6" pageNumber="65">D. pseudodelicata</emphasis>
</taxonomicName>
with the same potential.
</paragraph>
<paragraph id="8BD036E1FF964E57FF6428D113258EEA" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="65">
<taxonomicName id="4C6F4D62FF964E57FF6428D116318DEE" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[189,365,410,436]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FF6428D116318DEE" box="[189,365,410,436]" italics="true" pageId="6" pageNumber="65">Dyckia delicata</emphasis>
</taxonomicName>
possesses a series of singularities among the species of the 
<taxonomicName id="4C6F4D62FF964E57FBD128D013318DEE" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[1032,1133,410,436]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FBD128D013318DEE" box="[1032,1133,410,436]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex with regards to the morphology of its inflorescences and flowers. The sparsely branched architecture of the inflorescence, being simple or compound, once-branched, with branches only at the base of the fertile portion (when compound), robust branches, relatively large flowers with spiraled stigmatic lobes in pre-anthesis, and patent ovules positioned in relation to the longitudinal axis of the ovary, are features that are absent from most of the remaining species of the 
<taxonomicName id="4C6F4D62FF964E57FB392B6012198E1E" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[1248,1349,554,580]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FB392B6012198E1E" box="[1248,1349,554,580]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex but widely observed among other 
<taxonomicName id="4C6F4D62FF964E57FDDC2B0515E88E32" box="[517,692,590,616]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91BEAF3FF964E57FDDC2B0515088E32" box="[517,596,590,616]" italics="true" pageId="6" pageNumber="65">Dyckia</emphasis>
species.
</taxonomicName>
Despite the great morphological similarity between 
<taxonomicName id="4C6F4D62FF964E57FAD92B0412208E32" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[1280,1404,590,616]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FAD92B0412208E32" box="[1280,1404,590,616]" italics="true" pageId="6" pageNumber="65">D. delicata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6F4D62FF964E57FF512B38160F8ED6" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[136,339,626,652]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF964E57FF512B38160F8ED6" box="[136,339,626,652]" italics="true" pageId="6" pageNumber="65">D. pseudodelicata</emphasis>
</taxonomicName>
, the latter does not possess the morphological singularities mentioned above, having instead various flower and inflorescence character states typical of most species of the 
<taxonomicName id="4C6F4D62FF964E57FC7C2BDC13568EEA" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[933,1034,662,688]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FC7C2BDC13568EEA" box="[933,1034,662,688]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex.
</paragraph>
<paragraph id="8BD036E1FF964E57FF642BF114EA8866" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="65">
<taxonomicName id="4C6F4D62FF964E57FF642BF116E28E8E" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[189,446,698,724]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF964E57FF642BF116E28E8E" box="[189,446,698,724]" italics="true" pageId="6" pageNumber="65">Dyckia pseudodelicata</emphasis>
</taxonomicName>
basically differs from 
<taxonomicName id="4C6F4D62FF964E57FD642BF014658E8E" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[701,825,698,724]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FD642BF014658E8E" box="[701,825,698,724]" italics="true" pageId="6" pageNumber="65">D. delicata</emphasis>
</taxonomicName>
in the fertile aspect (
<tableCitation id="C6ED035AFF964E57FBFF2BF113378E8E" box="[1062,1131,698,724]" captionStart="TABLE 1" captionStartId="7.[136,224,376,400]" captionTargetPageId="7" captionText="TABLE 1. Principal characters distinguishing Dyckia pseudodelicata sp. nov. and Dyckia delicata." httpUri="http://table.plazi.org/id/DF106669FF974E56FF51283313FB8DCA" pageId="6" pageNumber="65" tableUuid="DF106669FF974E56FF51283313FB8DCA">Tab. 1</tableCitation>
) by having an inflorescence with thin (
<quantity id="4C979B04FF964E57FED92B95162A8EA3" box="[256,374,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.4" metricValueMin="2.0" pageId="6" pageNumber="65" unit="mm" value="2.7" valueMax="3.4" valueMin="2.0">2–3.4 mm</quantity>
<emphasis id="B91BEAF3FF964E57FEA62B9416C18EA2" box="[383,413,735,760]" italics="true" pageId="6" pageNumber="65">vs.</emphasis>
robust, 
<quantity id="4C979B04FF964E57FE242B9515008EA3" box="[509,604,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="65" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
diam. [
<figureCitation id="13542A64FF964E57FD6C2B95140B8EA3" box="[693,855,734,761]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2F and P</figureCitation>
]) and numerous first order branches (up to 18 [
<figureCitation id="13542A64FF964E57FAA42B9517F08F47" captionStart="FIGURE 1" captionStartId="2.[136,229,1799,1821]" captionTargetBox="[151,1436,187,1772]" captionTargetId="figure-25@2.[151,1436,187,1772]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Dyckia pseudodelicata (A H.M. Büneker 635 et al.; B–D Büneker 913 et al.) in habitat. A. Vegetative habit. B. Leaves detail. C. Fertile habit. D. Detail of apical portion of the inflorescence." figureDoi="http://doi.org/10.5281/zenodo.6630636" httpUri="https://zenodo.org/record/6630636/files/figure.png" pageId="6" pageNumber="65">Fig. 1D</figureCitation>
] 
<emphasis id="B91BEAF3FF964E57FF642A4817888F46" box="[189,212,771,796]" italics="true" pageId="6" pageNumber="65">vs</emphasis>
. absent or up to 7), flowers in the apical region of the branches subverticillate (
<emphasis id="B91BEAF3FF964E57FBA92A4813D28F46" box="[1136,1166,771,796]" italics="true" pageId="6" pageNumber="65">vs.</emphasis>
flowers spaced [
<figureCitation id="13542A64FF964E57FA882A49178D8F1B" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2H and R</figureCitation>
]) and smaller flowers (
<quantity id="4C979B04FF964E57FE062A6D15178F1B" box="[479,587,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="65" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 cm</quantity>
<emphasis id="B91BEAF3FF964E57FD8C2A6C15308F1A" box="[597,620,807,832]" italics="true" pageId="6" pageNumber="65">vs</emphasis>
. 
<quantity id="4C979B04FF964E57FDA72A6D15B58F1B" box="[638,745,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="6" pageNumber="65" unit="cm" value="1.7" valueMax="2.0" valueMin="1.4">1.4–2 cm</quantity>
in length [
<figureCitation id="13542A64FF964E57FCB32A6D13458F1B" box="[874,1049,806,833]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2G and Q</figureCitation>
]), anthers without a rostrum at the apex (
<emphasis id="B91BEAF3FF964E57FF162A0017B28F3E" box="[207,238,843,868]" italics="true" pageId="6" pageNumber="65">vs.</emphasis>
with a rostrum [
<figureCitation id="13542A64FF964E57FE762A0115168F3F" box="[431,586,842,869]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2I and S</figureCitation>
]) and thecae with sporangia of different lengths (
<emphasis id="B91BEAF3FF964E57FB5D2A0013FE8F3E" box="[1156,1186,843,868]" italics="true" pageId="6" pageNumber="65">vs.</emphasis>
equal [
<figureCitation id="13542A64FF964E57FB212A0112C88F3F" box="[1272,1428,842,869]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2I and S</figureCitation>
]), stigmatic lobes that are erect (
<emphasis id="B91BEAF3FF964E57FE002A2416AB8FD2" box="[473,503,879,904]" italics="true" pageId="6" pageNumber="65">vs.</emphasis>
spiraled) at anthesis and smaller (
<quantity id="4C979B04FF964E57FCAD2A2514BB8FD3" box="[884,999,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="6" pageNumber="65" unit="mm" value="1.2" valueMax="1.4" valueMin="1.0">1–1.4 mm</quantity>
<emphasis id="B91BEAF3FF964E57FC372A2413508FD2" box="[1006,1036,879,904]" italics="true" pageId="6" pageNumber="65">vs.</emphasis>
<quantity id="4C979B04FF964E57FBCA2A2513D98FD3" box="[1043,1157,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="6" pageNumber="65" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
in length [
<figureCitation id="13542A64FF964E57FB252A2512CA8FD3" box="[1276,1430,878,905]" captionStart="FIGURE 2" captionStartId="4.[136,229,1772,1794]" captionTargetBox="[170,1416,190,1748]" captionTargetId="figure-25@4.[170,1416,190,1748]" captionTargetPageId="4" captionText="FIGURE 2. A–J. Dyckia pseudodelicata (Büneker 913 et al.), K–T. Dyckia delicata (Büneker 395 et al.). A and K. Median region of leaf adaxial face. B and L. Median region of leaf abaxial face. C and M. Detail of adaxial face leaf margin with spines. D and N. Detail of abaxial face leaf margin with spines. E and O. Details of trichomes on the surface of the leaf adaxial surface. F and P. Main axis of the inflorescence. G and Q. Detail of flower in anthesis. H and R. Detail of buds. I and S. Anther. J and T. Stigma." figureDoi="http://doi.org/10.5281/zenodo.6630638" httpUri="https://zenodo.org/record/6630638/files/figure.png" pageId="6" pageNumber="65">Fig. 2J and T</figureCitation>
]). Some morphological characters of 
<taxonomicName id="4C6F4D62FF964E57FDD42AD915008FF6" box="[525,604,914,940]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF964E57FDD42AD915008FF6" box="[525,604,914,940]" italics="true" pageId="6" pageNumber="65">Dyckia</emphasis>
</taxonomicName>
have been little explored regarding their use in taxonomy, probably because they are inconspicuous, impractical for identification purposes and difficult to observe with the naked eye and/or in herborized material. Nonetheless, here we explore two unusual characters that proved to differentiate 
<taxonomicName id="4C6F4D62FF964E57FB262A9012278FAE" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[1279,1403,986,1012]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FB262A9012278FAE" box="[1279,1403,986,1012]" italics="true" pageId="6" pageNumber="65">D. delicata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6F4D62FF964E57FF512AB416088842" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[136,340,1022,1048]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF964E57FF512AB416088842" box="[136,340,1022,1048]" italics="true" pageId="6" pageNumber="65">D. pseudodelicata</emphasis>
</taxonomicName>
: the morphology of the trichomes on the margins of the petals and the position of ovules in relation to the longitudinal axis of the ovary and their disposition in the placenta.
</paragraph>
<paragraph id="8BD036E1FF964E57FF642D0D154089FE" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="65">
The margins of the upper portion of the petals of both species possess multicellular, narrowly triangular emergences; however, they are more numerous in 
<taxonomicName id="4C6F4D62FF964E57FDE52D20145788DE" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[572,779,1130,1156]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF964E57FDE52D20145788DE" box="[572,779,1130,1156]" italics="true" pageId="6" pageNumber="65">D. pseudodelicata</emphasis>
</taxonomicName>
, especially in the cucullate region of the apex, giving the margin a torn appearance. The petal margins of 
<taxonomicName id="4C6F4D62FF964E57FD752DC4147688F2" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[684,810,1166,1192]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FD752DC4147688F2" box="[684,810,1166,1192]" italics="true" pageId="6" pageNumber="65">D. delicata</emphasis>
</taxonomicName>
are straighter and the emergences are concentrated only in the margins of the apex (
<figureCitation id="13542A64FF964E57FE0B2DF9152B8897" box="[466,631,1202,1229]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="6" pageNumber="65">Fig. 3J and L</figureCitation>
). The margins of the petals of 
<taxonomicName id="4C6F4D62FF964E57FC3E2DF8133B8896" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[999,1127,1202,1228]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF964E57FC3E2DF8133B8896" box="[999,1127,1202,1228]" italics="true" pageId="6" pageNumber="65">D. delicata</emphasis>
</taxonomicName>
have few trichomes, which usually occur in greater numbers at the apex and are filiform, unicellular and not very elongate (
<figureCitation id="13542A64FF964E57FB352D9D12C388AB" box="[1260,1439,1238,1265]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="6" pageNumber="65">Fig. 3K and M</figureCitation>
). In contrast, 
<taxonomicName id="4C6F4D62FF964E57FECB2DB016BC894E" authorityName="Buneker &amp; Mariath" authorityYear="2022" box="[274,480,1274,1300]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF964E57FECB2DB016BC894E" box="[274,480,1274,1300]" italics="true" pageId="6" pageNumber="65">D. pseudodelicata</emphasis>
</taxonomicName>
has numerous trichomes in the margins of the petals and possesses three patterns of morphology and distribution: well-developed, dendritic, sparse in the proximal portion of the margins (
<figureCitation id="13542A64FF964E57FADC2C5517C38907" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="6" pageNumber="65">Fig. 3A–B and D</figureCitation>
); uniseriate-capitate, multicellular, sparse (and not very numerous) in the distal region of the margins and apex (
<figureCitation id="13542A64FF964E57FAA72C0917F589DA" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="6" pageNumber="65">Fig. 3F</figureCitation>
); and alternated by very numerous flabelliform trichomes (
<figureCitation id="13542A64FF964E57FCE72C2D14C889DA" box="[830,916,1382,1408]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="6" pageNumber="65">Fig. 3E</figureCitation>
), generally well developed, densely arranged in the margins near the apex (
<figureCitation id="13542A64FF964E57FE6E2CC1155389FE" box="[439,527,1418,1444]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="6" pageNumber="65">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8BD036E1FF964E57FF642CE513938B23" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="65">
The morphology and presence of these trichomes is an important diagnostic character to be analyzed in the context of the 
<taxonomicName id="4C6F4D62FF964E57FEF72C9816CA89B6" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[302,406,1490,1516]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FEF72C9816CA89B6" box="[302,406,1490,1516]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex, as it seems to be conserved among individuals of the same species and generally distinct between species. These trichomes are present in almost all species of the 
<taxonomicName id="4C6F4D62FF964E57FBC42CBC13DE8A4A" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[1053,1154,1526,1552]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FBC42CBC13DE8A4A" box="[1053,1154,1526,1552]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex, occurring in the margins and abaxial surfaces of perianth parts. However, they have been poorly characterized for most 
<taxonomicName id="4C6F4D62FF964E57FB272F5112FB8A6E" box="[1278,1447,1562,1588]" pageId="6" pageNumber="65">
<emphasis id="B91BEAF3FF964E57FB272F5112118A6E" box="[1278,1357,1562,1588]" italics="true" pageId="6" pageNumber="65">Dyckia</emphasis>
species
</taxonomicName>
, especially those of the petals, which are absent in most species not belonging to the 
<taxonomicName id="4C6F4D62FF964E57FBF52F7413CD8A02" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[1068,1169,1598,1624]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FBF52F7413CD8A02" box="[1068,1169,1598,1624]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex (
<bibRefCitation id="EFFE4B10FF964E57FADC2F75179F8A26" author="Carvalho, J. D. T. &amp; Essi, L. &amp; Oliveira, J. M. S." pageId="6" pageNumber="65" pagination="29 - 41" refId="ref6830" refString="Carvalho, J. D. T., Essi, L. &amp; Oliveira, J. M. S. (2017) Flower and floral trichome morphology of species of Dyckia Schult. f. (Bromeliaceae, Pitcairnioideae), and their importance to species characterization and genus taxonomy. Acta Botanica Brasilica 31: 29 - 41. https: // doi. org / 10.1590 / 0102 - 33062016 abb 0335" type="journal article" year="2017">
Carvalho 
<emphasis id="B91BEAF3FF964E57FAAD2F7412F08A02" box="[1396,1452,1598,1624]" italics="true" pageId="6" pageNumber="65">et al.</emphasis>
2017
</bibRefCitation>
; 
<bibRefCitation id="EFFE4B10FF964E57FF162F2916EC8A26" author="Buneker, H. M. &amp; Guarconi, E. A. E. &amp; Santos-Silva, F. &amp; Forzza, R. C." box="[207,432,1634,1660]" pageId="6" pageNumber="65" refId="ref6500" refString="Buneker, H. M., Guarconi, E. A. E., Santos-Silva, F. &amp; Forzza, R. C. (2020) Dyckia in Flora do Brasil 2020. Jardim Botanico do Rio de Janeiro. Available from: http: // floradobrasil. jbrj. gov. br / reflora / floradobrasil / FB 6046 (accessed 8 January 2021)." type="url" year="2020">
Büneker 
<emphasis id="B91BEAF3FF964E57FEEC2F2816328A26" box="[309,366,1634,1660]" italics="true" pageId="6" pageNumber="65">et al.</emphasis>
2020
</bibRefCitation>
, 
<bibRefCitation id="EFFE4B10FF964E57FE622F2915018A26" author="Buneker, H. M." box="[443,605,1634,1660]" pageId="6" pageNumber="65" refId="ref6350" refString="Buneker, H. M. (2021) Revisao taxonomica do complexo Dyckia selloa (Bromeliaceae, Pitcairnioideae) baseada em caracteres morfologicos. Masters dissertation, Universidade Federal do Rio Grande do Sul, Porto Alegre, 290 pp." type="book" year="2021">Büneker 2021</bibRefCitation>
). Trichomes like those of the petals of species of the 
<taxonomicName id="4C6F4D62FF964E57FB692F28124A8A26" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[1200,1302,1634,1660]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FB692F28124A8A26" box="[1200,1302,1634,1660]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex can also be observed in species of the 
<taxonomicName id="4C6F4D62FF964E57FDDD2FCC15C08AFA" authorityName="Mez" authorityYear="1896" box="[516,668,1670,1696]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="ferruginea">
<emphasis id="B91BEAF3FF964E57FDDD2FCC15C08AFA" box="[516,668,1670,1696]" italics="true" pageId="6" pageNumber="65">D. ferruginea</emphasis>
</taxonomicName>
complex (
<taxonomicName id="4C6F4D62FF964E57FCCA2FCC13628AFB" authority="Leme (1993: 6)" authorityName="Leme" authorityPageNumber="6" authorityYear="1993" box="[787,1086,1670,1697]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="B91BEAF3FF964E57FCCA2FCC14DB8AFA" box="[787,903,1670,1696]" italics="true" pageId="6" pageNumber="65">D. excelsa</emphasis>
<bibRefCitation id="EFFE4B10FF964E57FC572FCC13628AFB" author="Leme, E. M. C." box="[910,1086,1670,1697]" pageId="6" pageNumber="65" pagination="6 - 7" refId="ref7992" refString="Leme, E. M. C. (1993) Novas bromeliaceaes nativas do Brasil - XI. Pabstia 4: 6 - 7." type="journal article" year="1993">Leme (1993: 6)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C6F4D62FF964E57FB932FCC12FA8AFB" authority="Mez (1896: 533)" authorityName="Mez" authorityPageNumber="533" authorityYear="1896" box="[1098,1446,1670,1697]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="ferruginea">
<emphasis id="B91BEAF3FF964E57FB932FCC13BE8AFA" box="[1098,1250,1670,1696]" italics="true" pageId="6" pageNumber="65">D. ferruginea</emphasis>
<bibRefCitation id="EFFE4B10FF964E57FB302FCC12FA8AFB" author="Mez, C." box="[1257,1446,1670,1697]" pageId="6" pageNumber="65" refId="ref8341" refString="Mez, C. (1896) Bromeliaceae. In: De Candolle, C. (Ed.) Monographiae Phanerogamarum 9. Sumptibus G. Masson &amp; Cie., Paris, 990 pp." type="book" year="1896">Mez (1896: 533)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C6F4D62FF964E57FF512FE016B08A9F" authority="Hassler (1919: 316)" authorityName="Hassler" authorityPageNumber="316" authorityYear="1919" box="[136,492,1706,1733]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="insignis">
<emphasis id="B91BEAF3FF964E57FF512FE0165E8A9E" box="[136,258,1707,1732]" italics="true" pageId="6" pageNumber="65">D. insignis</emphasis>
<bibRefCitation id="EFFE4B10FF964E57FED32FE116B08A9F" author="Hassler, E." box="[266,492,1706,1733]" pageId="6" pageNumber="65" pagination="268 - 341" refId="ref7478" refString="Hassler, E. (1919) Bromeliacearum paraguariensiam conspectus. Annuair du Conservatoire et du Jardin Botaniques de Geneves 20: 268 - 341." type="journal article" year="1919">Hassler (1919: 316)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C6F4D62FF964E57FE202FE013028A9F" authority="(Pereira &amp; Martinelli 1982: 252) Forzza" authorityName="Forzza" baseAuthorityName="Pereira &amp; Martinelli" baseAuthorityPageNumber="252" baseAuthorityYear="1982" box="[505,1118,1706,1733]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="lymaniana">
<emphasis id="B91BEAF3FF964E57FE202FE015CE8A9E" box="[505,658,1706,1732]" italics="true" pageId="6" pageNumber="65">D. lymaniana</emphasis>
(
<bibRefCitation id="EFFE4B10FF964E57FD782FE1135F8A9F" author="Pereira, E. &amp; Martinelli, G." box="[673,1027,1706,1733]" pageId="6" pageNumber="65" pagination="251 - 260" refId="ref8453" refString="Pereira, E. &amp; Martinelli, G. (1982) Species novae in brasilia bromeliacearum XIX. Bradea 3: 251 - 260." type="journal article" year="1982">Pereira &amp; Martinelli 1982: 252</bibRefCitation>
) Forzza
</taxonomicName>
(
<emphasis id="B91BEAF3FF964E57FBB62FE013D98A9E" box="[1135,1157,1707,1732]" italics="true" pageId="6" pageNumber="65">in</emphasis>
Gomes-Da-Silva, Santos-Silva &amp; Forzza 2019: 486), and 
<taxonomicName id="4C6F4D62FF964E57FE222F8414AE8AB2" authority="Pereira &amp; Gouda (2018: 137))" authorityName="Pereira &amp; Gouda" authorityPageNumber="137" authorityYear="2018" box="[507,1010,1742,1768]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="splendida">
<emphasis id="B91BEAF3FF964E57FE222F8415D08AB2" box="[507,652,1742,1768]" italics="true" pageId="6" pageNumber="65">D. splendida</emphasis>
<bibRefCitation id="EFFE4B10FF964E57FD4C2F8514AE8AB2" author="Pereira, E. E. &amp; Gouda, E. J." box="[661,1010,1742,1768]" pageId="6" pageNumber="65" refId="ref8479" refString="Pereira, E. E. &amp; Gouda, E. J. (2018) A new Dyckia species (Bromeliaceae, Pitcairnioideae) from Brazilian Central-West region, Pantanal Mato-Grossense. Phytotaxa 333: 137 - 14. https: // doi. org / 10.11646 / phytotaxa. 333.1.12" type="journal volume" year="2018">Pereira &amp; Gouda (2018: 137))</bibRefCitation>
</taxonomicName>
. Species of the 
<taxonomicName id="4C6F4D62FF964E57FB712F84121E8AB2" authorityName="Mez" authorityYear="1896" box="[1192,1346,1742,1768]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="ferruginea">
<emphasis id="B91BEAF3FF964E57FB712F84121E8AB2" box="[1192,1346,1742,1768]" italics="true" pageId="6" pageNumber="65">D. ferruginea</emphasis>
</taxonomicName>
complex appear related to those of the 
<taxonomicName id="4C6F4D62FF964E57FE002FB815638B56" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[473,575,1778,1804]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FE002FB815638B56" box="[473,575,1778,1804]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex in the phylogenies of 
<bibRefCitation id="EFFE4B10FF964E57FC7C2FB813238B56" author="Krapp, F. &amp; Pinange, D. S. B. &amp; Benko-Iseppon, A. M. &amp; Leme, E. M. C. &amp; Weising, K." box="[933,1151,1778,1805]" pageId="6" pageNumber="65" pagination="1591 - 1614" refId="ref7816" refString="Krapp, F., Pinange, D. S. B., Benko-Iseppon, A. M., Leme, E. M. C. &amp; Weising, K. (2014) Phylogeny and evolution of Dyckia (Bromeliaceae) inferred from chloroplast and nuclear sequences. Plant Systematics and Evolution 300: 1591 - 1614. https: // doi. org / 10.1007 / s 00606 - 014 - 0985 - 0" type="journal article" year="2014">
Krapp 
<emphasis id="B91BEAF3FF964E57FC2A2FB813708B56" box="[1011,1068,1778,1804]" italics="true" pageId="6" pageNumber="65">et al.</emphasis>
(2014)
</bibRefCitation>
and 
<bibRefCitation id="EFFE4B10FF964E57FB6E2FB912F88B56" author="Pinange, D. S. B. &amp; Krapp, F. &amp; Zizka, G. &amp; Silvestro, D. &amp; Leme, E. M. C. &amp; Weising, K. &amp; Benko-Iseppon, A. M." box="[1207,1444,1778,1804]" pageId="6" pageNumber="65" pagination="39 - 56" refId="ref8531" refString="Pinange, D. S. B., Krapp, F., Zizka, G., Silvestro, D., Leme, E. M. C., Weising, K. &amp; Benko-Iseppon, A. M. (2017) Molecular phylogenetics, historical biogeography and character evolution in Dyckia (Bromeliaceae, Pitcairnioideae). Botanical Journal of the Linnean Society 183: 39 - 56. https: // doi. org / 10.1111 / boj. 12489" type="journal article" year="2017">
Pinangé 
<emphasis id="B91BEAF3FF964E57FAC02FB8120E8B56" box="[1305,1362,1778,1804]" italics="true" pageId="6" pageNumber="65">et al.</emphasis>
(2017)
</bibRefCitation>
, and the presence and similarity of petal trichomes in the species of these two complexes may indicative the degree of relationship between them, even though they appear very distinguished. Thus, these trichomes have great potential to be observed in the future under evolutionary, phylogenetic, morphological and functional aspects.
</paragraph>
<paragraph id="8BD036E1FF964E56FF642EC9168B8D12" blockId="6.[136,1452,158,2093]" lastBlockId="7.[136,1452,158,329]" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
The position of the ovules in relation to the longitudinal axis of the ovary and their disposition in the placenta (number of rows) have been little studied in 
<taxonomicName id="4C6F4D62FF964E57FDB72EED15E18B9A" box="[622,701,1958,1984]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF964E57FDB72EED15E18B9A" box="[622,701,1958,1984]" italics="true" pageId="6" pageNumber="65">Dyckia</emphasis>
</taxonomicName>
and are not covered in detail in the descriptive taxonomic literature. 
<bibRefCitation id="EFFE4B10FF964E57FF512E8016508BBF" author="Mez, C." box="[136,268,1994,2021]" pageId="6" pageNumber="65" refId="ref8378" refString="Mez, C. (1935) Bromeliaceae. In: Engler, A. (Ed.) Das Pflanzenreich, regni vegetabilis conspectus 4 (32). W. Engelmann, Leipzig, 667 pp." type="book" year="1935">Mez (1935)</bibRefCitation>
reports that 
<emphasis id="B91BEAF3FF964E57FE442E8115198BBE" box="[413,581,1994,2020]" italics="true" pageId="6" pageNumber="65">Prionophyllum</emphasis>
(genus name, currently synonymous with 
<taxonomicName id="4C6F4D62FF964E57FBF02E8113248BBE" box="[1065,1144,1994,2020]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF964E57FBF02E8113248BBE" box="[1065,1144,1994,2020]" italics="true" pageId="6" pageNumber="65">Dyckia</emphasis>
</taxonomicName>
, which is equivalent to the 
<taxonomicName id="4C6F4D62FF964E57FF512EA417B08452" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[136,236,2030,2056]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FF512EA417B08452" box="[136,236,2030,2056]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
complex) has few ovules and presents an illustration with the longitudinal section of the ovary of 
<taxonomicName id="4C6F4D62FF964E57FAF22EA412D38452" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[1323,1423,2030,2056]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF964E57FAF22EA412D38452" box="[1323,1423,2030,2056]" italics="true" pageId="6" pageNumber="65">D. selloa</emphasis>
</taxonomicName>
in which the ovules are suberect in relation to the axis of the ovary and inserted at the axillary base of the carpel. 
<taxonomicName id="4C6F4D62FF964E56FA852159166E8CE2" authorityName="Buneker &amp; Mariath" authorityYear="2022" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" lastPageId="7" lastPageNumber="66" order="Poales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF964E56FA852159166E8CE2" italics="true" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">Dyckia pseudodelicata</emphasis>
</taxonomicName>
, like 
<taxonomicName id="4C6F4D62FF974E56FEA829D4168A8CE2" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[369,470,158,184]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF974E56FEA829D4168A8CE2" box="[369,470,158,184]" italics="true" pageId="7" pageNumber="66">D. selloa</emphasis>
</taxonomicName>
, possesses ovules that are suberect in relation to the longitudinal axis of the ovary (that is, oriented obliquely to the placentation region) and in a reduced number, which was also verified for most species of the 
<taxonomicName id="4C6F4D62FF974E56FF6B29AC16448D5A" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[178,280,230,256]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF974E56FF6B29AC16448D5A" box="[178,280,230,256]" italics="true" pageId="7" pageNumber="66">D. selloa</emphasis>
</taxonomicName>
complex. One of the exceptions is 
<taxonomicName id="4C6F4D62FF974E56FD7329AC147B8D5A" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[682,807,230,256]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF974E56FD7329AC147B8D5A" box="[682,807,230,256]" italics="true" pageId="7" pageNumber="66">D. delicata</emphasis>
</taxonomicName>
, which possesses ovules that are patent in relation to the longitudinal axis of the ovary and in a considerably larger number, as was usually verified for species of 
<taxonomicName id="4C6F4D62FF974E56FAF3284112258D7E" box="[1322,1401,266,292]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF974E56FAF3284112258D7E" box="[1322,1401,266,292]" italics="true" pageId="7" pageNumber="66">Dyckia</emphasis>
</taxonomicName>
that do not belong to the complex.
</paragraph>
<caption id="DF106669FF974E56FF51283313FB8DCA" ID-Table-UUID="DF106669FF974E56FF51283313FB8DCA" box="[136,1191,376,400]" httpUri="http://table.plazi.org/id/DF106669FF974E56FF51283313FB8DCA" pageId="7" pageNumber="66" startId="7.[136,224,376,400]" targetBox="[149,1393,428,1294]" targetIsTable="true" targetPageId="7">
<paragraph id="8BD036E1FF974E56FF51283313FB8DCA" blockId="7.[136,1191,376,400]" box="[136,1191,376,400]" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF51283317A78DCA" bold="true" box="[136,251,376,400]" pageId="7" pageNumber="66">TABLE 1.</emphasis>
Principal characters distinguishing 
<emphasis id="B91BEAF3FF974E56FDA0283314998DCA" box="[633,965,376,400]" italics="true" pageId="7" pageNumber="66">
<taxonomicName id="4C6F4D62FF974E56FDA0283314308DCA" authority="Büneker &amp; Mariath, 2022" authorityName="Büneker &amp; Mariath" authorityYear="2022" box="[633,876,376,400]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="pseudodelicata" status="sp. nov.">Dyckia pseudodelicata</taxonomicName>
<taxonomicNameLabel id="A2285788FF974E56FCAD283214998DCA" box="[884,965,377,400]" pageId="7" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C6F4D62FF974E56FC21283313FC8DCA" authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[1016,1184,376,400]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF974E56FC21283313FC8DCA" box="[1016,1184,376,400]" italics="true" pageId="7" pageNumber="66">Dyckia delicata</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD036E1FF974E56FF4C28E7126A8954" pageId="7" pageNumber="66">
<table id="F96FC441FF97B1AEFF4C28E7122D8954" box="[149,1393,428,1294]" gridcols="4" gridrows="17" pageId="7" pageNumber="66">
<tr id="355F34A3FF97B1AEFF4C28E7122D8D98" box="[149,1393,428,450]" gridrow="0" pageId="7" pageNumber="66" rowspan-1="1">
<th id="768E5DDFFF97B1AEFF4C28E716E68D98" box="[149,442,428,450]" gridcol="0" gridrow="0" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C28E716548D98" bold="true" box="[149,264,428,450]" pageId="7" pageNumber="66">Characters</emphasis>
</th>
<th id="768E5DDFFF97B1AEFCFE28E7130A8D98" box="[807,1110,428,450]" gridcol="2" gridrow="0" pageId="7" pageNumber="66">
<taxonomicName id="4C6F4D62FF974E56FCFE28E713598D98" authorityName="Büneker &amp; Mariath" authorityYear="2022" box="[807,1029,428,450]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF974E56FCFE28E713598D98" bold="true" box="[807,1029,428,450]" italics="true" pageId="7" pageNumber="66">Dyckia pseudodelicata</emphasis>
</taxonomicName>
</th>
<th id="768E5DDFFF97B1AEFBAE28E7122D8D98" box="[1143,1393,428,450]" gridcol="3" gridrow="0" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FBA128E7124D8D98" bold="true" box="[1144,1297,428,450]" italics="true" pageId="7" pageNumber="66">Dyckia delicata</emphasis>
</th>
</tr>
<tr id="355F34A3FF97B1AEFF4C28A1122D8E5A" box="[149,1393,490,512]" gridrow="1" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D28A115A38E5A" box="[484,767,490,512]" gridcol="1" gridrow="1" pageId="7" pageNumber="66">Diameter</td>
<td id="768E5DDFFF97B1AEFCFE28A1130A8E5A" box="[807,1110,490,512]" gridcol="2" gridrow="1" pageId="7" pageNumber="66">2–3.4 mm</td>
<td id="768E5DDFFF97B1AEFBAE28A1122D8E5A" box="[1143,1393,490,512]" gridcol="3" gridrow="1" pageId="7" pageNumber="66">4–5 mm</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2B42122D8E45" box="[149,1393,521,543]" gridrow="2" pageId="7" pageNumber="66" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="768E5DDFFF97B1AEFF4C2B4216E68E45" box="[149,442,521,543]" gridcol="0" gridrow="2" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C2B4216DC8E44" bold="true" box="[149,384,521,543]" pageId="7" pageNumber="66">Inflorescence branches</emphasis>
</th>
</tr>
<tr id="355F34A3FF97B1AEFF4C2B63122D8E64" box="[149,1393,552,574]" gridrow="3" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2B6315A38E64" box="[484,767,552,574]" gridcol="1" gridrow="3" pageId="7" pageNumber="66">Number</td>
<td id="768E5DDFFF97B1AEFCFE2B63130A8E64" box="[807,1110,552,574]" gridcol="2" gridrow="3" pageId="7" pageNumber="66">up to 18</td>
<td id="768E5DDFFF97B1AEFBAE2B63122D8E64" box="[1143,1393,552,574]" gridcol="3" gridrow="3" pageId="7" pageNumber="66">absent or up to 7</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2B2D122D8EF0" box="[149,1393,614,682]" gridrow="4" pageId="7" pageNumber="66">
<th id="768E5DDFFF97B1AEFF4C2B2D16E68EF0" box="[149,442,614,682]" gridcol="0" gridrow="4" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C2BDE17BA8EF0" bold="true" box="[149,230,661,682]" pageId="7" pageNumber="66">Flowers</emphasis>
</th>
<td id="768E5DDFFF97B1AEFE3D2B2D15A38EF0" box="[484,767,614,682]" gridcol="1" gridrow="4" pageId="7" pageNumber="66">Arrangement in the apical region of the branches</td>
<td id="768E5DDFFF97B1AEFCFE2B2D130A8EF0" box="[807,1110,614,682]" gridcol="2" gridrow="4" pageId="7" pageNumber="66">subverticillate</td>
<td id="768E5DDFFF97B1AEFBAE2B2D122D8EF0" box="[1143,1393,614,682]" gridcol="3" gridrow="4" pageId="7" pageNumber="66">spaced, not subverticillate</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2B8F122D8E80" box="[149,1393,708,730]" gridrow="5" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2B8F15A38E80" box="[484,767,708,730]" gridcol="1" gridrow="5" pageId="7" pageNumber="66">Size</td>
<td id="768E5DDFFF97B1AEFCFE2B8F130A8E80" box="[807,1110,708,730]" gridcol="2" gridrow="5" pageId="7" pageNumber="66">1–1.2 cm</td>
<td id="768E5DDFFF97B1AEFBAE2B8F122D8E80" box="[1143,1393,708,730]" gridcol="3" gridrow="5" pageId="7" pageNumber="66">1.4–2 cm</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2A49122D8F42" box="[149,1393,770,792]" gridrow="6" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2A4915A38F42" box="[484,767,770,792]" gridcol="1" gridrow="6" pageId="7" pageNumber="66">Cells number</td>
<td id="768E5DDFFF97B1AEFCFE2A49130A8F42" box="[807,1110,770,792]" gridcol="2" gridrow="6" pageId="7" pageNumber="66">multicellular</td>
<td id="768E5DDFFF97B1AEFBAE2A49122D8F42" box="[1143,1393,770,792]" gridcol="3" gridrow="6" pageId="7" pageNumber="66">unicellular</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2A6A122D8F6D" box="[149,1393,801,823]" gridrow="7" pageId="7" pageNumber="66" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="768E5DDFFF97B1AEFF4C2A6A16E68F6D" box="[149,442,801,823]" gridcol="0" gridrow="7" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C2A6A16E68F6D" bold="true" box="[149,442,801,823]" pageId="7" pageNumber="66">Trichomes on the margins of</emphasis>
</th>
</tr>
<tr id="355F34A3FF97B1AEFF4C2A0B122D8F2C" box="[149,1393,832,886]" gridrow="8" pageId="7" pageNumber="66">
<th id="768E5DDFFF97B1AEFF4C2A0B16E68F2C" box="[149,442,832,886]" gridcol="0" gridrow="8" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C2A0A17AB8F0C" bold="true" box="[149,247,833,854]" pageId="7" pageNumber="66">the petals</emphasis>
</th>
<td id="768E5DDFFF97B1AEFE3D2A0B15A38F2C" box="[484,767,832,886]" gridcol="1" gridrow="8" pageId="7" pageNumber="66">Shape</td>
<td id="768E5DDFFF97B1AEFCFE2A0B130A8F2C" box="[807,1110,832,886]" gridcol="2" gridrow="8" pageId="7" pageNumber="66">flabelliform, uniseriate-capitate and dendritic</td>
<td id="768E5DDFFF97B1AEFBAE2A0B122D8F2C" box="[1143,1393,832,886]" gridcol="3" gridrow="8" pageId="7" pageNumber="66">filiform</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2AD5122D8FEE" box="[149,1393,926,948]" gridrow="9" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2AD515A38FEE" box="[484,767,926,948]" gridcol="1" gridrow="9" pageId="7" pageNumber="66">Rostrum</td>
<td id="768E5DDFFF97B1AEFCFE2AD5130A8FEE" box="[807,1110,926,948]" gridcol="2" gridrow="9" pageId="7" pageNumber="66">absent</td>
<td id="768E5DDFFF97B1AEFBAE2AD5122D8FEE" box="[1143,1393,926,948]" gridcol="3" gridrow="9" pageId="7" pageNumber="66">present</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2AF6122D8F88" box="[149,1393,957,978]" gridrow="10" pageId="7" pageNumber="66" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="768E5DDFFF97B1AEFF4C2AF616E68F88" box="[149,442,957,978]" gridcol="0" gridrow="10" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C2AF617B48F88" bold="true" box="[149,232,957,978]" pageId="7" pageNumber="66">Anthers</emphasis>
</th>
</tr>
<tr id="355F34A3FF97B1AEFF4C2A97122D8FA8" box="[149,1393,988,1010]" gridrow="11" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2A9715A38FA8" box="[484,767,988,1010]" gridcol="1" gridrow="11" pageId="7" pageNumber="66">Length of thecae sporangia</td>
<td id="768E5DDFFF97B1AEFCFE2A97130A8FA8" box="[807,1110,988,1010]" gridcol="2" gridrow="11" pageId="7" pageNumber="66">unequal</td>
<td id="768E5DDFFF97B1AEFBAE2A97122D8FA8" box="[1143,1393,988,1010]" gridcol="3" gridrow="11" pageId="7" pageNumber="66">equal</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2D51122D886A" box="[149,1393,1050,1072]" gridrow="12" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2D5115A3886A" box="[484,767,1050,1072]" gridcol="1" gridrow="12" pageId="7" pageNumber="66">Position of stigmatic lobe</td>
<td id="768E5DDFFF97B1AEFCFE2D51130A886A" box="[807,1110,1050,1072]" gridcol="2" gridrow="12" pageId="7" pageNumber="66">erect or suberect</td>
<td id="768E5DDFFF97B1AEFBAE2D51122D886A" box="[1143,1393,1050,1072]" gridcol="3" gridrow="12" pageId="7" pageNumber="66">spiraled</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2D72122D8815" box="[149,1393,1081,1103]" gridrow="13" pageId="7" pageNumber="66" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="768E5DDFFF97B1AEFF4C2D7216E68815" box="[149,442,1081,1103]" gridcol="0" gridrow="13" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C2D7217818815" bold="true" box="[149,221,1081,1103]" pageId="7" pageNumber="66">Stigma</emphasis>
</th>
</tr>
<tr id="355F34A3FF97B1AEFF4C2D13122D8834" box="[149,1393,1112,1134]" gridrow="14" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2D1315A38834" box="[484,767,1112,1134]" gridcol="1" gridrow="14" pageId="7" pageNumber="66">Length of stigmatic lobe</td>
<td id="768E5DDFFF97B1AEFCFE2D13130A8834" box="[807,1110,1112,1134]" gridcol="2" gridrow="14" pageId="7" pageNumber="66">1–1.4 mm</td>
<td id="768E5DDFFF97B1AEFBAE2D13122D8834" box="[1143,1393,1112,1134]" gridcol="3" gridrow="14" pageId="7" pageNumber="66">2–2.5 mm</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2DD3122D8887" box="[149,1393,1176,1245]" gridrow="15" pageId="7" pageNumber="66">
<th id="768E5DDFFF97B1AEFF4C2DD316E68887" box="[149,442,1176,1245]" gridcol="0" gridrow="15" pageId="7" pageNumber="66">
<emphasis id="B91BEAF3FF974E56FF4C2D8C17808887" bold="true" box="[149,220,1223,1245]" pageId="7" pageNumber="66">Ovules</emphasis>
</th>
<td id="768E5DDFFF97B1AEFE3D2DD315A38887" box="[484,767,1176,1245]" gridcol="1" gridrow="15" pageId="7" pageNumber="66">Position in relation to the longitudinal axis of the ovary</td>
<td id="768E5DDFFF97B1AEFCFE2DD3130A8887" box="[807,1110,1176,1245]" gridcol="2" gridrow="15" pageId="7" pageNumber="66">suberect</td>
<td id="768E5DDFFF97B1AEFBAE2DD3122D8887" box="[1143,1393,1176,1245]" gridcol="3" gridrow="15" pageId="7" pageNumber="66">patent</td>
</tr>
<tr id="355F34A3FF97B1AEFF4C2DB3122D8954" box="[149,1393,1272,1294]" gridrow="16" pageId="7" pageNumber="66" rowspan-0="1">
<td id="768E5DDFFF97B1AEFE3D2DB315A38954" box="[484,767,1272,1294]" gridcol="1" gridrow="16" pageId="7" pageNumber="66">Disposition</td>
<td id="768E5DDFFF97B1AEFCFE2DB3130A8954" box="[807,1110,1272,1294]" gridcol="2" gridrow="16" pageId="7" pageNumber="66">2 rows per locule</td>
<td id="768E5DDFFF97B1AEFBAE2DB3122D8954" box="[1143,1393,1272,1294]" gridcol="3" gridrow="16" pageId="7" pageNumber="66">2–4 rows per locule</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD036E1FF974E56FF642C01146E8A96" blockId="7.[136,1452,1353,2064]" pageId="7" pageNumber="66">
We hypothesize that the suberect position for the ovule was a response to carpel size reduction in the common ancestor of the 
<taxonomicName id="4C6F4D62FF974E56FEE22C2516FF89DD" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[315,419,1389,1415]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF974E56FEE22C2516FF89DD" box="[315,419,1389,1415]" italics="true" pageId="7" pageNumber="66">D. selloa</emphasis>
</taxonomicName>
complex, if the 
<taxonomicName id="4C6F4D62FF974E56FDBA2C25159789DD" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[611,715,1389,1415]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF974E56FDBA2C25159789DD" box="[611,715,1389,1415]" italics="true" pageId="7" pageNumber="66">D. selloa</emphasis>
</taxonomicName>
complex represents an evolutionary lineage. This reduction and suberect position of the ovule has yet to be widely verified among other lineages of the xeric 
<taxonomicName id="4C6F4D62FF974E56FB532CDA126A89F1" authorityName="Harms" authorityYear="1930" box="[1162,1334,1425,1451]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="subFamily" subFamily="Pitcairnioideae">Pitcairnioideae</taxonomicName>
. However, in the earlier divergent lineages of the xeric 
<taxonomicName id="4C6F4D62FF974E56FD4E2CFE141E8995" authorityName="Harms" authorityYear="1930" box="[663,834,1461,1487]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="subFamily" subFamily="Pitcairnioideae">Pitcairnioideae</taxonomicName>
, such as those that compose 
<taxonomicName id="4C6F4D62FF974E56FB452CFE12748995" box="[1180,1320,1461,1487]" class="Liliopsida" family="Bromeliaceae" genus="Encholirium" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF974E56FB452CFE12748995" box="[1180,1320,1461,1487]" italics="true" pageId="7" pageNumber="66">Encholirium</emphasis>
</taxonomicName>
(having as references the phylogenies of 
<bibRefCitation id="EFFE4B10FF974E56FE052C9115EA89AE" author="Krapp, F. &amp; Pinange, D. S. B. &amp; Benko-Iseppon, A. M. &amp; Leme, E. M. C. &amp; Weising, K." box="[476,694,1497,1524]" pageId="7" pageNumber="66" pagination="1591 - 1614" refId="ref7816" refString="Krapp, F., Pinange, D. S. B., Benko-Iseppon, A. M., Leme, E. M. C. &amp; Weising, K. (2014) Phylogeny and evolution of Dyckia (Bromeliaceae) inferred from chloroplast and nuclear sequences. Plant Systematics and Evolution 300: 1591 - 1614. https: // doi. org / 10.1007 / s 00606 - 014 - 0985 - 0" type="journal article" year="2014">
Krapp 
<emphasis id="B91BEAF3FF974E56FDF32C91153F89A9" box="[554,611,1497,1523]" italics="true" pageId="7" pageNumber="66">et al.</emphasis>
(2014)
</bibRefCitation>
and 
<bibRefCitation id="EFFE4B10FF974E56FD372C9214B489AE" author="Pinange, D. S. B. &amp; Krapp, F. &amp; Zizka, G. &amp; Silvestro, D. &amp; Leme, E. M. C. &amp; Weising, K. &amp; Benko-Iseppon, A. M." box="[750,1000,1497,1524]" pageId="7" pageNumber="66" pagination="39 - 56" refId="ref8531" refString="Pinange, D. S. B., Krapp, F., Zizka, G., Silvestro, D., Leme, E. M. C., Weising, K. &amp; Benko-Iseppon, A. M. (2017) Molecular phylogenetics, historical biogeography and character evolution in Dyckia (Bromeliaceae, Pitcairnioideae). Botanical Journal of the Linnean Society 183: 39 - 56. https: // doi. org / 10.1111 / boj. 12489" type="journal article" year="2017">
Pinangé 
<emphasis id="B91BEAF3FF974E56FC882C9114D689A9" box="[849,906,1497,1523]" italics="true" pageId="7" pageNumber="66">et al.</emphasis>
(2017))
</bibRefCitation>
, the locules of the carpels are relatively large and the position and disposition of ovules of most species are identical to that observed in species of 
<taxonomicName id="4C6F4D62FF974E56FAF22CB612268A4D" box="[1323,1402,1533,1559]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF974E56FAF22CB612268A4D" box="[1323,1402,1533,1559]" italics="true" pageId="7" pageNumber="66">Dyckia</emphasis>
</taxonomicName>
that do not belong to the 
<taxonomicName id="4C6F4D62FF974E56FEA92F6916898A61" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[368,469,1569,1595]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF974E56FEA92F6916898A61" box="[368,469,1569,1595]" italics="true" pageId="7" pageNumber="66">D. selloa</emphasis>
</taxonomicName>
complex. These characters, however, need to be further investigated. A further increase in carpel size during evolution, as in 
<taxonomicName id="4C6F4D62FF974E56FDF72F0D15F78A05" box="[558,683,1605,1631]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF974E56FDF72F0D15F78A05" box="[558,683,1605,1631]" italics="true" pageId="7" pageNumber="66">D. delicata</emphasis>
</taxonomicName>
, could be interpreted as a reversal to the ancestral state of the xeric 
<taxonomicName id="4C6F4D62FF974E56FF512F2216688AD9" authorityName="Harms" authorityYear="1930" box="[136,308,1641,1667]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="subFamily" subFamily="Pitcairnioideae">Pitcairnioideae</taxonomicName>
. With greater availability of locular space, the position of the ovules in relation to the longitudinal axis of the ovary became clear again. Thus, the disposition of ovules in a greater number of rows may have arisen in association with this larger space, as observed in 
<taxonomicName id="4C6F4D62FF974E56FD762FF914778A91" box="[687,811,1713,1739]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF974E56FD762FF914778A91" box="[687,811,1713,1739]" italics="true" pageId="7" pageNumber="66">D. delicata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD036E1FF974E59FF642F9E15B78D7E" blockId="7.[136,1452,1353,2064]" lastBlockId="8.[136,1452,158,725]" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
The ovules of 
<taxonomicName id="4C6F4D62FF974E56FE862F9D15778AB5" authorityName="Büneker &amp; Mariath" authorityYear="2022" box="[351,555,1749,1775]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF974E56FE862F9D15778AB5" box="[351,555,1749,1775]" italics="true" pageId="7" pageNumber="66">D. pseudodelicata</emphasis>
</taxonomicName>
are disposed in two rows per locule, with one row per placental attachment line (
<figureCitation id="13542A64FF974E56FF492FB2166B8B4E" box="[144,311,1785,1812]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="7" pageNumber="66">Fig. 3G and H</figureCitation>
), as is the case for most of the species of 
<taxonomicName id="4C6F4D62FF974E56FCC92FB214038B49" box="[784,863,1785,1811]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF974E56FCC92FB214038B49" box="[784,863,1785,1811]" italics="true" pageId="7" pageNumber="66">Dyckia</emphasis>
</taxonomicName>
studied to date. On the other hand, 
<taxonomicName id="4C6F4D62FF974E56FB2F2FB1122F8B49" box="[1270,1395,1785,1811]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF974E56FB2F2FB1122F8B49" box="[1270,1395,1785,1811]" italics="true" pageId="7" pageNumber="66">D. delicata</emphasis>
</taxonomicName>
may have ovules disposed in 2–4 rows per locule, with one or two rows per placental attachment line (
<figureCitation id="13542A64FF974E56FB2A2E5612C28B62" box="[1267,1438,1821,1848]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="7" pageNumber="66">Fig. 3N and O</figureCitation>
). Analysis of Endress’s (2011) review of the diversity, development and evolution of angiosperm ovules reveals that both ovule position in relation to the longitudinal axis of the ovary and ovule arrangement (number of rows) seem to be related to the proportion of available locular space and ovule size, ovule number and direction of ovule initiation sequence in placentae. 
<bibRefCitation id="EFFE4B10FF974E56FE5C2EE6150C8B92" author="Kuhn, S. A. &amp; Nogueira, F. M. &amp; Chauveau, O. &amp; Mariath, J. E." box="[389,592,1965,1992]" pageId="7" pageNumber="66" pagination="151691" refId="ref7895" refString="Kuhn, S. A., Nogueira, F. M., Chauveau, O. &amp; Mariath, J. E. deA. (2020) Diversity and evolutionary trends of the ovary and ovule in Bromeliaceae. Flora 272: 151691. https: // doi. org / 10.1016 / j. flora. 2020.151691" type="journal article" year="2020">
Kuhn 
<emphasis id="B91BEAF3FF974E56FE102EE5155C8B9D" box="[457,512,1965,1991]" italics="true" pageId="7" pageNumber="66">et al.</emphasis>
(2020)
</bibRefCitation>
reported that ovules arranged in a few rows per placental attachment line is an ancestral character in 
<taxonomicName id="4C6F4D62FF974E56FEA22E9A15458BB1" box="[379,537,2001,2027]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
. Transitions toward many ovule rows occurred at least twice at the base of a clade, including all genera of 
<taxonomicName id="4C6F4D62FF974E56FE5E2EBE156A8455" authorityName="Burnett" authorityYear="1835" box="[391,566,2037,2063]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="subFamily" subFamily="Tillandsioideae">Tillandsioideae</taxonomicName>
, except 
<taxonomicName id="4C6F4D62FF974E56FD562EBD15AD844A" box="[655,753,2038,2064]" class="Liliopsida" family="Bromeliaceae" genus="Catopsis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF974E56FD562EBD15AD844A" box="[655,753,2038,2064]" italics="true" pageId="7" pageNumber="66">Catopsis</emphasis>
</taxonomicName>
, on the one hand, and among Puyoideae and 
<taxonomicName id="4C6F4D62FF974E56FB3D2EBE12D58455" authorityName="Burnett" authorityYear="1835" box="[1252,1417,2037,2063]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="66" phylum="Tracheophyta" rank="subFamily" subFamily="Bromelioideae">Bromelioideae</taxonomicName>
on the other hand. These transitions are followed by at least three reversals in 
<taxonomicName id="4C6F4D62FF984E59FC1329D5132A8CE2" authorityName="Burnett" authorityYear="1835" box="[970,1142,158,184]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="subFamily" subFamily="Tillandsioideae">Tillandsioideae</taxonomicName>
and one, but perhaps more, in 
<taxonomicName id="4C6F4D62FF984E59FF7D298916118C86" authorityName="Burnett" authorityYear="1835" box="[164,333,194,220]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="subFamily" subFamily="Bromelioideae">Bromelioideae</taxonomicName>
.A transition from many towards lots ovule rows per each placental attachment line is observed at the base of the 
<taxonomicName id="4C6F4D62FF984E59FED029AD16298D5A" authorityName="J.R.Grant" authorityYear="1995" box="[265,373,230,256]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF984E59FED029AD16298D5A" box="[265,373,230,256]" italics="true" pageId="8" pageNumber="67">Werauhia</emphasis>
</taxonomicName>
clade within 
<taxonomicName id="4C6F4D62FF984E59FDD429AD15E08D5A" authorityName="Burnett" authorityYear="1835" box="[525,700,230,256]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="subFamily" subFamily="Tillandsioideae">Tillandsioideae</taxonomicName>
, while another transition from few to many ovule rows occurred at the base of the 
<taxonomicName id="4C6F4D62FF984E59FEEB284016F88D7E" box="[306,420,267,292]" class="Liliopsida" family="Bromeliaceae" genus="Pitcairnia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF984E59FEEB284016F88D7E" box="[306,420,267,292]" italics="true" pageId="8" pageNumber="67">Pitcairnia</emphasis>
</taxonomicName>
clade within 
<taxonomicName id="4C6F4D62FF984E59FDE2284115BB8D7E" authorityName="Harms" authorityYear="1930" box="[571,743,266,292]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="subFamily" subFamily="Pitcairnioideae">Pitcairnioideae</taxonomicName>
.
</paragraph>
<paragraph id="8BD036E1FF984E59FF642865157B8ED6" blockId="8.[136,1452,158,725]" pageId="8" pageNumber="67">
Despite these differences in the position and disposition of ovules between 
<taxonomicName id="4C6F4D62FF984E59FBC7286413C08D12" box="[1054,1180,302,328]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis id="B91BEAF3FF984E59FBC7286413C08D12" box="[1054,1180,302,328]" italics="true" pageId="8" pageNumber="67">D. delicata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6F4D62FF984E59FB0E286412F98D12" authorityName="Büneker &amp; Mariath" authorityYear="2022" box="[1239,1445,302,328]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis id="B91BEAF3FF984E59FB0E286412F98D12" box="[1239,1445,302,328]" italics="true" pageId="8" pageNumber="67">D. pseudodelicata</emphasis>
</taxonomicName>
, their seeds are morphologically similar (
<figureCitation id="13542A64FF984E59FD8A281915B78D37" box="[595,747,338,365]" captionStart="FIGURE 3" captionStartId="5.[136,229,1819,1841]" captionTargetBox="[183,1404,281,1794]" captionTargetId="figure-72@5.[183,1404,281,1794]" captionTargetPageId="5" captionText="FIGURE 3. A–I. Dyckia pseudodelicata (Büneker 913 et al.), J–O. Dyckia delicata (Büneker 395 et al.). A–F and J–M. Petal margin. A. Trichomes of the proximal margin. B. Trichomes of median margin. C. Trichomes of the distal margin. D. Dendritic trichome on DIC. E. Flabelliform trichome on DIC. F. Uniseriate-capitate trichome (left) and flabelliform trichome (right) on DIC. J. Emergences of the distal margin. K. Unicellular trichomes of the apical region. L. Emergences on DIC. M. Unicellular filiform trichomes on DIC. G and N. Transverse section of the median region of ovary. H and O. Detail of transverse section of median region of ovary, where two rows of ovules can be seen in H and four in O. I and P. Seed. E, G, H, L, M, N and O. Stained with Safranin. D and F. Stained with Toluidine Blue." figureDoi="http://doi.org/10.5281/zenodo.6630640" httpUri="https://zenodo.org/record/6630640/files/figure.png" pageId="8" pageNumber="67">Fig. 3I and P</figureCitation>
), being oblanceoloid as for other species of 
<taxonomicName id="4C6F4D62FF984E59FB3D2819126F8D36" box="[1252,1331,338,364]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF984E59FB3D2819126F8D36" box="[1252,1331,338,364]" italics="true" pageId="8" pageNumber="67">Dyckia</emphasis>
</taxonomicName>
belonging to the complex and possessing a chalazal appendix which constitutes a poorly developed wing (
<emphasis id="B91BEAF3FF984E59FB0E283C13AA8DCA" box="[1239,1270,375,400]" italics="true" pageId="8" pageNumber="67">vs.</emphasis>
well developed for the rest of 
<taxonomicName id="4C6F4D62FF984E59FEFE28D1162A8DEE" box="[295,374,410,436]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B91BEAF3FF984E59FEFE28D1162A8DEE" box="[295,374,410,436]" italics="true" pageId="8" pageNumber="67">Dyckia</emphasis>
</taxonomicName>
). The anatomy, number, position in relation to the longitudinal axis of the ovary and disposition of ovules are characters to be investigated more widely in the xeric 
<taxonomicName id="4C6F4D62FF984E59FC5628F5136B8D82" authorityName="Harms" authorityYear="1930" box="[911,1079,446,472]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="subFamily" subFamily="Pitcairnioideae">Pitcairnioideae</taxonomicName>
as they seem to have taxonomic importance as diagnostic characters and may be directly related to the unique morphology of the seeds of the 
<taxonomicName id="4C6F4D62FF984E59FA4928A817978E7A" authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis id="B91BEAF3FF984E59FA4928A817978E7A" italics="true" pageId="8" pageNumber="67">D. selloa</emphasis>
</taxonomicName>
complex. They are oblanceoloid with a poorly developed chalazal appendix (
<emphasis id="B91BEAF3FF984E59FBE82B4C13148E7A" box="[1073,1096,519,544]" italics="true" pageId="8" pageNumber="67">vs</emphasis>
. flated with developed chalazal appendix forming a subfalciform wing in most 
<emphasis id="B91BEAF3FF984E59FD452B61144F8E1E" box="[668,787,554,580]" italics="true" pageId="8" pageNumber="67">
<taxonomicName id="4C6F4D62FF984E59FD452B6115B78E1E" box="[668,747,554,580]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="genus">Dyckia</taxonomicName>
l.s.
</emphasis>
), a characteristic that appears to be a synapomorphy of the complex. Furthermore, the study of embryological characters can be an important tool for evolutionary analysis and help in phylogenetic reconstructions.
</paragraph>
</subSubSection>
<subSubSection id="C375656AFF984E59FF642BDD126A8E8E" pageId="8" pageNumber="67" type="materials_examined">
<paragraph id="8BD036E1FF984E59FF642BDD126A8E8E" blockId="8.[136,1452,158,725]" pageId="8" pageNumber="67">
<emphasis id="B91BEAF3FF984E59FF642BDD15BE8EEA" bold="true" box="[189,738,662,688]" pageId="8" pageNumber="67">
<collectingRegion id="49ABF803FF984E59FF642BDD156B8EEA" box="[189,567,662,688]" pageId="8" pageNumber="67">Additional Specimen Examined</collectingRegion>
(
<typeStatus id="54D48843FF984E59FD9F2BDD15EA8EEA" box="[582,694,662,688]" pageId="8" pageNumber="67" type="paratype">paratype</typeStatus>
):—
</emphasis>
<materialsCitation id="3B073CBCFF984E59FD3B2BDD126E8E8E" ID-GBIF-Occurrence="3807302301" collectingDate="2017-08-03" collectingDateMax="2020-01" collectingDateMin="2017-08-03" collectionCode="H" country="Brazil" isTailEnumeration="true" location="Toropi-mirim" pageId="8" pageNumber="67" specimenCount="1" stateProvince="Rio Grande do Sul">
<collectingCountry id="F3787671FF984E59FD3B2BDD14178EEA" box="[738,843,662,688]" name="Brazil" pageId="8" pageNumber="67">BRAZIL</collectingCountry>
. 
<collectingRegion id="49ABF803FF984E59FC8C2BDD13768EEA" box="[853,1066,662,688]" country="Brazil" name="Rio Grande do Sul" pageId="8" pageNumber="67">Rio Grande do Sul</collectingRegion>
: 
<emphasis id="B91BEAF3FF984E59FBEC2BDD13398EEA" bold="true" box="[1077,1125,662,688]" pageId="8" pageNumber="67">Jari</emphasis>
, encosta rochosa próxima ao arroio 
<location id="8EB0603AFF984E59FF082BF116308E8E" LSID="urn:lsid:plazi:treatment:03C687F7FF914E59FF512E5F126A8E8E:8EB0603AFF984E59FF082BF116308E8E" box="[209,364,698,724]" country="Brazil" name="Toropi-mirim" pageId="8" pageNumber="67" stateProvince="Rio Grande do Sul">Toropi-mirim</location>
, 
<date id="FFD11021FF984E59FEAE2BF115428E8E" box="[375,542,698,724]" pageId="8" pageNumber="67" value="2017-08-03">
<collectingDate id="EF95E9C9FF984E59FEAE2BF115428E8E" box="[375,542,698,724]" pageId="8" pageNumber="67" value="2017-08-03">3 August 2017</collectingDate>
</date>
, floresceu em cultivo em 
<date id="FFD11021FF984E59FCE72BF0148B8E8E" box="[830,983,698,725]" pageId="8" pageNumber="67" value="2020-01">
<collectingDate id="EF95E9C9FF984E59FCE72BF0148B8E8E" box="[830,983,698,725]" pageId="8" pageNumber="67" value="2020-01">January 2020</collectingDate>
</date>
, 
<emphasis id="B91BEAF3FF984E59FC3B2BF013AC8E8E" box="[994,1264,698,724]" italics="true" pageId="8" pageNumber="67">
<collectionCode id="ED7EAE24FF984E59FC3B2BF014A68E8E" box="[994,1018,699,724]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="8" pageNumber="67" type="Herbarium">H</collectionCode>
. 
<collectionCode id="ED7EAE24FF984E59FC242BF013448E8E" box="[1021,1048,699,724]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="8" pageNumber="67" type="Herbarium">M</collectionCode>
. 
<location id="8EB0603AFF984E59FBFB2BF113238E8E" LSID="urn:lsid:plazi:treatment:03C687F7FF914E59FF512E5F126A8E8E:8EB0603AFF984E59FBFB2BF113238E8E" box="[1058,1151,698,724]" country="Brazil" name="Buneker" pageId="8" pageNumber="67" stateProvince="Jari">Büneker</location>
635 et al.
</emphasis>
(
<collectionCode id="ED7EAE24FF984E59FB262BF112758E8E" box="[1279,1321,698,724]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="8" pageNumber="67" type="Herbarium">RB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>