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<document id="69E04D6089AA2D11DE6E913AE9258D7F" ID-DOI="10.5281/zenodo.204914" ID-GBIF-Dataset="035006f1-9010-4f0b-a5fb-83fed142d245" ID-ISSN="1175-5326" ID-Zenodo-Dep="204914" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460221844229" checkinUser="plazi" docAuthor="Ciccotto, Patrick J., Konings, Adrianus &amp; Stauffer, Jay R." docDate="2011" docId="03BB942A0D6483749DEFFD9EFB63FD59" docLanguage="en" docName="zt02738p025.pdf" docOrigin="Zootaxa 2738" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Metriaclima nkhunguensis Ciccotto, Konings &amp; Stauffer, 2011, new species" docType="treatment" docVersion="11" lastPageNumber="19" masterDocId="FF82EC520D7483669D78FF83FFCBFFFE" masterDocTitle="Descriptions of five new species in the genus Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa" masterLastPageNumber="25" masterPageNumber="1" pageNumber="17" updateTime="1698237790047" updateUser="plazi">
<mods:mods id="57D858FEAB7C6CCAA1621445557E0A4A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EA6041651B26F73EE1E6035C969918C3">Descriptions of five new species in the genus Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa</mods:title>
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<mods:name id="4C9F6C131A94BDF51F3062E90642C2FC" type="personal">
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<mods:namePart id="6577BC92D9DDBE3739F6BF1A184441E1">Ciccotto, Patrick J.</mods:namePart>
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<mods:namePart id="4161BF29EF9D18FDBD6CE242310BD564">Konings, Adrianus</mods:namePart>
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<mods:namePart id="3F27C132BE66235EC817DE28D255D434">Stauffer, Jay R.</mods:namePart>
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<mods:typeOfResource id="6F7B3FBCA978D49F15935512ED2DA0E4">text</mods:typeOfResource>
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<mods:title id="7C976DBCD302B265B324019D1D536CA3">Zootaxa</mods:title>
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<mods:date id="29005450AAEBAD3C62870DB8C6D22ED8">2011</mods:date>
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<mods:number id="9758F0F24EB20DBBCFD9561E7D4AA7F3">2738</mods:number>
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<mods:identifier id="3D6668BCD4F1D3F790B2E918098E5ACD" type="DOI">10.5281/zenodo.204914</mods:identifier>
<mods:identifier id="F4CE1D9DE036B45079814EE88A473C1C" type="GBIF-Dataset">035006f1-9010-4f0b-a5fb-83fed142d245</mods:identifier>
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<mods:identifier id="C0671CF2AB90CC655F4093CD345A62E1" type="Zenodo-Dep">204914</mods:identifier>
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<treatment id="03BB942A0D6483749DEFFD9EFB63FD59" ID-DOI="http://doi.org/10.5281/zenodo.6192258" ID-GBIF-Taxon="119385420" ID-Zenodo-Dep="6192258" LSID="urn:lsid:zoobank.org:act:274F4DA4-707C-48B7-BF32-310F5E6BBE96" httpUri="http://treatment.plazi.org/id/03BB942A0D6483749DEFFD9EFB63FD59" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C30876B70D6483769DEFFD9EFEA4FDA9" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BAD253C0D6483769DEFFD9EFDB7FDC9" blockId="16.[151,636,541,599]" box="[151,636,541,567]" pageId="16" pageNumber="17">
<heading id="D0E592500D6483769DEFFD9EFDB7FDC9" bold="true" box="[151,636,541,567]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B966F92E0D6483769DEFFD9EFDB7FDC9" bold="true" box="[151,636,541,567]" pageId="16" pageNumber="17">
<taxonomicName id="4C125EBF0D6483769DEFFD9EFE16FDC9" ID-CoL="42HWM" box="[151,477,541,567]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nkhunguensis" status="sp. nov.">
<emphasis id="B966F92E0D6483769DEFFD9EFE16FDC9" bold="true" box="[151,477,541,567]" italics="true" pageId="16" pageNumber="17">Metriaclima nkhunguensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25544550D6483769C94FD9EFDB7FDC9" box="[492,636,541,567]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD253C0D6483769DEFFDBCFEA4FDA9" blockId="16.[151,636,541,599]" box="[151,367,575,599]" pageId="16" pageNumber="17">
<figureCitation id="132939B90D6483769DEFFDBCFF26FDA9" box="[151,237,575,599]" captionStart="FIGURE 8. A" captionStartId="16.[151,250,1943,1966]" captionTargetBox="[276,1311,1188,1922]" captionTargetId="figure@16.[276,1312,1151,1922]" captionTargetPageId="16" captionText="FIGURE 8. A. Metriaclima nkhunguensis, holotype, PSU 4603, 72.8 mm SL; Mozambique: Lake MalaŴi: Nkhungu Reef. B. M. nkhunguensis, male in habitat at Nkhungu Reef, Mozambique; approx. 60 mm SL; not preserved. C. M. nkhunguensis, female in habitat at Nkhungu Reef, Mozambique; not preserved." httpUri="https://zenodo.org/record/204922/files/figure.png" pageId="16" pageNumber="17">Fig. 8A</figureCitation>
–C;
<tableCitation id="C69010870D6483769C64FDBCFEA4FDA9" box="[284,367,575,599]" captionStart="TABLE 6" captionStartId="17.[151,235,152,173]" captionTargetBox="[151,1436,314,1990]" captionTargetPageId="17" captionText="TABLE 6. Morphometric and meristic values of Metriaclima nkhunguensis from Nkhungu Reef. Mean and range include holotype." httpUri="http://table.plazi.org/id/DF6D75B40D6583779DEFFF1BFF02FF32" pageId="16" pageNumber="17" tableUuid="DF6D75B40D6583779DEFFF1BFF02FF32">Table 6</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D6483769DEFFD0BFDBEFD60" box="[151,629,648,670]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BAD253C0D6483769DEFFD0BFDBEFD60" blockId="16.[151,629,648,670]" box="[151,629,648,670]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB0202120D6483769DEFFD0BFDBEFD60" box="[151,629,648,670]" pageId="16" pageNumber="17">
<taxonomicName id="4C125EBF0D6483769DEFFD0BFEDFFD60" box="[151,276,648,670]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D6483769DEFFD0BFEDFFD60" box="[151,276,648,670]" italics="true" pageId="16" pageNumber="17">Metriaclima</emphasis>
</taxonomicName>
sp. &quot;blue reef&quot;:
<treatmentCitation id="0AB3032D0D6483769CC1FD0BFDBAFD60" author="Konings" box="[441,625,648,670]" page="58" pageId="16" pageNumber="17" year="2001">
<bibRefCitation id="EF8358CD0D6483769CC1FD0BFD80FD63" author="Konings" box="[441,587,648,670]" pageId="16" pageNumber="25" refString="Konings, A. (2001) Mala w i cichlids in their natural habitat, Third edition. Cichlid Press, El Paso, Texas, 352 pp." type="book" year="2001">Konings 2001</bibRefCitation>
: 58
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D6483769DEFFD48FC63FCD4" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BAD253C0D6483769DEFFD48FDF7FCF8" blockId="16.[151,1437,714,1135]" pageId="16" pageNumber="17">
<emphasis id="B966F92E0D6483769DEFFD48FEC0FD1D" bold="true" box="[151,267,715,739]" pageId="16" pageNumber="17">
<typeStatus id="54A99B9E0D6483769DEFFD48FECDFD1D" box="[151,262,715,739]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED03BDF90D6483769C69FD49FE8FFD1C" box="[273,324,714,738]" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="16" pageNumber="17">PSU</collectionCode>
4603,
<quantity id="4CEA88D90D6483769CE9FD49FE3EFD1C" box="[401,501,714,738]" metricMagnitude="-2" metricUnit="m" metricValue="7.279999999999999" pageId="16" pageNumber="17" unit="mm" value="72.8">72.8 mm</quantity>
SL;
<collectingCountry id="F30565AC0D6483769F53FD49FD08FD1C" box="[555,707,714,738]" name="Mozambique" pageId="16" pageNumber="17">Mozambique</collectingCountry>
: Lake
<collectingCountry id="F30565AC0D6483769E74FD49FCADFD1D" box="[780,870,714,739]" name="Malawi" pageId="16" pageNumber="17">Malaŵi</collectingCountry>
: Nkhungu Reef,
<geoCoordinate id="EE2643FB0D6483769959FD49FB72FD1C" box="[1057,1209,714,738]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="-12.980017">12° 58.801' S</geoCoordinate>
,
<geoCoordinate id="EE2643FB0D64837699BDFD49FAABFD1C" box="[1221,1376,714,738]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="34.76395">34° 45.837' E</geoCoordinate>
; A.F. Konings &amp; J.R. Stauffer,
<date id="FFAC03FC0D6483769CCAFD6DFDF2FCF8" box="[434,569,750,774]" pageId="16" pageNumber="17" value="2006-03-01">1 Mar 2006</date>
.
</paragraph>
<paragraph id="8BAD253C0D6483769DBFFC90FC63FCD4" blockId="16.[151,1437,714,1135]" box="[199,936,786,811]" pageId="16" pageNumber="17">
<emphasis id="B966F92E0D6483769DBFFC90FE8DFCD5" bold="true" box="[199,326,787,811]" pageId="16" pageNumber="17">
<typeStatus id="54A99B9E0D6483769DBFFC90FE8AFCD5" box="[199,321,787,811]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED03BDF90D6483769C34FC91FEB4FCD4" box="[332,383,786,810]" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="16" pageNumber="17">PSU</collectionCode>
4604, 5;
<quantity id="4CEA88D90D6483769C92FC91FD47FCD4" box="[490,652,786,810]" metricMagnitude="-2" metricUnit="m" metricValue="6.93" metricValueMax="7.33" metricValueMin="6.529999999999999" pageId="16" pageNumber="17" unit="mm" value="69.3" valueMax="73.3" valueMin="65.3">65.3–73.3 mm</quantity>
SL; data as for
<typeStatus id="54A99B9E0D6483769E46FC91FC6FFCD4" box="[830,932,786,810]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D6483779DBFFCB5FD66FED9" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BAD253C0D6483769DBFFCB5FCD8FB90" blockId="16.[151,1437,714,1135]" pageId="16" pageNumber="17">
<emphasis id="B966F92E0D6483769DBFFCB5FE8AFCB1" bold="true" box="[199,321,822,847]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
A moderately-sloped head and bicuspid teeth in the outer row of the jaws place this species in
<taxonomicName id="4C125EBF0D6483769DEFFCD8FEEAFC8D" box="[151,289,859,883]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D6483769DEFFCD8FEEAFC8D" box="[151,289,859,883]" italics="true" pageId="16" pageNumber="17">Metriaclima</emphasis>
</taxonomicName>
. Both males and females are light blue, fading to white ventrally, distinguishing them from all other members of
<taxonomicName id="4C125EBF0D6483769C5BFCFDFE66FC68" box="[291,429,894,918]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D6483769C5BFCFDFE66FC68" box="[291,429,894,918]" italics="true" pageId="16" pageNumber="17">Metriaclima</emphasis>
</taxonomicName>
, except
<taxonomicName id="4C125EBF0D6483769F74FC03FD53FC68" box="[524,664,894,919]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="callainos">
<emphasis id="B966F92E0D6483769F74FC03FD53FC68" box="[524,664,894,919]" italics="true" pageId="16" pageNumber="17">M. callainos</emphasis>
</taxonomicName>
. Male
<taxonomicName id="4C125EBF0D6483769F9EFC03FC6FFC68" box="[742,932,894,919]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6483769F9EFC03FC6FFC68" box="[742,932,894,919]" italics="true" pageId="16" pageNumber="17">M. nkhunguensis</emphasis>
</taxonomicName>
are distinguished from those of the similarly colored
<taxonomicName id="4C125EBF0D6483769D8CFC27FE7BFC45" box="[244,432,931,955]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chrysomallos">
<emphasis id="B966F92E0D6483769D8CFC27FE7BFC45" box="[244,432,931,955]" italics="true" pageId="16" pageNumber="17">M. chrysomallos</emphasis>
</taxonomicName>
from the Gome population,
<taxonomicName id="4C125EBF0D6483769F81FC27FCB4FC45" box="[761,895,931,955]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="estherae">
<emphasis id="B966F92E0D6483769F81FC27FCB4FC45" box="[761,895,931,955]" italics="true" pageId="16" pageNumber="17">M. estherae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C125EBF0D6483769EB9FC27FB84FC45" box="[961,1103,931,955]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="callainos">
<emphasis id="B966F92E0D6483769EB9FC27FB84FC45" box="[961,1103,931,955]" italics="true" pageId="16" pageNumber="17">M. callainos</emphasis>
</taxonomicName>
, the latter two species of the sediment-free rocky habitat, by the yellow base of the pectoral fin (vs. no yellow pigment in males of the other three species). Female
<taxonomicName id="4C125EBF0D6483769CE1FC6FFD9CFBFD" box="[409,599,1003,1027]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6483769CE1FC6FFD9CFBFD" box="[409,599,1003,1027]" italics="true" pageId="16" pageNumber="17">M. nkhunguensis</emphasis>
</taxonomicName>
are light blue, lighter than those of the only other known species with allblue females,
<taxonomicName id="4C125EBF0D6483769C40FB93FE0CFBD8" box="[312,455,1038,1063]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="callainos">
<emphasis id="B966F92E0D6483769C40FB93FE0CFBD8" box="[312,455,1038,1063]" italics="true" pageId="16" pageNumber="17">M. callainos</emphasis>
</taxonomicName>
, which are a darker cobalt-blue (female
<taxonomicName id="4C125EBF0D6483769EE5FB93FBEFFBD8" box="[925,1060,1038,1063]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="estherae">
<emphasis id="B966F92E0D6483769EE5FB93FBEFFBD8" box="[925,1060,1038,1063]" italics="true" pageId="16" pageNumber="17">M. estherae</emphasis>
</taxonomicName>
are either brown or orange/red).
<taxonomicName id="4C125EBF0D6483769DEFFBB0FE0FFBB5" box="[151,452,1075,1099]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6483769DEFFBB0FE0FFBB5" box="[151,452,1075,1099]" italics="true" pageId="16" pageNumber="17">Metriaclima nkhunguensis</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName id="4C125EBF0D6483769E59FBB7FC7AFBB5" box="[801,945,1075,1099]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="callainos">
<emphasis id="B966F92E0D6483769E59FBB7FC7AFBB5" box="[801,945,1075,1099]" italics="true" pageId="16" pageNumber="17">M. callainos</emphasis>
</taxonomicName>
by a shorter lower jaw (mean 31.2, range 29.4–34.1 % HL vs. mean 37.5, range 34.9–43.3 % HL).
</paragraph>
<caption id="DF6D75B40D6483769DEFF814FCEFF815" httpUri="https://zenodo.org/record/204922/files/figure.png" pageId="16" pageNumber="17" targetBox="[276,1311,1188,1922]" targetPageId="16">
<paragraph id="8BAD253C0D6483769DEFF814FCEFF815" blockId="16.[151,1435,1943,2027]" pageId="16" pageNumber="17">
<emphasis id="B966F92E0D6483769DEFF814FEF8F853" bold="true" box="[151,307,1943,1966]" pageId="16" pageNumber="17">FIGURE 8. A.</emphasis>
<taxonomicName id="4C125EBF0D6483769C41F81BFD8CF850" box="[313,583,1944,1966]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6483769C41F81BFD8CF850" box="[313,583,1944,1966]" italics="true" pageId="16" pageNumber="17">Metriaclima nkhunguensis</emphasis>
</taxonomicName>
, holotype, PSU 4603, 72.8 mm SL; Mozambique: Lake MalaŴi: Nkhungu Reef.
<emphasis id="B966F92E0D64837698FCF81BFA50F853" bold="true" box="[1412,1435,1944,1965]" pageId="16" pageNumber="17">B.</emphasis>
<taxonomicName id="4C125EBF0D6483769DEFF834FE8EF832" box="[151,325,1974,1996]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6483769DEFF834FE8EF832" box="[151,325,1974,1996]" italics="true" pageId="16" pageNumber="17">M. nkhunguensis</emphasis>
</taxonomicName>
, male in habitat at Nkhungu Reef, Mozambique; approx. 60 mm SL; not preserved.
<emphasis id="B966F92E0D64837699BDF835FB15F833" bold="true" box="[1221,1246,1974,1997]" pageId="16" pageNumber="17">C.</emphasis>
<taxonomicName id="4C125EBF0D648376999FF834FA5EF832" box="[1255,1429,1974,1996]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D648376999FF834FA5EF832" box="[1255,1429,1974,1996]" italics="true" pageId="16" pageNumber="17">M. nkhunguensis</emphasis>
</taxonomicName>
, female in habitat at Nkhungu Reef, Mozambique; not preserved.
</paragraph>
</caption>
<caption id="DF6D75B40D6583779DEFFF1BFF02FF32" ID-Table-UUID="DF6D75B40D6583779DEFFF1BFF02FF32" httpUri="http://table.plazi.org/id/DF6D75B40D6583779DEFFF1BFF02FF32" pageId="17" pageNumber="18" targetBox="[151,1436,314,1990]" targetIsTable="true" targetPageId="17">
<paragraph id="8BAD253C0D6583779DEFFF1BFF02FF32" blockId="17.[151,1436,152,295]" pageId="17" pageNumber="18">
<emphasis id="B966F92E0D6583779DEFFF1BFEC8FF53" bold="true" box="[151,259,152,173]" pageId="17" pageNumber="18">TABLE 6.</emphasis>
Morphometric and meristic values of
<taxonomicName id="4C125EBF0D6583779FFCFF1BFC59FF50" box="[644,914,152,174]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6583779FFCFF1BFC59FF50" box="[644,914,152,174]" italics="true" pageId="17" pageNumber="18">Metriaclima nkhunguensis</emphasis>
</taxonomicName>
from Nkhungu Reef. Mean and range include holotype.
</paragraph>
</caption>
<paragraph id="8BAD253C0D6583779FFDFF6BFC58FF00" blockId="17.[151,1436,152,295]" box="[645,915,232,254]" pageId="17" pageNumber="18">
<taxonomicName id="4C125EBF0D6583779FFDFF6BFC58FF00" box="[645,915,232,254]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6583779FFDFF6BFC58FF00" box="[645,915,232,254]" italics="true" pageId="17" pageNumber="18">Metriaclima nkhunguensis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD253C0D6583779FFDFE92FD66FED9" blockId="17.[151,1436,152,295]" box="[645,685,273,295]" pageId="17" pageNumber="18">n=6</paragraph>
</subSubSection>
<paragraph id="8BAD253C0D6583779DE7FEE0FB5EF83B" pageId="17" pageNumber="18">
<table id="F912D79C0D657C999DEFFEB9FA57F838" box="[151,1436,314,1990]" gridcols="4" gridrows="10" pageId="17" pageNumber="18">
<tr id="3522277E0D657C999DEFFEB9FA57FE32" box="[151,1436,314,460]" gridrow="0" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFEB9FDC2FE32" box="[151,521,314,460]" gridcol="0" gridrow="0" pageId="17" pageNumber="18">Standard length, mm Head length, mm Percentage of standard length</th>
<th id="76F34E020D657C999FFDFEB9FD16FE32" box="[645,733,314,460]" gridcol="1" gridrow="0" pageId="17" pageNumber="18">holotype 72.8 22.3</th>
<th id="76F34E020D657C999EE1FEB9FC13FE32" box="[921,984,314,460]" gridcol="2" gridrow="0" pageId="17" pageNumber="18">mean 69.6 20.9</th>
<th id="76F34E020D657C999916FEB9FA57FE32" box="[1134,1436,314,460]" gridcol="3" gridrow="0" pageId="17" pageNumber="18">range 65.3–73.3 19.8–22.3</th>
</tr>
<tr id="3522277E0D657C999DEFFE63FA57FD8F" box="[151,1436,480,625]" gridrow="1" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFE63FDC2FD8F" box="[151,521,480,625]" gridcol="0" gridrow="1" pageId="17" pageNumber="18">Head length Snout to dorsal Snout to pelvic Greatest body depth</th>
<td id="76F34E020D657C999FFDFE63FD16FD8F" box="[645,733,480,625]" gridcol="1" gridrow="1" pageId="17" pageNumber="18">30.6 29.8 38.1 31.9</td>
<td id="76F34E020D657C999EE1FE63FC13FD8F" box="[921,984,480,625]" gridcol="2" gridrow="1" pageId="17" pageNumber="18">30.1 31.1 37.5 32.9</td>
<td id="76F34E020D657C999916FE63FA57FD8F" box="[1134,1436,480,625]" gridcol="3" gridrow="1" pageId="17" pageNumber="18">29.1–31.0 29.8–33.1 36.5–38.1 31.6–34.1</td>
</tr>
<tr id="3522277E0D657C999DEFFD06FA57FD3A" box="[151,1436,645,708]" gridrow="2" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFD06FDC2FD3A" box="[151,521,645,708]" gridcol="0" gridrow="2" pageId="17" pageNumber="18">Caudal peduncle length Least caudal peduncle depth</th>
<td id="76F34E020D657C999FFDFD06FD16FD3A" box="[645,733,645,708]" gridcol="1" gridrow="2" pageId="17" pageNumber="18">14.3 12.3</td>
<td id="76F34E020D657C999EE1FD06FC13FD3A" box="[921,984,645,708]" gridcol="2" gridrow="2" pageId="17" pageNumber="18">14.0 12.2</td>
<td id="76F34E020D657C999916FD06FA57FD3A" box="[1134,1436,645,708]" gridcol="3" gridrow="2" pageId="17" pageNumber="18">12.9–15.3 11.7–12.7</td>
</tr>
<tr id="3522277E0D657C999DEFFD5BFA57FC42" box="[151,1436,728,956]" gridrow="3" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFD5BFDC2FC42" box="[151,521,728,956]" gridcol="0" gridrow="3" pageId="17" pageNumber="18">Dorsal-fin base length Anterior dorsal to anterior anal Anterior dorsal to posterior anal Posterior dorsal to anterior anal Posterior dorsal to posterior anal Anterior dorsal to pelvic-fin origin</th>
<td id="76F34E020D657C999FFDFD5BFD16FC42" box="[645,733,728,956]" gridcol="1" gridrow="3" pageId="17" pageNumber="18">62.5 53.7 64.9 29.5 15.9 36.2</td>
<td id="76F34E020D657C999EE1FD5BFC13FC42" box="[921,984,728,956]" gridcol="2" gridrow="3" pageId="17" pageNumber="18">61.4 52.6 63.7 29.6 15.2 36.1</td>
<td id="76F34E020D657C999916FD5BFA57FC42" box="[1134,1436,728,956]" gridcol="3" gridrow="3" pageId="17" pageNumber="18">58.5–62.6 51.5–53.7 62.1–64.9 28.0–32.0 14.1–16.4 34.6–37.5</td>
</tr>
<tr id="3522277E0D657C999DEFFC53FA57FB4A" box="[151,1436,976,1204]" gridrow="4" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFC53FDC2FB4A" box="[151,521,976,1204]" gridcol="0" gridrow="4" pageId="17" pageNumber="18">Posterior dorsal to pelvic-fin origin Percentage of head length Horizontal eye diameter Vertical eye diameter Snout length Postorbital head length</th>
<td id="76F34E020D657C999FFDFC53FD16FB4A" box="[645,733,976,1204]" gridcol="1" gridrow="4" pageId="17" pageNumber="18">59.8 32.6 33.1 29.4 39.4</td>
<td id="76F34E020D657C999EE1FC53FC13FB4A" box="[921,984,976,1204]" gridcol="2" gridrow="4" pageId="17" pageNumber="18">59.5 34.0 34.3 27.1 38.3</td>
<td id="76F34E020D657C999916FC53FA57FB4A" box="[1134,1436,976,1204]" gridcol="3" gridrow="4" pageId="17" pageNumber="18">58.3–60.2 32.6–35.8 33.1–35.3 24.6–29.4 35.6–40.6</td>
</tr>
<tr id="3522277E0D657C999DEFFB4BFA57FAA7" box="[151,1436,1224,1369]" gridrow="5" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFB4BFDC2FAA7" box="[151,521,1224,1369]" gridcol="0" gridrow="5" pageId="17" pageNumber="18">Preorbital depth Lower-jaw length Cheek depth Head depth</th>
<td id="76F34E020D657C999FFDFB4BFD16FAA7" box="[645,733,1224,1369]" gridcol="1" gridrow="5" pageId="17" pageNumber="18">23.5 29.7 24.5 91.6</td>
<td id="76F34E020D657C999EE1FB4BFC13FAA7" box="[921,984,1224,1369]" gridcol="2" gridrow="5" pageId="17" pageNumber="18">21.1 31.2 24.8 92.5</td>
<td id="76F34E020D657C999916FB4BFA57FAA7" box="[1134,1436,1224,1369]" gridcol="3" gridrow="5" pageId="17" pageNumber="18">19.0–23.5 29.4–34.1 23.0–26.3 89.4–96.3</td>
</tr>
<tr id="3522277E0D657C999DEFFAEEFA57FA52" box="[151,1436,1389,1452]" gridrow="6" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFAEEFDC2FA52" box="[151,521,1389,1452]" gridcol="0" gridrow="6" pageId="17" pageNumber="18">Meristics Dorsal-fin spines</th>
<td id="76F34E020D657C999FFDFAEEFD16FA52" box="[645,733,1389,1452]" gridcol="1" gridrow="6" pageId="17" pageNumber="18">17</td>
<td id="76F34E020D657C999EE1FAEEFC13FA52" box="[921,984,1389,1452]" gridcol="2" gridrow="6" pageId="17" pageNumber="18">mode 17, 18</td>
<td id="76F34E020D657C999916FAEEFA57FA52" box="[1134,1436,1389,1452]" gridcol="3" gridrow="6" pageId="17" pageNumber="18">range 17–18</td>
</tr>
<tr id="3522277E0D657C999DEFFA43FA57F95A" box="[151,1436,1472,1700]" gridrow="7" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFFA43FDC2F95A" box="[151,521,1472,1700]" gridcol="0" gridrow="7" pageId="17" pageNumber="18">Dorsal-fin rays Anal-fin spines Anal-fin rays Pelvic-fin rays Pectoral-fin rays Lateral line scales</th>
<td id="76F34E020D657C999FFDFA43FD16F95A" box="[645,733,1472,1700]" gridcol="1" gridrow="7" pageId="17" pageNumber="18">10 3 7 5 15 31</td>
<td id="76F34E020D657C999EE1FA43FC13F95A" box="[921,984,1472,1700]" gridcol="2" gridrow="7" pageId="17" pageNumber="18">9 3 8 5 14 30, 31</td>
<td id="76F34E020D657C999916FA43FA57F95A" box="[1134,1436,1472,1700]" gridcol="3" gridrow="7" pageId="17" pageNumber="18">9–10 3–3 7–8 5–5 14–15 30–31</td>
</tr>
<tr id="3522277E0D657C999DEFF93BFA57F862" box="[151,1436,1720,1948]" gridrow="8" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFF93BFDC2F862" box="[151,521,1720,1948]" gridcol="0" gridrow="8" pageId="17" pageNumber="18">Pored scales posterior to lateral line Cheek scales Gill rakers on first ceratobranchial Gill rakers on first epibranchial Teeth in outer row of left lower jaw Teeth rows on upper jaw</th>
<td id="76F34E020D657C999FFDF93BFD16F862" box="[645,733,1720,1948]" gridcol="1" gridrow="8" pageId="17" pageNumber="18">1 4 13 2 22 3</td>
<td id="76F34E020D657C999EE1F93BFC13F862" box="[921,984,1720,1948]" gridcol="2" gridrow="8" pageId="17" pageNumber="18">1 5 13 3 22 3</td>
<td id="76F34E020D657C999916F93BFA57F862" box="[1134,1436,1720,1948]" gridcol="3" gridrow="8" pageId="17" pageNumber="18">0–2 4–5 11–13 2–3 19–23 3–4</td>
</tr>
<tr id="3522277E0D657C999DEFF833FA57F838" box="[151,1436,1968,1990]" gridrow="9" pageId="17" pageNumber="18">
<th id="76F34E020D657C999DEFF833FDC2F838" box="[151,521,1968,1990]" gridcol="0" gridrow="9" pageId="17" pageNumber="18">Teeth rows on lower jaw</th>
<td id="76F34E020D657C999FFDF833FD16F838" box="[645,733,1968,1990]" gridcol="1" gridrow="9" pageId="17" pageNumber="18">3</td>
<td id="76F34E020D657C999EE1F833FC13F838" box="[921,984,1968,1990]" gridcol="2" gridrow="9" pageId="17" pageNumber="18">3</td>
<td id="76F34E020D657C999916F833FA57F838" box="[1134,1436,1968,1990]" gridcol="3" gridrow="9" pageId="17" pageNumber="18">3–3</td>
</tr>
</table>
</paragraph>
<subSubSection id="C30876B70D6683749DBFFF14FD3DFDA1" pageId="18" pageNumber="19" type="description">
<paragraph id="8BAD253C0D6683749DBFFF14FF3AFE79" blockId="18.[151,1437,151,680]" pageId="18" pageNumber="19">
<emphasis id="B966F92E0D6683749DBFFF14FE92FF4E" bold="true" box="[199,345,151,176]" pageId="18" pageNumber="19">Description.</emphasis>
Morphometric and meristic data in
<tableCitation id="C69010870D6683749E79FF14FC91FF51" box="[769,858,151,175]" captionStart="TABLE 6" captionStartId="17.[151,235,152,173]" captionTargetBox="[151,1436,314,1990]" captionTargetPageId="17" captionText="TABLE 6. Morphometric and meristic values of Metriaclima nkhunguensis from Nkhungu Reef. Mean and range include holotype." httpUri="http://table.plazi.org/id/DF6D75B40D6583779DEFFF1BFF02FF32" pageId="18" pageNumber="19" tableUuid="DF6D75B40D6583779DEFFF1BFF02FF32">Table 6</tableCitation>
. Dorsal snout profile slightly concave to straight; mouth cleft with slightly downward angle; jaws isognathous. Teeth on dentary in three rows, on premaxilla in three to four rows; outer row teeth typically bicuspid anteriorly and unicuspid posteriorly, inner rows tricuspid, innermost row unicuspid; lower pharyngeal jaw with numerous slender teeth with teeth in posterior row slightly larger. Portion of upper dental arcade normally visible in closed mouth. Tips of teeth in premaxilla and dentary in Vshaped line with anteriormost in upper and lower jaw furthest apart and separate in closed mouth. Lateral scales ctenoid.
</paragraph>
<paragraph id="8BAD253C0D6683749DBFFE17FA93FE0A" blockId="18.[151,1437,151,680]" pageId="18" pageNumber="19">Breeding males pale blue laterally fading to white-blue ventrally. Head entirely pale blue with white gular region. Dorsal fin light blue with white-blue lappets. Caudal-fin rays clear with blue membranes. Anal fin light blue and 1–5 orange ocelli. Pelvic fin light blue with white leading edge. Pectoral fin clear with orange base.</paragraph>
<paragraph id="8BAD253C0D6683749DBFFE7CFD3DFDA1" blockId="18.[151,1437,151,680]" pageId="18" pageNumber="19">Females light blue fading to white ventrally. Head light blue/gray, white gular region. Dorsal fin light blue with orange lappets. Caudal fin light blue. Anal fin light blue with 1-4 orange ocelli. Pelvic fin light blue/gray with white leading edge. Pectoral fin clear with white base.</paragraph>
</subSubSection>
<subSubSection id="C30876B70D6683749DBFFDE8FABAFD7A" box="[199,1393,619,644]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BAD253C0D6683749DBFFDE8FABAFD7A" blockId="18.[151,1437,151,680]" box="[199,1393,619,644]" pageId="18" pageNumber="19">
<emphasis id="B966F92E0D6683749DBFFDE8FEABFD7A" bold="true" box="[199,352,619,644]" pageId="18" pageNumber="19">Distribution.</emphasis>
<taxonomicName id="4C125EBF0D6683749C1FFDEFFD5AFD7A" box="[359,657,620,644]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="nkhunguensis">
<emphasis id="B966F92E0D6683749C1FFDEFFD5AFD7A" box="[359,657,620,644]" italics="true" pageId="18" pageNumber="19">Metriaclima nkhunguensis</emphasis>
</taxonomicName>
is endemic to Nkhungu and Minos reefs in
<collectingCountry id="F30565AC0D6683749903FDEFFAC4FD7A" box="[1147,1295,620,644]" name="Mozambique" pageId="18" pageNumber="19">Mozambique</collectingCountry>
(
<figureCitation id="132939B90D6683749866FDEFFAABFD7A" box="[1310,1376,620,644]" captionStart="FIGURE 1" captionStartId="1.[151,250,1266,1289]" captionTargetBox="[154,1435,196,1245]" captionTargetId="figure@1.[151,1436,193,1245]" captionTargetPageId="1" captionText="FIGURE 1. Map of Lake MalaŴi showing the locations of species discussed throughout the text." httpUri="https://zenodo.org/record/204915/files/figure.png" pageId="18" pageNumber="19">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D6683749DBFFD13FB63FD59" box="[199,1192,655,680]" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BAD253C0D6683749DBFFD13FB63FD59" blockId="18.[151,1437,151,680]" box="[199,1192,655,680]" pageId="18" pageNumber="19">
<emphasis id="B966F92E0D6683749DBFFD13FE85FD56" bold="true" box="[199,334,656,680]" pageId="18" pageNumber="19">Etymology.</emphasis>
The specific name refers to one of the reefs at which the species is endemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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