308 lines
37 KiB
XML
308 lines
37 KiB
XML
<document id="95DE17A43CB187768D8513249A442D73" ID="10.11646/zootaxa.4244.2.6SLASH10563" ID-CLB-Dataset="36238" ID-DOI="10.11646/zootaxa.4244.2.6" ID-GBIF-Dataset="bef234e7-eadb-483e-b574-e379c061a9fb" ID-ISSN="1175-5326" ID-Zenodo-Dep="400886" ID-ZooBank="4945BA57-0971-4F7B-B866-DAF157D25A08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490001616237" checkinUser="plazi" docAuthor="Pinto, Roberta R. & Fernandes, Ronaldo" docDate="2017" docId="03A987EDFF83FFD3FF3DFF7BFD3BB7B9" docLanguage="en" docName="zootaxa.4244.2.6SLASH10563.pdf" docOrigin="Zootaxa 4244 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trilepida affinis Boulenger 1884" docType="treatment" docVersion="7" lastPageNumber="251" masterDocId="FF90FF95FF80FFD6FFAAFFE3FFBFB26A" masterDocTitle="Morphological variation of Trilepida macrolepis (Peters 1857), with reappraisal of the taxonomic status of Rena affinis (Boulenger 1884) (Serpentes: Leptotyphlopidae: Epictinae)" masterLastPageNumber="260" masterPageNumber="246" pageNumber="249" updateTime="1698438173641" updateUser="ExternalLinkService">
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<mods:title id="98C76F2D149C49846B8A232D5F8B1097">Morphological variation of Trilepida macrolepis (Peters 1857), with reappraisal of the taxonomic status of Rena affinis (Boulenger 1884) (Serpentes: Leptotyphlopidae: Epictinae)</mods:title>
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<mods:namePart id="3FB7A66B5F8FA47C4AECBBE56B3C4248">Pinto, Roberta R.</mods:namePart>
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<mods:namePart id="4C7783B14DF5F676D5F195670CE60012">Fernandes, Ronaldo</mods:namePart>
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<mods:title id="BF3B42BEB7E739593E8FD1CDA5455028">Zootaxa</mods:title>
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<mods:date id="726CA73D735E3EAAE9941F590632B126">2017</mods:date>
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<treatment id="03A987EDFF83FFD3FF3DFF7BFD3BB7B9" ID-DOI="http://doi.org/10.5281/zenodo.6015154" ID-GBIF-Taxon="127848061" ID-Zenodo-Dep="6015154" LSID="urn:lsid:plazi:treatment:03A987EDFF83FFD3FF3DFF7BFD3BB7B9" httpUri="http://treatment.plazi.org/id/03A987EDFF83FFD3FF3DFF7BFD3BB7B9" lastPageId="5" lastPageNumber="251" pageId="3" pageNumber="249">
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<subSubSection id="C31A6570FF83FFD5FF3DFF7BFF66B2B9" pageId="3" pageNumber="249" type="nomenclature">
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<paragraph id="8BBF36FBFF83FFD5FF3DFF7BFD84B2D8" blockId="3.[151,571,152,211]" box="[151,571,152,178]" pageId="3" pageNumber="249">
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<heading id="D0F78197FF83FFD5FF3DFF7BFD84B2D8" bold="true" box="[151,571,152,178]" fontSize="11" level="1" pageId="3" pageNumber="249" reason="1">
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<taxonomicName id="4C004D78FF83FFD5FF3DFF7BFD84B2D8" authority="Boulenger, 1884" authorityName="Boulenger" authorityYear="1884" box="[151,571,152,178]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
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<emphasis id="B974EAE9FF83FFD5FF3DFF7BFD84B2D8" bold="true" box="[151,571,152,178]" pageId="3" pageNumber="249">
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<emphasis id="B974EAE9FF83FFD5FF3DFF7BFEE6B2D8" bold="true" box="[151,345,152,178]" italics="true" pageId="3" pageNumber="249">Trilepida affinis</emphasis>
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(
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<bibRefCitation id="EF914B0AFF83FFD5FEC2FF7BFD8CB2D8" author="Boulenger" box="[360,563,152,178]" pageId="3" pageNumber="249" refString="Boulenger, G. A. (1884) Descriptions of new species of reptiles and batrachians in the British Museum, Part. 2. Annals and Magazine of Natural History, 1884 (77), 396 - 398." type="journal article" year="1884">Boulenger, 1884</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BBF36FBFF83FFD5FF3DFF59FF66B2B9" blockId="3.[151,571,152,211]" box="[151,217,186,211]" pageId="3" pageNumber="249">
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<figureCitation id="133B2A7EFF83FFD5FF3DFF59FF66B2B9" box="[151,217,186,211]" captionStart="FIGURE 1" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[409,1178,445,1917]" captionTargetId="figure@4.[409,1178,445,1917]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal (A), lateral (B), and ventral (C) views of the head of Trilepida affinis holotype (BMNH 75.2.26.4). The arrow shows the divided base of first supralabial scale. Scale = 5 mm. Photos by R. Fernandes (MNRJ)." httpUri="https://zenodo.org/record/400887/files/figure.png" pageId="3" pageNumber="249">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A6570FF83FFD5FF3DFEE0FC1FB3D8" pageId="3" pageNumber="249" type="reference_group">
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<paragraph id="8BBF36FBFF83FFD5FF3DFEE0FD98B372" blockId="3.[151,928,258,434]" box="[151,551,258,280]" pageId="3" pageNumber="249">
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<treatmentCitationGroup id="AB1011D5FF83FFD5FF3DFEE0FD98B372" box="[151,551,258,280]" pageId="3" pageNumber="249">
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<taxonomicName id="4C004D78FF83FFD5FF3DFEE0FD9CB372" ID-CoL="896MX" authority="Boulenger 1884:396" authorityName="Boulenger" authorityPageNumber="396" authorityYear="1884" box="[151,547,258,280]" class="Squamata" family="Leptotyphlopidae" genus="Stenostoma" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affine">
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<emphasis id="B974EAE9FF83FFD5FF3DFEE0FEF4B372" box="[151,331,258,280]" italics="true" pageId="3" pageNumber="249">Stenostoma affine</emphasis>
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<treatmentCitation id="0AA110EAFF83FFD5FEFBFEE1FD9CB372" author="Boulenger" box="[337,547,258,280]" page="396" pageId="3" pageNumber="249" year="1884">
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<bibRefCitation id="EF914B0AFF83FFD5FEFBFEE1FE48B372" author="Boulenger" box="[337,503,258,280]" pageId="3" pageNumber="249" refString="Boulenger, G. A. (1884) Descriptions of new species of reptiles and batrachians in the British Museum, Part. 2. Annals and Magazine of Natural History, 1884 (77), 396 - 398." type="journal article" year="1884">Boulenger 1884</bibRefCitation>
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:396
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BBF36FBFF83FFD5FF3DFEC2FD95B35D" blockId="3.[151,928,258,434]" box="[151,554,289,311]" pageId="3" pageNumber="249">
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<treatmentCitationGroup id="AB1011D5FF83FFD5FF3DFEC2FD95B35D" box="[151,554,289,311]" pageId="3" pageNumber="249">
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<taxonomicName id="4C004D78FF83FFD5FF3DFEC2FEF8B35D" ID-CoL="3G6HL" box="[151,327,289,311]" class="Squamata" family="Leptotyphlopidae" genus="Glauconia" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
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<emphasis id="B974EAE9FF83FFD5FF3DFEC2FEF8B35D" box="[151,327,289,311]" italics="true" pageId="3" pageNumber="249">Glauconia affinis</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AA110EAFF83FFD5FECBFEC2FD99B35D" author="Boulenger" box="[353,550,289,311]" page="62" pageId="3" pageNumber="249" year="1893">
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<bibRefCitation id="EF914B0AFF83FFD5FECBFEC2FDB9B35D" author="Boulenger" box="[353,518,289,311]" pageId="3" pageNumber="249" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Natural History). Fol. I. Containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae, and Colubridae Aglyphae. British Museum of Natural History, London, pp. 57 - 71." type="book chapter" year="1893">Boulenger 1893</bibRefCitation>
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:62
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BBF36FBFF83FFD5FF3DFEA3FD7CB31E" blockId="3.[151,928,258,434]" pageId="3" pageNumber="249">
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<treatmentCitationGroup id="AB1011D5FF83FFD5FF3DFEA3FD7CB31E" pageId="3" pageNumber="249">
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<taxonomicName id="4C004D78FF83FFD5FF3DFEA3FED4B33C" box="[151,363,320,342]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
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<emphasis id="B974EAE9FF83FFD5FF3DFEA3FED4B33C" box="[151,363,320,342]" italics="true" pageId="3" pageNumber="249">Leptotyphlops affinis</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AA110EAFF83FFD5FE2CFEA3FCA9B33C" author="McDiarmid" box="[390,790,320,342]" page="19" pageId="3" pageNumber="249" year="1999">
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<bibRefCitation id="EF914B0AFF83FFD5FE2CFEA3FD46B33C" author="McDiarmid" box="[390,761,320,342]" pageId="3" pageNumber="249" refString="McDiarmid, R. W., Campbell, J. A. & Tour, T. (1999) Snake species of the world. A Taxonomic and Geographic Reference. Fol. 1. The Herpetologists' League Published, Washington, D. C., 511 pp." type="book" year="1999">McDiarmid, Campbell & Tour 1999</bibRefCitation>
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:19
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</treatmentCitation>
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.
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<taxonomicName id="4C004D78FF83FFD5FF3DFEBDFED4B31E" box="[151,363,350,372]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
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<emphasis id="B974EAE9FF83FFD5FF3DFEBDFED4B31E" box="[151,363,350,372]" italics="true" pageId="3" pageNumber="249">Leptotyphlops affinis</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AA110EAFF83FFD5FE2CFEBCFD00B31E" author="La" box="[390,703,350,372]" page="136" pageId="3" pageNumber="249" year="2004">
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<bibRefCitation id="EF914B0AFF83FFD5FE2CFEBCFD2BB31E" author="La" box="[390,660,350,372]" pageId="3" pageNumber="249" refString="La Marca, E. & Soriano, P. (2004) Reptiles de los Andes de Fenezuela. Catalogo Zoologico de Fenezuela. Fol. 2. Biogeos, Merida, 173 pp." type="book" year="2004">La Marca & Soriano 2004</bibRefCitation>
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:136
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BBF36FBFF83FFD5FF3DFE9DFC1FB3D8" blockId="3.[151,928,258,434]" pageId="3" pageNumber="249">
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<treatmentCitationGroup id="AB1011D5FF83FFD5FF3DFE9DFC1FB3D8" pageId="3" pageNumber="249">
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<taxonomicName id="4C004D78FF83FFD5FF3DFE9DFEAFB3F9" box="[151,272,381,403]" class="Reptilia" family="Leptotyphlopidae" genus="Rena" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
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<emphasis id="B974EAE9FF83FFD5FF3DFE9DFEAFB3F9" box="[151,272,381,403]" italics="true" pageId="3" pageNumber="249">Rena affinis</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AA110EAFF83FFD5FE81FE9EFCF2B3F9" author="Adalsteinsson" box="[299,845,381,403]" page="10" pageId="3" pageNumber="249" year="2009">
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<bibRefCitation id="EF914B0AFF83FFD5FE81FE9EFC8FB3F9" author="Adalsteinsson" box="[299,816,381,403]" pageId="3" pageNumber="249" refString="Adalsteinsson, S. A., Branch, W. R., Trape, S., Vitt, L. J. & Hedges, S. B. (2009) Molecular phylogeny, classification, and biogeography of snakes of the family Leptotyphlopidae (Reptilia, Squamata). Zootaxa, 2244 (2244), 1 - 50." type="journal article" year="2009">Adalsteinsson, Branch, Trape, Vitt & Hedges 2009</bibRefCitation>
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:10
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</treatmentCitation>
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<taxonomicName id="4C004D78FF83FFD5FF3DFE7FFE86B3D8" box="[151,313,412,434]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
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<emphasis id="B974EAE9FF83FFD5FF3DFE7FFE86B3D8" box="[151,313,412,434]" italics="true" pageId="3" pageNumber="249">Trilepida affinis</emphasis>
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</taxonomicName>
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—Nattera-Mummak,
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<treatmentCitation id="0AA110EAFF83FFD5FDBEFE7FFCD2B3D8" author="Gonzalez" box="[532,877,412,434]" page="296" pageId="3" pageNumber="249" year="2015">
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<bibRefCitation id="EF914B0AFF83FFD5FDBEFE7FFC85B3D8" author="Gonzalez" box="[532,826,412,434]" pageId="3" pageNumber="249" refString="Natera-Mumaw, M., Gonzalez, L. F. E. & Fernandez, M. C. (2015) Atlas Serpientes de Fenezuela. Una Fision Actual de Su Diversidad. Dimacofi Negocios Avanzados S. A., Santiago, 441 pp." type="book" year="2015">González & Fernández 2015</bibRefCitation>
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: 296
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</treatmentCitation>
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-297.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A6570FF83FFD5FF3DFE3AFE5CB07C" pageId="3" pageNumber="249" type="materials_examined">
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<paragraph id="8BBF36FBFF83FFD5FF3DFE3AFE5CB07C" blockId="3.[151,1436,473,2010]" pageId="3" pageNumber="249">
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<materialsCitation id="3B683CA6FF83FFD5FF3DFE3AFE5CB07C" ID-GBIF-Occurrence="1440684151" collectionCode="BMNH" country="Venezuela" location="Tachira" pageId="3" pageNumber="249" specimenCode="BMNH 75.2" specimenCount="1" stateProvince="Tachira" typeStatus="holotype">
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<emphasis id="B974EAE9FF83FFD5FF3DFE3AFEB4B398" bold="true" box="[151,267,473,498]" pageId="3" pageNumber="249">
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<typeStatus id="54BB8859FF83FFD5FF3DFE3AFEB9B398" box="[151,262,473,498]" pageId="3" pageNumber="249" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult male (
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<figureCitation id="133B2A7EFF83FFD5FE0BFE3AFE5CB39B" box="[417,483,473,498]" captionStart="FIGURE 1" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[409,1178,445,1917]" captionTargetId="figure@4.[409,1178,445,1917]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal (A), lateral (B), and ventral (C) views of the head of Trilepida affinis holotype (BMNH 75.2.26.4). The arrow shows the divided base of first supralabial scale. Scale = 5 mm. Photos by R. Fernandes (MNRJ)." httpUri="https://zenodo.org/record/400887/files/figure.png" pageId="3" pageNumber="249">Fig. 1</figureCitation>
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),
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<specimenCode id="DBA69E80FF83FFD5FE56FE39FD34B398" box="[508,651,473,498]" collectionCode="BMNH" pageId="3" pageNumber="249">
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<collectionCode id="ED11AE3EFF83FFD5FE56FE39FDEFB39B" box="[508,592,474,497]" pageId="3" pageNumber="249">BMNH</collectionCode>
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75.2
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</specimenCode>
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.26.4 (now
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<collectionCode id="ED11AE3EFF83FFD5FCA0FE39FCE1B39B" box="[778,862,474,497]" pageId="3" pageNumber="249">BMNH</collectionCode>
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1946.1.11.16), from state of
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<collectingRegion id="49C4F819FF83FFD5FB03FE3AFB42B398" box="[1193,1277,473,498]" country="Venezuela" name="Tachira" pageId="3" pageNumber="249">Tachira</collectingRegion>
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in
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<collectingCountry id="F317766BFF83FFD5FA88FE3AFA27B398" box="[1314,1432,473,498]" name="Venezuela" pageId="3" pageNumber="249">Venezuela</collectingCountry>
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, without specific locality data.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A6570FF83FFD5FF6DFDC2FEAFB658" pageId="3" pageNumber="249" type="diagnosis">
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<paragraph id="8BBF36FBFF83FFD5FF6DFDC2FBBCB17B" blockId="3.[151,1436,473,2010]" pageId="3" pageNumber="249">
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<emphasis id="B974EAE9FF83FFD5FF6DFDC2FEFEB050" bold="true" box="[199,321,545,570]" pageId="3" pageNumber="249">Diagnosis.</emphasis>
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<taxonomicName id="4C004D78FF83FFD5FEE1FDC2FDBDB053" box="[331,514,545,570]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
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<emphasis id="B974EAE9FF83FFD5FEE1FDC2FDBDB053" box="[331,514,545,570]" italics="true" pageId="3" pageNumber="249">Trilepida affinis</emphasis>
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</taxonomicName>
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|
can be distinguished from all other congeners by the following combination of characters: (1) snout truncate in dorsal and ventral views, and rounded in lateral view; (2) supraocular present; (3) rostral subtriangular in dorsal view, not reaching the anterior limit of ocular scales; (4) rostral shorter than supranasals; (5) frontal scale longer than other middorsal cephalic shields; (6) frontal longer than supraocular scale; (7) two supralabials (1+1); (8) four infralabials; (9) scales at midtail 10; (10) fused caudals present; (11) middorsals 214; (12) midventrals 202; (13) subcaudals 18; and (14) seven dorsal scales rows brown, contrasting with seven ventral rows cream, and head slightly lighter in dorsal view than body pattern.
|
|
</paragraph>
|
|
<paragraph id="8BBF36FBFF83FFD5FF6DFCFEFEAFB658" blockId="3.[151,1436,473,2010]" pageId="3" pageNumber="249">
|
|
<emphasis id="B974EAE9FF83FFD5FF6DFCFEFED2B15C" bold="true" box="[199,365,797,822]" pageId="3" pageNumber="249">Comparisons.</emphasis>
|
|
<taxonomicName id="4C004D78FF83FFD5FEDFFCFEFD94B15C" box="[373,555,797,822]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="affinis">
|
|
<emphasis id="B974EAE9FF83FFD5FEDFFCFEFD94B15C" box="[373,555,797,822]" italics="true" pageId="3" pageNumber="249">Trilepida affinis</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="4C004D78FF83FFD5FD6FFCFDFCDBB15C" box="[709,868,797,822]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="brasiliensis">
|
|
<emphasis id="B974EAE9FF83FFD5FD6FFCFDFCDBB15C" box="[709,868,797,822]" italics="true" pageId="3" pageNumber="249">T. brasiliensis</emphasis>
|
|
</taxonomicName>
|
|
by having supraocular scales (vs. absent); differs from
|
|
<taxonomicName id="4C004D78FF83FFD5FF7FFCA1FE26B130" box="[213,409,833,858]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="guayaguilensis">
|
|
<emphasis id="B974EAE9FF83FFD5FF7FFCA1FE26B130" box="[213,409,833,858]" italics="true" pageId="3" pageNumber="249">T. guayaguilensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C004D78FF83FFD5FE0DFCA1FDA9B130" box="[423,534,833,858]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="joshuai">
|
|
<emphasis id="B974EAE9FF83FFD5FE0DFCA1FDA9B130" box="[423,534,833,858]" italics="true" pageId="3" pageNumber="249">T. joshuai</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C004D78FF83FFD5FD8EFCA1FD03B130" box="[548,700,833,858]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="macrolepis">
|
|
<emphasis id="B974EAE9FF83FFD5FD8EFCA1FD03B130" box="[548,700,833,858]" italics="true" pageId="3" pageNumber="249">T. macrolepis</emphasis>
|
|
</taxonomicName>
|
|
(rarely), and
|
|
<taxonomicName id="4C004D78FF83FFD5FCFCFCA1FC77B133" box="[854,968,834,857]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="pastusa">
|
|
<emphasis id="B974EAE9FF83FFD5FCFCFCA1FC77B133" box="[854,968,834,857]" italics="true" pageId="3" pageNumber="249">T. pastusa</emphasis>
|
|
</taxonomicName>
|
|
by having ten scales rows on midtail (vs. 12); differs from
|
|
<taxonomicName id="4C004D78FF83FFD5FEFDFC85FE54B114" box="[343,491,869,894]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="anthracina">
|
|
<emphasis id="B974EAE9FF83FFD5FEFDFC85FE54B114" box="[343,491,869,894]" italics="true" pageId="3" pageNumber="249">T. anthracina</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C004D78FF83FFD5FE51FC85FDCDB114" box="[507,626,869,894]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="dugandi">
|
|
<emphasis id="B974EAE9FF83FFD5FE51FC85FDCDB114" box="[507,626,869,894]" italics="true" pageId="3" pageNumber="249">T. dugandi</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C004D78FF83FFD5FDD5FC85FCB4B114" box="[639,779,869,894]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="fuliginosa">
|
|
<emphasis id="B974EAE9FF83FFD5FDD5FC85FCB4B114" box="[639,779,869,894]" italics="true" pageId="3" pageNumber="249">T. fuliginosa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C004D78FF83FFD5FCB2FC85FC36B114" box="[792,905,869,894]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="koppesi">
|
|
<emphasis id="B974EAE9FF83FFD5FCB2FC85FC36B114" box="[792,905,869,894]" italics="true" pageId="3" pageNumber="249">T. koppesi</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C004D78FF83FFD5FC3CFC85FB92B114" box="[918,1069,869,894]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="macrolepis">
|
|
<emphasis id="B974EAE9FF83FFD5FC3CFC85FB92B114" box="[918,1069,869,894]" italics="true" pageId="3" pageNumber="249">T. macrolepis</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="4C004D78FF83FFD5FBC0FC85FB4AB114" box="[1130,1269,869,894]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="salgueiroi">
|
|
<emphasis id="B974EAE9FF83FFD5FBC0FC85FB4AB114" box="[1130,1269,869,894]" italics="true" pageId="3" pageNumber="249">T. salgueiroi</emphasis>
|
|
</taxonomicName>
|
|
by having two supralabials (vs. three); differs
|
|
<taxonomicName id="4C004D78FF83FFD5FDAFFC69FDC3B1C8" box="[517,636,906,930]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="nicefori">
|
|
<emphasis id="B974EAE9FF83FFD5FDAFFC69FDC3B1C8" box="[517,636,906,930]" italics="true" pageId="3" pageNumber="249">T. nicefori</emphasis>
|
|
</taxonomicName>
|
|
by having four infralabials (vs. three); differs from
|
|
<taxonomicName id="4C004D78FF83FFD5FB43FC69FACAB1C8" box="[1257,1397,905,930]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="dimidiata">
|
|
<emphasis id="B974EAE9FF83FFD5FB43FC69FACAB1C8" box="[1257,1397,905,930]" italics="true" pageId="3" pageNumber="249">T. dimidiata</emphasis>
|
|
</taxonomicName>
|
|
by having snout truncate in dorsal and ventral views (vs. rounded); differs from
|
|
<taxonomicName id="4C004D78FF83FFD5FC44FC4DFB3EB1AC" box="[1006,1153,941,966]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="brevissima">
|
|
<emphasis id="B974EAE9FF83FFD5FC44FC4DFB3EB1AC" box="[1006,1153,941,966]" italics="true" pageId="3" pageNumber="249">T. brevissima</emphasis>
|
|
</taxonomicName>
|
|
by having snout rounded on lateral view (vs. obtuse-rounded); differs from
|
|
<taxonomicName id="4C004D78FF83FFD5FD42FC31FC8BB183" box="[744,820,978,1001]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="249" phylum="Chordata" rank="species" species="jani">
|
|
<emphasis id="B974EAE9FF83FFD5FD42FC31FC8BB183" box="[744,820,978,1001]" italics="true" pageId="3" pageNumber="249">T. jani</emphasis>
|
|
</taxonomicName>
|
|
by having frontal scale longer than other middorsal cephalic shields (vs. with similar size). Refers to Table 1 for additional meristic and color differences between congeners.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C31A6570FF83FFD3FF6DFBDEFD3BB7B9" lastPageId="5" lastPageNumber="251" pageId="3" pageNumber="249" type="description">
|
|
<paragraph id="8BBF36FBFF83FFD2FF6DFBDEFBC6B3E2" blockId="3.[151,1436,473,2010]" lastBlockId="4.[151,1436,151,392]" lastPageId="4" lastPageNumber="250" pageId="3" pageNumber="249">
|
|
<emphasis id="B974EAE9FF83FFD5FF6DFBDEFD37B63C" bold="true" box="[199,648,1085,1110]" pageId="3" pageNumber="249">
|
|
Redescription of the
|
|
<typeStatus id="54BB8859FF83FFD5FE14FBDEFD9AB63C" box="[446,549,1085,1110]" pageId="3" pageNumber="249" type="holotype">holotype</typeStatus>
|
|
(
|
|
<figureCitation id="133B2A7EFF83FFD5FD9EFBDEFDC7B63C" box="[564,632,1085,1110]" captionStart="FIGURE 1" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[409,1178,445,1917]" captionTargetId="figure@4.[409,1178,445,1917]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal (A), lateral (B), and ventral (C) views of the head of Trilepida affinis holotype (BMNH 75.2.26.4). The arrow shows the divided base of first supralabial scale. Scale = 5 mm. Photos by R. Fernandes (MNRJ)." httpUri="https://zenodo.org/record/400887/files/figure.png" pageId="3" pageNumber="249">Fig. 1</figureCitation>
|
|
).
|
|
</emphasis>
|
|
Adult male, TL
|
|
<quantity id="4CF89B1EFF83FFD5FCEEFBDDFC1CB63F" box="[836,931,1086,1110]" metricMagnitude="-1" metricUnit="m" metricValue="2.18" pageId="3" pageNumber="249" unit="mm" value="218.0">218 mm</quantity>
|
|
, TAL
|
|
<quantity id="4CF89B1EFF83FFD5FC43FBDDFB84B63F" box="[1001,1083,1086,1110]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="249" unit="mm" value="24.0">24 mm</quantity>
|
|
; MB
|
|
<quantity id="4CF89B1EFF83FFD5FBD3FBDDFB6DB63F" box="[1145,1234,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="249" unit="mm" value="3.9">3.9 mm</quantity>
|
|
; MT
|
|
<quantity id="4CF89B1EFF83FFD5FAA4FBDEFAD8B63F" box="[1294,1383,1085,1110]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="249" unit="mm" value="3.5">3.5 mm</quantity>
|
|
; TL/ TAL 9.1; TL/MB 55.9; TAL/MT 6.9; HL
|
|
<quantity id="4CF89B1EFF83FFD5FDD0FB82FD6AB613" box="[634,725,1121,1146]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="249" unit="mm" value="5.3">5.3 mm</quantity>
|
|
; HW
|
|
<quantity id="4CF89B1EFF83FFD5FCB1FB81FCC9B613" box="[795,886,1122,1146]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="249" unit="mm" value="2.8">2.8 mm</quantity>
|
|
; eye diameter
|
|
<quantity id="4CF89B1EFF83FFD5FBB4FB81FBC5B613" box="[1054,1146,1122,1146]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="249" unit="mm" value="3.4">3.4 mm</quantity>
|
|
. Body subcylindrical and robust; head subcylindrical slightly distinct from body, almost twice longer than wide. Snout slightly truncate in dorsal and ventral views, and rounded in lateral view; rostral subtriangular in dorsal view, not reaching an imaginary transverse line between anterior border of ocular; rostral contacting supranasal and infranasal laterally and frontal dorsally; nasal completely divided horizontally by oblique suture crossing nostril; nostril elliptical, obliquely oriented and placed in the middle of nasal suture; supranasal as high as long, bordering rostral anteriorly, infranasal ventrally, first supralabial and ocular posteriorly, and frontal dorsally; supranasal base longer as upper border of infranasal scale; upper lip composed by rostral, infranasal, anterior supralabial, ocular and posterior supralabial (1+1); temporal scale distinct from lateral dorsal scales; first supralabial crossing nostril and reaching eye level; first supralabial on the right side divided; second supralabial trapezoidal, reaching nostril level, lower than first supralabial and in broad contact with temporal scale posteriorly; ocular enlarged, subhexagonal with straight apex, slightly rounded at eye level, as high as long, contacting posterior borders of supranasal and first supralabial anteriorly, parietal and second supralabial posteriorly, and supraocular dorsally; eyes distinct (
|
|
<quantity id="4CF89B1EFF83FFD5FA9FF9F2FA2FB440" box="[1333,1424,1553,1578]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="249" unit="mm" value="0.5">0.5 mm</quantity>
|
|
), placed centrally of expanded upper part of ocular shield, displaced above the level of nostril; supraocular as long as wide, smaller than frontal, contacting supranasal anteriorly, frontal, postfrontal and ocular laterally, and parietal posteriorly; middorsal cephalic shields (postfrontal and interparietal) subequal, hexagonal in dorsal view, weakly imbricate, as long as wide; frontal enlarged, longer than wide, longer than other middorsal, contacting rostral and supranasal anteriorly, supraocular laterally and postfrontal posteriorly; postfrontal as wide as long, contacting frontal and supraoculars anteriorly, parietals laterally and interparietal posteriorly; interparietal as wide as long, contacting postfrontal and parietals anteriorly, occipitals laterally and interoccipital posteriorly; interocciptal shorter than other middorsal, nearly twice wider than long, contacting interparietal and occipitals anteriorly, dorsal scales laterally and first middorsal scale posteriorly; parietal and occipital subequal in shape, irregularly pentagonal; parietal almost twice wider than long, lower margin contacting upper border of second supralabial, posterior margin contacting temporal, occipital and interparietal, anterior border in broad contact with ocular and supraocular and postfrontal laterally; occipital shorter than parietal, wider than long, its lower limit not reaching the level of upper margin of second supralabial; occipital separated of second supralabial by temporal; symphysial slightly cycloid, anterior and posterior borders respectively straight and slightly convex, as long as wide; four infralabials behind symphysial on both sides; first two infralabials subequal, nearly longer than wide; third infralabial slightly wider than first two; fourth infralabial distinctively longer than others. Middorsal scales 214; midventral scales 202; scale rows around midbody 14, reducing to 10 rows in the middle of the tail; cloacal shield enlarged and subtriangular; subcaudals 18; fused caudals present; terminal spine large and conical. Dorsal scales homogeneous, cycloid, smooth, weekly imbricate, and nearly two times wider than long.
|
|
</paragraph>
|
|
<caption id="DF7F6673FF84FFD2FF3DF871FB0CB5AD" httpUri="https://zenodo.org/record/400887/files/figure.png" pageId="4" pageNumber="250" targetBox="[409,1178,445,1917]" targetPageId="4">
|
|
<paragraph id="8BBF36FBFF84FFD2FF3DF871FB0CB5AD" blockId="4.[151,1435,1938,1991]" pageId="4" pageNumber="250">
|
|
<emphasis id="B974EAE9FF84FFD2FF3DF871FEAAB5C2" bold="true" box="[151,277,1938,1960]" pageId="4" pageNumber="250">FIGURE 1.</emphasis>
|
|
Dorsal (A), lateral (B), and ventral (C) views of the head of
|
|
<taxonomicName id="4C004D78FF84FFD2FC3EF871FB86B5C2" box="[916,1081,1938,1960]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="250" phylum="Chordata" rank="species" species="affinis">
|
|
<emphasis id="B974EAE9FF84FFD2FC3EF871FB86B5C2" box="[916,1081,1938,1960]" italics="true" pageId="4" pageNumber="250">Trilepida affinis</emphasis>
|
|
</taxonomicName>
|
|
holotype (BMNH 75.2.26.4). The arrow shows the divided base of first supralabial scale. Scale = 5 mm. Photos by R. Fernandes (MNRJ).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="DF7F6673FF85FFD3FF3DFB81FB9BB6FD" httpUri="https://zenodo.org/record/400888/files/figure.png" pageId="5" pageNumber="251" targetBox="[160,1424,201,1093]" targetPageId="5">
|
|
<paragraph id="8BBF36FBFF85FFD3FF3DFB81FB9BB6FD" blockId="5.[151,1436,1122,1175]" pageId="5" pageNumber="251">
|
|
<emphasis id="B974EAE9FF85FFD3FF3DFB81FEACB612" bold="true" box="[151,275,1122,1144]" pageId="5" pageNumber="251">FIGURE 2.</emphasis>
|
|
Geographic distribution of
|
|
<taxonomicName id="4C004D78FF85FFD3FD9AFB81FD6CB612" box="[560,723,1122,1144]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="251" phylum="Chordata" rank="species" species="affinis">
|
|
<emphasis id="B974EAE9FF85FFD3FD9AFB81FD6CB612" box="[560,723,1122,1144]" italics="true" pageId="5" pageNumber="251">Trilepida affinis</emphasis>
|
|
</taxonomicName>
|
|
(grid) and
|
|
<taxonomicName id="4C004D78FF85FFD3FCEDFB81FBA4B612" box="[839,1051,1122,1144]" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="251" phylum="Chordata" rank="species" species="macrolepis">
|
|
<emphasis id="B974EAE9FF85FFD3FCEDFB81FBA4B612" box="[839,1051,1122,1144]" italics="true" pageId="5" pageNumber="251">Trilepida macrolepis</emphasis>
|
|
</taxonomicName>
|
|
(circles). Type locality is represented by open symbol, examined specimens by close symbol and literature records by triangles.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BBF36FBFF85FFD3FF6DFB5DFE32B729" blockId="5.[151,1436,1214,1491]" pageId="5" pageNumber="251">
|
|
<emphasis id="B974EAE9FF85FFD3FF6DFB5DFDB5B6BD" bold="true" box="[199,522,1214,1239]" pageId="5" pageNumber="251">Coloration in preservative.</emphasis>
|
|
Body color pattern with seven dorsal scale rows brown and seven cream ventral scale rows; coloration of the head is slightly lighter than body dorsally; head color pale brown until lower limits of supralabials and first middorsal scale; lower limits of supralabials and infralabials cream; cloacal shield and terminal spine brown.
|
|
</paragraph>
|
|
<paragraph id="8BBF36FBFF85FFD3FF6DFAADFD3BB7B9" blockId="5.[151,1436,1214,1491]" pageId="5" pageNumber="251">
|
|
<materialsCitation id="3B683CA6FF85FFD3FF6DFAADFD3BB7B9" ID-GBIF-Occurrence="1440684150" collectionCode="MPL" location="Merida" pageId="5" pageNumber="251" specimenCount="1" stateProvince="Merida" typeStatus="holotype">
|
|
<emphasis id="B974EAE9FF85FFD3FF6DFAADFE50B70D" bold="true" box="[199,495,1358,1383]" pageId="5" pageNumber="251">Geographic distribution.</emphasis>
|
|
The species is known to date only from its
|
|
<typeStatus id="54BB8859FF85FFD3FC7CFAADFB84B70D" box="[982,1083,1358,1383]" pageId="5" pageNumber="251" type="holotype">holotype</typeStatus>
|
|
, which does not have a specific location of collection (
|
|
<figureCitation id="133B2A7EFF85FFD3FE38FA91FE68B7E0" box="[402,471,1394,1419]" captionStart="FIGURE 2" captionStartId="5.[151,250,1122,1144]" captionTargetBox="[160,1424,201,1093]" captionTargetId="figure@5.[151,1436,193,1101]" captionTargetPageId="5" captionText="FIGURE 2. Geographic distribution of Trilepida affinis (grid) and Trilepida macrolepis (circles). Type locality is represented by open symbol, examined specimens by close symbol and literature records by triangles." httpUri="https://zenodo.org/record/400888/files/figure.png" pageId="5" pageNumber="251">Fig. 2</figureCitation>
|
|
). Although
|
|
<collectionCode id="ED11AE3EFF85FFD3FDF1FA90FD2AB7E0" box="[603,661,1395,1418]" country="Mauritius" httpUri="http://grbio.org/cool/fjh7-aqdi" name="Musee de Port Louis" pageId="5" pageNumber="251">MPL</collectionCode>
|
|
0 1 and other specimens mentioned in literature suggest that
|
|
<taxonomicName id="4C004D78FF85FFD3FA9FFA91FF62B7C4" class="Reptilia" family="Leptotyphlopidae" genus="Trilepida" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="251" phylum="Chordata" rank="species" species="affinis">
|
|
<emphasis id="B974EAE9FF85FFD3FA9FFA91FF62B7C4" italics="true" pageId="5" pageNumber="251">Trilepida affinis</emphasis>
|
|
</taxonomicName>
|
|
may occurs along the
|
|
<collectingRegion id="49C4F819FF85FFD3FE44FA75FD80B7C5" box="[494,575,1430,1455]" country="Venezuela" name="Merida" pageId="5" pageNumber="251">Merida</collectingRegion>
|
|
Mountain Range, all these records need for confirmation on the basis of detailed examination of voucher specimens.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |