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<document id="D6839918B0EAA1343B8D5253948D6BB2" ID-DOI="10.11646/zootaxa.5492.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13234967" ID-ZooBank="47F51BFA-AC3B-42BB-BBBC-E5ACE29C3F2F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722933112560" checkinUser="plazi" docAuthor="Naik, Santhosh &amp; Shashank, P. R." docDate="2024" docId="03A987C2FFD7DA40FF23461BFC0D3900" docLanguage="en" docName="zootaxa.5492.3.7.pdf" docOrigin="Zootaxa 5492 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5492.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Choristoneura ribhoiensis Naik &amp; Shashank, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="412" masterDocId="FF90FFBAFFD6DA43FFB44008FFC73A15" masterDocTitle="Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India" masterLastPageNumber="420" masterPageNumber="409" pageNumber="410" updateTime="1722933579633" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="53C83EB94C6BC2F2EC3AA2FDC2618634">Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India</mods:title>
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<mods:name id="F94A19A42667F0B9479D113FE6F940AB" type="personal">
<mods:role id="08D5D4FFA4A67F503FC671E3CCA51ECD">
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<mods:namePart id="6884839DA963B9F2D24FB4EAB98517D1">Naik, Santhosh</mods:namePart>
<mods:affiliation id="45B5B7A4BEDB4E6279FD7069F819B8DA">National Pusa Collection, Division of Entomology, ICAR- Indian Agricultural Research Institute, New Delhi, 110012, India</mods:affiliation>
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<mods:namePart id="716798B62BE7688F123F7D29A143B461">Shashank, P. R.</mods:namePart>
<mods:nameIdentifier id="8DD0899CBC10A62B2CEE24F77C73A770" type="ORCID">0000-0002-8177-6091</mods:nameIdentifier>
<mods:nameIdentifier id="83B82537B4584726B3146FB3D293E9AC" type="email">spathour@gmail.com</mods:nameIdentifier>
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<mods:title id="A14E2E131D34184DF9904EDCCD9D9EB6">Zootaxa</mods:title>
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<mods:date id="3FE35295A5FB7D2F51A3033B56299746">2024</mods:date>
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<mods:number id="A0DE4295225810DED903E26131332635">2024-08-06</mods:number>
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<mods:number id="82554402A88D1D44676B756087C4F4FD">5492</mods:number>
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<mods:identifier id="FD08F2CC37C21B1C259719C34DB1794B" type="DOI">10.11646/zootaxa.5492.3.7</mods:identifier>
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<treatment id="03A987C2FFD7DA40FF23461BFC0D3900" LSID="urn:lsid:plazi:treatment:03A987C2FFD7DA40FF23461BFC0D3900" httpUri="http://treatment.plazi.org/id/03A987C2FFD7DA40FF23461BFC0D3900" lastPageId="3" lastPageNumber="412" pageId="1" pageNumber="410">
<subSubSection id="C31A655FFFD7DA42FF23461BFC913C38" box="[151,854,1555,1582]" pageId="1" pageNumber="410" type="nomenclature">
<paragraph id="8BBF36D4FFD7DA42FF23461BFC913C38" blockId="1.[151,854,1555,1617]" box="[151,854,1555,1582]" pageId="1" pageNumber="410">
<heading id="D0F781B8FFD7DA42FF23461BFC913C38" bold="true" box="[151,854,1555,1582]" fontSize="11" level="1" pageId="1" pageNumber="410" reason="1">
<emphasis id="B974EAC6FFD7DA42FF23461BFC913C38" bold="true" box="[151,854,1555,1582]" pageId="1" pageNumber="410">
<taxonomicName id="4C004D57FFD7DA42FF23461BFE123C3B" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[151,469,1555,1582]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="ribhoiensis" status="sp. nov.">
<emphasis id="B974EAC6FFD7DA42FF23461BFE123C3B" bold="true" box="[151,469,1555,1582]" italics="true" pageId="1" pageNumber="410">Choristoneura ribhoiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757BDFFD7DA42FE68461BFDFE3C38" box="[476,569,1555,1581]" pageId="1" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
Shashank &amp; Santhosh
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD7DA42FF23463FFEA03CA7" pageId="1" pageNumber="410" type="description">
<paragraph id="8BBF36D4FFD7DA42FF23463FFE193C44" blockId="1.[151,854,1555,1617]" box="[151,478,1591,1617]" pageId="1" pageNumber="410">
(
<figureCitation id="133B2A51FFD7DA42FF2B463FFEC43C44" box="[159,259,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1472,1496]" captionTargetBox="[162,1425,284,1438]" captionTargetId="figure-62@2.[151,1436,273,1449]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus illustrations. A. Choristoneura ribhoiensis sp. nov. (holotype, male); B. Clepsis laetornata (male); C. Clepsis laetornata (female); D. Diplocalyptis ferruginimixta (male); E. Diplocalyptis operosa (male); F. Terthreutis subrectangulus sp. nov. (holotype, male). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13234971" httpUri="https://zenodo.org/record/13234971/files/figure.png" pageId="1" pageNumber="410">Figs. 1A</figureCitation>
,
<figureCitation id="133B2A51FFD7DA42FEBB463FFEF43C44" box="[271,307,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1692,1716]" captionTargetBox="[151,1432,818,1662]" captionTargetId="figure-241@3.[151,1437,813,1669]" captionTargetPageId="3" captionText="FIGURE 2. Labial palpi. A. Choristoneura ribhoiensis sp. nov. (holotype, male); B. Clepsis laetornata (male); C. Diplocalyptis ferruginimixta (male); D. Diplocalyptis operosa (male); E. Terthreutis subrectangulus sp. nov. (holotype, male). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13234975" httpUri="https://zenodo.org/record/13234975/files/figure.png" pageId="1" pageNumber="410">2A</figureCitation>
,
<figureCitation id="133B2A51FFD7DA42FE8B463FFE423C44" box="[319,389,1591,1617]" captionStart="FIGURE 3" captionStartId="7.[151,250,1915,1939]" captionTargetBox="[162,1425,192,1880]" captionTargetId="figure-17@7.[151,1436,181,1891]" captionTargetPageId="7" captionText="FIGURE 3. Wing venation; Choristoneura ribhoiensis sp. nov. (holotype, male) (A. Forewing, B. Hindwing); Clepsis laetornata (C. Forewing, D. Hindwing)); Diplocalyptis ferruginimixta (E. Forewing, F. Hindwing); Diplocalyptis operosa (G. Forewing, H. Hindwing); Terthreutis subrectangulus sp. nov. (holotype, male) (I. Forewing, J. Hindwing). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13234977" httpUri="https://zenodo.org/record/13234977/files/figure.png" pageId="1" pageNumber="410">3A–B</figureCitation>
,
<figureCitation id="133B2A51FFD7DA42FE24463FFE113C44" box="[400,470,1591,1617]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[166,1420,192,1908]" captionTargetId="figure-23@8.[156,1431,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Choristoneura ribhoiensis sp. nov. (A. Male genitalia, B. Uncus, C. Transtilla process, D. Phallus); Clepsis laetornata (E. Male genitalia, F. Female genitalia). Scale bar A, E, F= 0.5 mm; B, C, D= 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13234979" httpUri="https://zenodo.org/record/13234979/files/figure.png" pageId="1" pageNumber="410">4A–D</figureCitation>
)
</paragraph>
<paragraph id="8BBF36D4FFD7DA42FF234674FD863C81" blockId="1.[151,577,1660,1714]" box="[151,577,1660,1684]" pageId="1" pageNumber="410">
Type locality:
<collectingCountry id="F3177644FFD7DA42FE9D4674FE9A3C81" box="[297,349,1660,1684]" name="India" pageId="1" pageNumber="410">India</collectingCountry>
(Ri Bhoi,
<collectingRegion id="49C4F836FFD7DA42FE7C4674FDFD3C81" box="[456,570,1660,1684]" country="India" name="Meghalaya" pageId="1" pageNumber="410">Meghalaya</collectingRegion>
)
</paragraph>
<paragraph id="8BBF36D4FFD7DA42FF234692FEA03CA7" blockId="1.[151,577,1660,1714]" box="[151,359,1690,1714]" pageId="1" pageNumber="410">
<materialsCitation id="3B683C89FFD7DA42FF234692FEA03CA7" box="[151,359,1690,1714]" collectionCode="INPC" pageId="1" pageNumber="410" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54BB8876FFD7DA42FF234692FF3E3CA7" box="[151,249,1690,1714]" pageId="1" pageNumber="410" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED11AE11FFD7DA42FEB74692FEFA3CA7" box="[259,317,1690,1714]" country="India" httpUri="http://grbio.org/cool/qfpa-5ik3" name="National Pusa Collections" pageId="1" pageNumber="410">INPC</collectionCode>
(
<specimenCount id="9D06FD5DFFD7DA42FEFF4692FE983CA7" box="[331,351,1690,1714]" count="1" pageId="1" pageNumber="410" type="male">♂</specimenCount>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD7DA41FF2346E8FD583AEC" lastPageId="2" lastPageNumber="411" pageId="1" pageNumber="410" type="diagnosis">
<paragraph id="8BBF36D4FFD7DA41FF2346E8FD583AEC" blockId="1.[151,1436,1760,2039]" lastBlockId="2.[151,1436,151,249]" lastPageId="2" lastPageNumber="411" pageId="1" pageNumber="410">
<emphasis id="B974EAC6FFD7DA42FF2346E8FED53CEF" bold="true" box="[151,274,1760,1786]" pageId="1" pageNumber="410">Diagnosis.</emphasis>
<taxonomicName id="4C004D57FFD7DA42FEAF46E8FD843CEF" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[283,579,1760,1786]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="ribhoiensis" status="sp. nov.">
<emphasis id="B974EAC6FFD7DA42FEAF46E8FD843CEF" box="[283,579,1760,1786]" italics="true" pageId="1" pageNumber="410">Choristoneura ribhoiensis</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFD7DA42FDF946E9FD6E3CEE" bold="true" box="[589,681,1761,1787]" pageId="1" pageNumber="410">
<taxonomicNameLabel id="A24757BDFFD7DA42FDF946E9FD6E3CEE" box="[589,681,1761,1787]" pageId="1" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is closely related to
<taxonomicName id="4C004D57FFD7DA42FC1646E8FAA83CEE" authority="Razowski, 2008" authorityName="Razowski" authorityYear="2008" box="[930,1391,1760,1787]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="chapana">
<emphasis id="B974EAC6FFD7DA42FC1646E8FB683CEF" box="[930,1199,1760,1786]" italics="true" pageId="1" pageNumber="410">Choristoneura chapana</emphasis>
<bibRefCitation id="EF914B25FFD7DA42FB0C46E8FAA83CEE" author="Razowski, J." box="[1208,1391,1760,1787]" pageId="1" pageNumber="410" pagination="283 - 299" refId="ref7112" refString="Razowski, J. (2008) On two South Asian genera Ceramea Diakonoff and Terthreutis Meyrick (Lepidoptera: Tortricidae). Polish Journal of Entomology, 4 (77), 283 - 299." type="journal article" year="2008">Razowski, 2008</bibRefCitation>
</taxonomicName>
but differ in genitalia characters. Male genitalia of
<taxonomicName id="4C004D57FFD7DA42FD01470DFC933D0B" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[693,852,1796,1823]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="ribhoiensis">
<emphasis id="B974EAC6FFD7DA42FD01470DFC933D0B" box="[693,852,1796,1823]" italics="true" pageId="1" pageNumber="410">C. ribhoiensis</emphasis>
</taxonomicName>
have the uncus longer, tegumen relatively smaller, well developed transtilla process, and sacculus not prominent. Whereas genitalia of
<taxonomicName id="4C004D57FFD7DA42FBD64721FB213D57" authorityName="Razowski" authorityYear="2008" box="[1122,1254,1832,1859]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="chapana">
<emphasis id="B974EAC6FFD7DA42FBD64721FB213D57" box="[1122,1254,1832,1859]" italics="true" pageId="1" pageNumber="410">C. chapana</emphasis>
</taxonomicName>
have the uncus relatively smaller, tegumen wider and bigger, poorly developed transtilla process and prominent sacculus running up to the outer margin of valva. Fifty-three species of
<taxonomicName id="4C004D57FFD7DA42FD434778FC5D3D9F" authorityName="Lederer" authorityYear="1859" box="[759,922,1904,1930]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAC6FFD7DA42FD434778FC5D3D9F" box="[759,922,1904,1930]" italics="true" pageId="1" pageNumber="410">Choristoneura</emphasis>
</taxonomicName>
are currently recognised globally (
<bibRefCitation id="EF914B25FFD7DA42FA974778FF3A3DBA" author="Gilligan, T. M. &amp; Baixeras, J. &amp; Brown, J. W." pageId="1" pageNumber="410" refId="ref6544" refString="Gilligan, T. M., Baixeras, J. &amp; Brown, J. W. (2018) T @ RTS: Online World Catalogue of the Tortricidae. Version. 4.0. Available from: http: // www. tortricid. net / Catalogue. asp (accessed 12 April 2023)" type="url" year="2018">
Gilligan
<emphasis id="B974EAC6FFD7DA42FA3C4779FF733DBB" italics="true" pageId="1" pageNumber="410">et al.</emphasis>
, 2018
</bibRefCitation>
), out of which six species are recorded from
<collectingCountry id="F3177644FFD7DA42FD44479CFCEB3DBB" box="[752,812,1940,1966]" name="India" pageId="1" pageNumber="410">India</collectingCountry>
, namely:
<taxonomicName id="4C004D57FFD7DA42FC23479DFB003DBB" authority="Razowski, 2006" authorityName="Razowski" authorityYear="2006" box="[919,1223,1940,1967]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="colyma">
<emphasis id="B974EAC6FFD7DA42FC23479DFBCF3DBB" box="[919,1032,1940,1967]" italics="true" pageId="1" pageNumber="410">C. colyma</emphasis>
Razowski, 2006
</taxonomicName>
;
<taxonomicName id="4C004D57FFD7DA42FB65479DFE933DC7" authority="Obraztsov, 1968" authorityName="Obraztsov" authorityYear="1968" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="ferrugininotata">
<emphasis id="B974EAC6FFD7DA42FB65479DFA5B3DBA" box="[1233,1436,1941,1967]" italics="true" pageId="1" pageNumber="410">C. ferrugininotata</emphasis>
Obraztsov, 1968
</taxonomicName>
;
<taxonomicName id="4C004D57FFD7DA42FED647B1FD0F3DC6" authority="(Meyrick, 1924)" baseAuthorityName="Meyrick" baseAuthorityYear="1924" box="[354,712,1976,2003]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="griseicoma">
<emphasis id="B974EAC6FFD7DA42FED647B1FDC63DC7" box="[354,513,1977,2003]" italics="true" pageId="1" pageNumber="410">C. griseicoma</emphasis>
(Meyrick, 1924)
</taxonomicName>
;
<taxonomicName id="4C004D57FFD7DA42FD6247B1FB833DC6" authority="(Meyrick, 1932)" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[726,1092,1976,2003]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="neurophaea">
<emphasis id="B974EAC6FFD7DA42FD6247B1FCB93DC7" box="[726,894,1976,2003]" italics="true" pageId="1" pageNumber="410">C. neurophaea</emphasis>
(Meyrick, 1932)
</taxonomicName>
;
<taxonomicName id="4C004D57FFD7DA42FBE647B1FA5B3DC6" authority="Razowski, 1992" authorityName="Razowski" authorityYear="1992" box="[1106,1436,1976,2003]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="propensa">
<emphasis id="B974EAC6FFD7DA42FBE647B1FB1B3DC7" box="[1106,1244,1977,2003]" italics="true" pageId="1" pageNumber="410">C. propensa</emphasis>
Razowski, 1992
</taxonomicName>
and
<taxonomicName id="4C004D57FFD7DA42FF7347D5FD623DE2" authority="Ganai &amp; Khan, 2017" authorityName="Ganai &amp; Khan" authorityYear="2017" box="[199,677,2012,2039]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="410" phylum="Arthropoda" rank="species" species="pseudofumiferana">
<emphasis id="B974EAC6FFD7DA42FF7347D5FE753DE3" box="[199,434,2012,2039]" italics="true" pageId="1" pageNumber="410">C. pseudofumiferana</emphasis>
Ganai &amp; Khan, 2017
</taxonomicName>
[nomen nudum] (
<bibRefCitation id="EF914B25FFD7DA42FCC447D4FB993DE2" author="Pathania, P. C. &amp; Das, A. &amp; Brown, J. &amp; Chandra, K." box="[880,1118,2012,2039]" pageId="1" pageNumber="410" pagination="1 - 95" refId="ref7020" refString="Pathania, P. C., Das, A., Brown, J. &amp; Chandra, K. (2020) Catalogue of Tortricidae Latreille, 1802 (Lepidoptera: Tortricoidea) of India. Zootaxa, 4757 (1), 1 - 95. https: // doi. org / 10.11646 / zootaxa. 4757.1.1" type="journal article" year="2020">
Pathania
<emphasis id="B974EAC6FFD7DA42FC6D47D5FBD53DE3" box="[985,1042,2012,2038]" italics="true" pageId="1" pageNumber="410">et al.</emphasis>
, 2020
</bibRefCitation>
). These species are different from
<taxonomicName id="4C004D57FFD4DA41FF62409FFEB33AA4" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[214,372,151,177]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="ribhoiensis" status="sp. nov">
<emphasis id="B974EAC6FFD4DA41FF62409FFEB33AA4" box="[214,372,151,177]" italics="true" pageId="2" pageNumber="411">C. ribhoiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757BDFFD4DA41FEC9409FFE083AA4" box="[381,463,151,177]" pageId="2" pageNumber="411" rank="species">sp. nov</taxonomicNameLabel>
but present some similar elements, such as an an incomplete transtilla divided in two labies and gnathos moderately sclerotized lateral bands with well-developed sclerotised gnathal hooks arises from tegumen shoulder and are connected medially.
</paragraph>
</subSubSection>
<caption id="DF7F665CFFD4DA41FF2345C8FD3A3C35" ID-DOI="http://doi.org/10.5281/zenodo.13234971" ID-Zenodo-Dep="13234971" httpUri="https://zenodo.org/record/13234971/files/figure.png" pageId="2" pageNumber="411" startId="2.[151,250,1472,1496]" targetBox="[162,1425,284,1438]" targetPageId="2" targetType="figure">
<paragraph id="8BBF36D4FFD4DA41FF2345C8FD3A3C35" blockId="2.[151,1437,1472,1569]" pageId="2" pageNumber="411">
<emphasis id="B974EAC6FFD4DA41FF2345C8FEDE3FCD" bold="true" box="[151,281,1472,1496]" pageId="2" pageNumber="411">FIGURE 1.</emphasis>
Dorsal habitus illustrations.
<emphasis id="B974EAC6FFD4DA41FDE045C8FDAA3FCD" bold="true" box="[596,621,1472,1496]" pageId="2" pageNumber="411">A.</emphasis>
<taxonomicName id="4C004D57FFD4DA41FDCD45C9FC4D3FCC" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[633,906,1473,1497]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="ribhoiensis" status="sp. nov.">
<emphasis id="B974EAC6FFD4DA41FDCD45C9FC4D3FCC" box="[633,906,1473,1497]" italics="true" pageId="2" pageNumber="411">Choristoneura ribhoiensis</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFD4DA41FC2145C9FC2B3FCC" bold="true" box="[917,1004,1473,1497]" pageId="2" pageNumber="411">
<taxonomicNameLabel id="A24757BDFFD4DA41FC2145C9FC2B3FCC" box="[917,1004,1473,1497]" pageId="2" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, male);
<emphasis id="B974EAC6FFD4DA41FB0D45C9FB173FCC" bold="true" box="[1209,1232,1473,1497]" pageId="2" pageNumber="411">B.</emphasis>
<taxonomicName id="4C004D57FFD4DA41FB6845C9FA5A3FCC" baseAuthorityName="Wang Li &amp; Wang" baseAuthorityYear="2003" box="[1244,1437,1473,1497]" class="Insecta" family="Tortricidae" genus="Clepsis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="laetornata">
<emphasis id="B974EAC6FFD4DA41FB6845C9FA5A3FCC" box="[1244,1437,1473,1497]" italics="true" pageId="2" pageNumber="411">Clepsis laetornata</emphasis>
</taxonomicName>
(male);
<emphasis id="B974EAC6FFD4DA41FF5345ECFF383FE9" bold="true" box="[231,255,1508,1532]" pageId="2" pageNumber="411">C.</emphasis>
<taxonomicName id="4C004D57FFD4DA41FEB245EDFE063FE8" baseAuthorityName="Wang Li &amp; Wang" baseAuthorityYear="2003" box="[262,449,1509,1533]" class="Insecta" family="Tortricidae" genus="Clepsis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="laetornata">
<emphasis id="B974EAC6FFD4DA41FEB245EDFE063FE8" box="[262,449,1509,1533]" italics="true" pageId="2" pageNumber="411">Clepsis laetornata</emphasis>
</taxonomicName>
(female);
<emphasis id="B974EAC6FFD4DA41FD9F45EDFD843FE8" bold="true" box="[555,579,1509,1533]" pageId="2" pageNumber="411">D.</emphasis>
<taxonomicName id="4C004D57FFD4DA41FDFE45EDFCAC3FE8" authorityName="Razowski" authorityYear="2009" box="[586,875,1509,1533]" class="Insecta" family="Tortricidae" genus="Diplocalyptis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="ferruginimixta">
<emphasis id="B974EAC6FFD4DA41FDFE45EDFCAC3FE8" box="[586,875,1509,1533]" italics="true" pageId="2" pageNumber="411">Diplocalyptis ferruginimixta</emphasis>
</taxonomicName>
(male);
<emphasis id="B974EAC6FFD4DA41FC7545EDFC1E3FE8" bold="true" box="[961,985,1509,1533]" pageId="2" pageNumber="411">E.</emphasis>
<taxonomicName id="4C004D57FFD4DA41FC6B45EDFB783FE9" baseAuthorityName="Meyrick" baseAuthorityYear="1908" box="[991,1215,1509,1533]" class="Insecta" family="Tortricidae" genus="Diplocalyptis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="operosa">
<emphasis id="B974EAC6FFD4DA41FC6B45EDFB783FE9" box="[991,1215,1509,1533]" italics="true" pageId="2" pageNumber="411">Diplocalyptis operosa</emphasis>
</taxonomicName>
(male);
<emphasis id="B974EAC6FFD4DA41FAA145EDFAEF3FE8" bold="true" box="[1301,1320,1509,1533]" pageId="2" pageNumber="411">F.</emphasis>
<taxonomicName id="4C004D57FFD4DA41FA9B45EDFEF63C34" authorityName="Naik &amp; Shashank" authorityYear="2024" class="Insecta" family="Tortricidae" genus="Terthreutis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="subrectangulus" status="sp. nov.">
<emphasis id="B974EAC6FFD4DA41FA9B45EDFEF63C34" italics="true" pageId="2" pageNumber="411">Terthreutis subrectangulus</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFD4DA41FE834601FE4F3C34" bold="true" box="[311,392,1545,1569]" pageId="2" pageNumber="411">
<taxonomicNameLabel id="A24757BDFFD4DA41FE834601FE4F3C34" box="[311,392,1545,1569]" pageId="2" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, male). Scale bar = 2 mm.
</paragraph>
</caption>
<subSubSection id="C31A655FFFD4DA40FF73467BFE65380C" lastPageId="3" lastPageNumber="412" pageId="2" pageNumber="411" type="description">
<paragraph id="8BBF36D4FFD4DA41FF73467BFE533D70" blockId="2.[151,1437,1651,2037]" pageId="2" pageNumber="411">
<emphasis id="B974EAC6FFD4DA41FF73467BFE9E3C98" bold="true" box="[199,345,1651,1677]" pageId="2" pageNumber="411">Description.</emphasis>
Head: Ocellus moderate, well developed and brownish compound eyes. Head whitish dusted with light grey, face fuscous. Vertex and dorsal part of frons clothed with long pale ochreous rough, upwardly projected scales. Ventral part of frons region covered with small, flat, upwardly raised pale ochreous scales. Labial palpi clothed with forwardly projected pale yellowish ochreous scales; first segment short, ochreous, tinged with whitish cream; second segment long and broad, ochreous interspersed with pale yellow along dorsal margin from base to near apex; small third segment, porrect with long ochreous scale. Antenna filiform, pale ochreous, cilia not visible. Chaetosemata present.
</paragraph>
<paragraph id="8BBF36D4FFD4DA40FF734767FA873B08" blockId="2.[151,1437,1651,2037]" lastBlockId="3.[151,1437,151,789]" lastPageId="3" lastPageNumber="412" pageId="2" pageNumber="411">
Thorax: Smooth, pronotal collar, tegulae, mesonotum, and posterior crest grey mixed with dark brown scales; legs unmodified, middle legs with a pair of tibial spurs; hind tibia covered with dense white scales interspersed with light grey scales, inner margin with tuft of ochreous grey scales. Wings: Forewing (wingspan
<quantity id="4CF89B31FFD4DA41FAA547BFFAA33DC4" box="[1297,1380,1975,2001]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="411" unit="mm" value="14.0">14 mm</quantity>
) sub rectangular, slightly widening distally, slightly sinuate and slightly convex in outer half, apex falcate, with cream ochreous scales as background interspersed with dark grey, tinged with brown ochreous and orange-grey scales; Hindwing subtriangular, dark brown, with pale brownish grey apex and greyish brown cilia along the apical and anal margin. Wing venation: Forewing with all the veins separated beyond the discal cell, R4 and R5 stalked, CuP remain at margin. Hindwing with Rs and M1 connate basally, M3 and CuA2 connate basally, CuP distinct.
</paragraph>
<paragraph id="8BBF36D4FFD5DA40FF73412FFD023BE0" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="412">Abdomen: Light grey ochreous brown. Male genitalia: Uncus long, subtriangular, broad base obtuse apically; tegumen broad, subrectangular; socius not prominent; gnathos moderately sclerotized, lateral bands; gnathal hooks sclerotized, from tegumen shoulder, connected medially; valva medium, ovate, spinose; sacculus small sclerotized with minute setae; transtilla incomplete, formed by two sclerotized lobes with minute spines, connected medially; juxta rather small, round basally; vinculum medium sclerotized. Phallus broad base tapering towards apex; vesica usually with three lanceolate cornuti, caulis short.</paragraph>
<paragraph id="8BBF36D4FFD5DA40FF7341F7FE65380C" blockId="3.[151,1437,151,789]" box="[199,418,511,537]" pageId="3" pageNumber="412">
<emphasis id="B974EAC6FFD5DA40FF7341F7FEE2380C" bold="true" box="[199,293,511,537]" pageId="3" pageNumber="412">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD5DA40FF73422AFAC43874" pageId="3" pageNumber="412" type="materials_examined">
<paragraph id="8BBF36D4FFD5DA40FF73422AFAC43874" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="412">
<emphasis id="B974EAC6FFD5DA40FF73422AFE533829" bold="true" box="[199,404,546,572]" pageId="3" pageNumber="412">Material studied.</emphasis>
<materialsCitation id="3B683C89FFD5DA40FE2F422BFED43874" collectingDate="2021-03-16" country="India" latitude="25.05611" location="Umiam" longLatPrecision="21" longitude="91.91167" municipality="Ri Bhoi" pageId="3" pageNumber="412" specimenCount="1" specimenCount-male="1" stateProvince="Meghalaya">
<collectingCountry id="F3177644FFD5DA40FE2F422BFE1E3828" box="[411,473,547,573]" name="India" pageId="3" pageNumber="412">India</collectingCountry>
:
<collectingRegion id="49C4F836FFD5DA40FE57422BFDA33828" box="[483,612,547,573]" country="India" name="Meghalaya" pageId="3" pageNumber="412">Meghalaya</collectingRegion>
:
<collectingMunicipality id="6BDBACAEFFD5DA40FDDA422BFD0D3828" box="[622,714,547,573]" pageId="3" pageNumber="412">Ri Bhoi</collectingMunicipality>
: CPGSAS,
<location id="8EDF600FFFD5DA40FCF8422BFC673828" LSID="urn:lsid:plazi:treatment:03A987C2FFD7DA40FF23461BFC0D3900:8EDF600FFFD5DA40FCF8422BFC673828" box="[844,928,547,573]" country="India" latitude="25.05611" longLatPrecision="21" longitude="91.91167" municipality="Ri Bhoi" name="Umiam" pageId="3" pageNumber="412" stateProvince="Meghalaya">Umiam</location>
(
<geoCoordinate id="EE345013FFD5DA40FC1B422AFB873828" box="[943,1088,546,573]" degrees="25" direction="north" minutes="03" orientation="latitude" pageId="3" pageNumber="412" precision="15" seconds="22" value="25.05611">25°03’22’’N</geoCoordinate>
,
<geoCoordinate id="EE345013FFD5DA40FBFF422AFB103828" box="[1099,1239,546,573]" degrees="91" direction="east" minutes="54" orientation="longitude" pageId="3" pageNumber="412" precision="15" seconds="42" value="91.91167">91°54’42’’E</geoCoordinate>
, 1520 MSL),
<specimenCount id="9D06FD5DFFD5DA40FAC6422BFA503828" box="[1394,1431,547,573]" count="1" pageId="3" pageNumber="412" type="male">1♂</specimenCount>
,
<date id="FFBE1014FFD5DA40FF23424FFED43874" box="[151,275,583,609]" pageId="3" pageNumber="412" value="2021-03-16">
<collectingDate id="EFFAE9FCFFD5DA40FF23424FFED43874" box="[151,275,583,609]" pageId="3" pageNumber="412" value="2021-03-16">16.iii.2021</collectingDate>
</date>
</materialsCitation>
, Mercury vapour lamp coll. Santhosh Naik G &amp; Sunil; genitalia slide no. L00051352 (
<specimenCount id="9D06FD5DFFD5DA40FB6A424FFB323874" box="[1246,1269,583,609]" count="1" pageId="3" pageNumber="412" type="male">♂</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD5DA40FF734263FEA438BC" pageId="3" pageNumber="412" type="etymology">
<paragraph id="8BBF36D4FFD5DA40FF734263FEA438BC" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="412">
<emphasis id="B974EAC6FFD5DA40FF734263FE893890" bold="true" box="[199,334,619,645]" pageId="3" pageNumber="412">Etymology.</emphasis>
The new species
<taxonomicName id="4C004D57FFD5DA40FD954263FC8D3890" authorityName="Naik &amp; Shashank" authorityYear="2024" box="[545,842,619,645]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="412" phylum="Arthropoda" rank="species" species="ribhoiensis" status="sp. nov.">
<emphasis id="B974EAC6FFD5DA40FD954263FC8D3890" box="[545,842,619,645]" italics="true" pageId="3" pageNumber="412">Choristoneura ribhoiensis</emphasis>
</taxonomicName>
<emphasis id="B974EAC6FFD5DA40FCE14263FC763890" bold="true" box="[853,945,619,645]" pageId="3" pageNumber="412">
<taxonomicNameLabel id="A24757BDFFD5DA40FCE14263FC763890" box="[853,945,619,645]" pageId="3" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is named after the
<typeStatus id="54BB8876FFD5DA40FB294263FB0A3890" box="[1181,1229,619,645]" pageId="3" pageNumber="412">type</typeStatus>
locality, Ri Bhoi,
<collectingRegion id="49C4F836FFD5DA40FF234287FED138BC" box="[151,278,655,681]" country="India" name="Meghalaya" pageId="3" pageNumber="412">Meghalaya</collectingRegion>
,
<collectingCountry id="F3177644FFD5DA40FE964287FE9838BC" box="[290,351,655,681]" name="India" pageId="3" pageNumber="412">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD5DA40FF7342BAFD8638D8" box="[199,577,690,717]" pageId="3" pageNumber="412" type="distribution">
<paragraph id="8BBF36D4FFD5DA40FF7342BAFD8638D8" blockId="3.[151,1437,151,789]" box="[199,577,690,717]" pageId="3" pageNumber="412">
<emphasis id="B974EAC6FFD5DA40FF7342BAFEA438D9" bold="true" box="[199,355,690,716]" pageId="3" pageNumber="412">Distribution:</emphasis>
<collectingCountry id="F3177644FFD5DA40FEDE42BBFE6338D8" box="[362,420,691,717]" name="India" pageId="3" pageNumber="412">India</collectingCountry>
(
<collectingRegion id="49C4F836FFD5DA40FE0642BBFDF238D8" box="[434,565,691,717]" country="India" name="Meghalaya" pageId="3" pageNumber="412">Meghalaya</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A655FFFD5DA40FF7342DFFC0D3900" pageId="3" pageNumber="412" type="discussion">
<paragraph id="8BBF36D4FFD5DA40FF7342DFFC0D3900" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="412">
<emphasis id="B974EAC6FFD5DA40FF7342DFFEFA38E4" bold="true" box="[199,317,727,753]" pageId="3" pageNumber="412">Remarks:</emphasis>
This new species is placed in
<taxonomicName id="4C004D57FFD5DA40FDCA42DFFCE638E4" authorityName="Lederer" authorityYear="1859" box="[638,801,727,753]" class="Insecta" family="Tortricidae" genus="Choristoneura" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="412" phylum="Arthropoda" rank="genus">
<emphasis id="B974EAC6FFD5DA40FDCA42DFFCE638E4" box="[638,801,727,753]" italics="true" pageId="3" pageNumber="412">Choristoneura</emphasis>
</taxonomicName>
based on adult resemblance and male genitalia characters. The
<typeStatus id="54BB8876FFD5DA40FF7E42F3FEEB3900" box="[202,300,763,789]" pageId="3" pageNumber="412" type="holotype">holotype</typeStatus>
was collected from a deciduous forest in
<collectingRegion id="49C4F836FFD5DA40FD4A42F3FCBA3900" box="[766,893,763,789]" country="India" name="Meghalaya" pageId="3" pageNumber="412">Meghalaya</collectingRegion>
,
<collectingCountry id="F3177644FFD5DA40FC3D42F3FC013900" box="[905,966,763,789]" name="India" pageId="3" pageNumber="412">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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