147 lines
18 KiB
XML
147 lines
18 KiB
XML
<document id="63DE31C9F3C545ADE2D78784A7AA8521" ID-CLB-Dataset="31500" ID-DOI="10.11646/zootaxa.4079.5.3" ID-GBIF-Dataset="7f223778-dd55-45a1-9878-88ae789e57ac" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050879" ID-ZooBank="CC01F586-597F-4C42-9F87-DBD81CF5AAC2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510811724910" checkinUser="plazi" docAuthor="Alwin-Kownacka, Alicja, Szadziewski, Ryszard & Szwedo, Jacek" docDate="2016" docId="03A987B447486E6EFF0A1952FAC4FB0E" docLanguage="en" docName="zootaxa.4079.5.3.pdf" docOrigin="Zootaxa 4079 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Allohelea Kieffer 1917" docType="treatment" docVersion="11" lastPageNumber="554" masterDocId="FF90FFCC474B6E6DFF9D1B12FFA0FFA8" masterDocTitle="Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species" masterLastPageNumber="572" masterPageNumber="551" pageNumber="554" updateTime="1698496611241" updateUser="ExternalLinkService">
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<mods:title id="C87E5DC6F345065FEE3A181A33B03538">Biting midges of the tribe Ceratopogonini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species</mods:title>
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<mods:namePart id="7823D6E90A8FC78EE9A245305CDE9EEB">Alwin-Kownacka, Alicja</mods:namePart>
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<mods:namePart id="6410A0265CF0AABB095150493F371721">Szadziewski, Ryszard</mods:namePart>
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<mods:namePart id="9AA0DA9DBDF6D6A638A4164F79B8D315">Szwedo, Jacek</mods:namePart>
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<mods:typeOfResource id="19D592005CB836A64998A2B3C927699D">text</mods:typeOfResource>
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<mods:title id="5D914CAE94261121AA12EE8629B53F00">Zootaxa</mods:title>
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<mods:part id="1639857258F0239DECFABE3DC4DCBC9D">
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<mods:date id="B688185B60213297C62199B7D02CFF19">2016</mods:date>
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<mods:number id="F53506D2193033C272ED7B91B4ADCBE5">2016-02-16</mods:number>
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<mods:number id="8CB1BE7D9843EB5A0EA4C9D2C19E6FF3">4079</mods:number>
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<treatment id="03A987B447486E6EFF0A1952FAC4FB0E" ID-DOI="http://doi.org/10.5281/zenodo.5700837" ID-GBIF-Taxon="137307439" ID-Zenodo-Dep="5700837" LSID="urn:lsid:plazi:treatment:03A987B447486E6EFF0A1952FAC4FB0E" httpUri="http://treatment.plazi.org/id/03A987B447486E6EFF0A1952FAC4FB0E" lastPageNumber="554" pageId="3" pageNumber="554">
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<subSubSection id="C31A652947486E6EFF0A1952FDA9FDF2" box="[151,521,575,602]" pageId="3" pageNumber="554" type="nomenclature">
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<paragraph id="8BBF36A247486E6EFF0A1952FDA9FDF2" blockId="3.[151,521,575,602]" box="[151,521,575,602]" pageId="3" pageNumber="554">
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<heading id="D0F781CE47486E6EFF0A1952FDA9FDF2" bold="true" box="[151,521,575,602]" fontSize="11" level="1" pageId="3" pageNumber="554" reason="1">
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<emphasis id="B974EAB047486E6EFF0A1952FDA9FDF2" bold="true" box="[151,521,575,602]" pageId="3" pageNumber="554">
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Genus
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<taxonomicName id="4C004D2147486E6EFF73192DFDA9FDF2" ID-CoL="SCV" ID-ENA="1874478" authority="Kieffer, 1917" authorityName="Kieffer" authorityYear="1917" box="[238,521,575,602]" class="Insecta" family="Ceratopogonidae" genus="Allohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EAB047486E6EFF73192DFEC2FDF1" bold="true" box="[238,354,575,601]" italics="true" pageId="3" pageNumber="554">Allohelea</emphasis>
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Kieffer, 1917
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A652947486E6EFF0A1995FE09FD71" pageId="3" pageNumber="554" type="materials_examined">
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<paragraph id="8BBF36A247486E6EFF0A1995FE09FD71" blockId="3.[151,1436,646,730]" pageId="3" pageNumber="554">
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Type species.
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<taxonomicName id="4C004D2147486E6EFEB81994FD69FD34" authority="Kieffer, 1911" authorityName="Kieffer" authorityYear="1911" box="[293,713,646,668]" class="Insecta" family="Ceratopogonidae" genus="Sphaeromias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B974EAB047486E6EFEB81994FD9AFD34" box="[293,570,646,668]" italics="true" pageId="3" pageNumber="554">Sphaeromias pulchripennis</emphasis>
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Kieffer, 1911
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</taxonomicName>
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; by original designation.
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<taxonomicName id="4C004D2147486E6EFF0A19B7FDE4FD13" authority="Clastrier & Delecolle, 1990" authorityName="Clastrier & Delecolle" authorityYear="1990" box="[151,580,677,699]" class="Insecta" family="Ceratopogonidae" genus="Boreohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EAB047486E6EFF0A19B7FEABFD13" box="[151,267,677,699]" italics="true" pageId="3" pageNumber="554">Boreohelea</emphasis>
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<bibRefCitation id="EF914B5347486E6EFE8A19B7FDE4FD13" author="Clastrier" box="[279,580,677,699]" pageId="3" pageNumber="554" refString="Clastrier, J. & Delecolle, J. C. (1990) Description d'un nouveau genre et de nouvelles especes africaines des genres Allohelea Kieffer, Monohelea Kieffer, Downeshelea Wirth & Grogan, Boreohelea nov. gen. (Diptera: Ceratopogonidae). Annales de la Societe Entomologique de France, 26, 129 - 157." type="journal article" year="1990">Clastrier & Delécolle, 1990</bibRefCitation>
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</taxonomicName>
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. Type species.
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<taxonomicName id="4C004D2147486E6EFD7019B7FAB8FD13" authority="Clastrier & Delecolle, 1990" authorityName="Clastrier & Delecolle" authorityYear="1990" box="[749,1304,677,699]" class="Insecta" family="Ceratopogonidae" genus="Boreohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="afrotropica">
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<emphasis id="B974EAB047486E6EFD7019B7FC7FFD13" box="[749,991,677,699]" italics="true" pageId="3" pageNumber="554">Boreohelea afrotropica</emphasis>
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<bibRefCitation id="EF914B5347486E6EFC7619B7FAB8FD13" author="Clastrier" box="[1003,1304,677,699]" pageId="3" pageNumber="554" refString="Clastrier, J. & Delecolle, J. C. (1990) Description d'un nouveau genre et de nouvelles especes africaines des genres Allohelea Kieffer, Monohelea Kieffer, Downeshelea Wirth & Grogan, Boreohelea nov. gen. (Diptera: Ceratopogonidae). Annales de la Societe Entomologique de France, 26, 129 - 157." type="journal article" year="1990">Clastrier & Delécolle, 1990</bibRefCitation>
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</taxonomicName>
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; by original designation.
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<emphasis id="B974EAB047486E6EFED519D1FE09FD71" bold="true" box="[328,425,707,730]" pageId="3" pageNumber="554">Syn. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A652947486E6EFF0A1813FEA0FC02" pageId="3" pageNumber="554" type="diagnosis">
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<paragraph id="8BBF36A247486E6EFF0A1813FEA0FC02" blockId="3.[151,1437,769,1190]" pageId="3" pageNumber="554">
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<emphasis id="B974EAB047486E6EFF0A1813FEB1FCB2" bold="true" box="[151,273,769,794]" pageId="3" pageNumber="554">Diagnosis.</emphasis>
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Species of this genus are easily recognized by having following features: tarsal hind claws in both sexes single and long; third segment of palpus swollen, with small rounded sensory pit; wing with or without dark pattern; hind femur and tibia swollen, unarmed, with distinct spines; two seminal capsules; aedeagus with proximal and distal pieces. This worldwide distributed genus now includes 59 species. From the Middle East two species are recorded.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A652947486E6EFF5A18A7FAC4FB0E" pageId="3" pageNumber="554" type="discussion">
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<paragraph id="8BBF36A247486E6EFF5A18A7FAC4FB0E" blockId="3.[151,1437,769,1190]" pageId="3" pageNumber="554">
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<emphasis id="B974EAB047486E6EFF5A18A7FEECFC66" bold="true" box="[199,332,949,974]" pageId="3" pageNumber="554">Discussion.</emphasis>
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The genus
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<taxonomicName id="4C004D2147486E6EFE7918A7FC1DFC66" authority="Clastrier & Delecolle, 1990" authorityName="Clastrier & Delecolle" authorityYear="1990" box="[484,957,949,974]" class="Insecta" family="Ceratopogonidae" genus="Boreohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EAB047486E6EFE7918A7FDC4FC66" box="[484,612,949,974]" italics="true" pageId="3" pageNumber="554">Boreohelea</emphasis>
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<bibRefCitation id="EF914B5347486E6EFDEF18A7FC1DFC66" author="Clastrier" box="[626,957,949,974]" pageId="3" pageNumber="554" refString="Clastrier, J. & Delecolle, J. C. (1990) Description d'un nouveau genre et de nouvelles especes africaines des genres Allohelea Kieffer, Monohelea Kieffer, Downeshelea Wirth & Grogan, Boreohelea nov. gen. (Diptera: Ceratopogonidae). Annales de la Societe Entomologique de France, 26, 129 - 157." type="journal article" year="1990">Clastrier & Delécolle, 1990</bibRefCitation>
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</taxonomicName>
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is proposed for
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<taxonomicName id="4C004D2147486E6EFB0D18A7FE6EFC5A" authority="Clastrier & Delecolle, 1990" authorityName="Clastrier & Delecolle" authorityYear="1990" class="Insecta" family="Ceratopogonidae" genus="Boreohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="afrotropica">
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<emphasis id="B974EAB047486E6EFB0D18A7FA3BFC66" box="[1168,1435,949,974]" italics="true" pageId="3" pageNumber="554">Boreohelea afrotropica</emphasis>
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<bibRefCitation id="EF914B5347486E6EFF0A18CBFE6EFC5A" author="Clastrier" box="[151,462,985,1010]" pageId="3" pageNumber="554" refString="Clastrier, J. & Delecolle, J. C. (1990) Description d'un nouveau genre et de nouvelles especes africaines des genres Allohelea Kieffer, Monohelea Kieffer, Downeshelea Wirth & Grogan, Boreohelea nov. gen. (Diptera: Ceratopogonidae). Annales de la Societe Entomologique de France, 26, 129 - 157." type="journal article" year="1990">Clastrier & Delécolle, 1990</bibRefCitation>
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</taxonomicName>
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described from a female with pale wings and subcylindrical fourth tarsomeres on fore and mid legs (
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<bibRefCitation id="EF914B5347486E6EFEA618EFFDD6FBBE" author="Clastrier" box="[315,630,1021,1046]" pageId="3" pageNumber="554" refString="Clastrier, J. & Delecolle, J. C. (1990) Description d'un nouveau genre et de nouvelles especes africaines des genres Allohelea Kieffer, Monohelea Kieffer, Downeshelea Wirth & Grogan, Boreohelea nov. gen. (Diptera: Ceratopogonidae). Annales de la Societe Entomologique de France, 26, 129 - 157." type="journal article" year="1990">Clastrier & Delécolle 1990</bibRefCitation>
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). All other characters as in
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<taxonomicName id="4C004D2147486E6EFC2818EFFB80FBBE" authorityName="Kieffer" authorityYear="1917" box="[949,1056,1021,1046]" class="Insecta" family="Ceratopogonidae" genus="Allohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EAB047486E6EFC2818EFFB80FBBE" box="[949,1056,1021,1046]" italics="true" pageId="3" pageNumber="554">Allohelea</emphasis>
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</taxonomicName>
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. Subsequently described male of
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<taxonomicName id="4C004D2147486E6EFF0A1F33FD2FFB92" authority="Szadziewski et al. 2011" authorityName="Szadziewski et al." authorityYear="2011" box="[151,655,1057,1082]" class="Insecta" family="Ceratopogonidae" genus="Allohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="vespertilio">
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<emphasis id="B974EAB047486E6EFF0A1F33FEDFFB92" box="[151,383,1057,1082]" italics="true" pageId="3" pageNumber="554">Allohelea vespertilio</emphasis>
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<bibRefCitation id="EF914B5347486E6EFE1B1F33FD2FFB92" author="Szadziewski" box="[390,655,1057,1082]" pageId="3" pageNumber="554" refString="Szadziewski, R., Gwizdalska-Kentzer, M. & Gilka, W. (2011) Order Diptera, family Ceratopogonidae. In: Sheikh Tahnoon Bin Zayed Al Nahyan, H. H. (Patron) & van Harten, A. (Ed.), Arthropod fauna of the United Arab Emirates. Volume 4. Dar Al Ummah Printing, Publishing, Distribution & Advertising, Abu Dhabi, UAE, pp. 636 - 653." type="book" year="2011">
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Szadziewski
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<emphasis id="B974EAB047486E6EFD841F30FDF2FB92" box="[537,594,1057,1082]" italics="true" pageId="3" pageNumber="554">et al.</emphasis>
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2011
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</bibRefCitation>
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</taxonomicName>
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with pale wings has also subcylindrical fourth tarsomeres of fore and mid legs like
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<taxonomicName id="4C004D2147486E6EFEA11F57FDE4FBF6" authorityName="Clastrier & Delecolle" authorityYear="1990" box="[316,580,1093,1118]" class="Insecta" family="Ceratopogonidae" genus="Boreohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="afrotropica">
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<emphasis id="B974EAB047486E6EFEA11F57FDE4FBF6" box="[316,580,1093,1118]" italics="true" pageId="3" pageNumber="554">Boreohelea afrotropica</emphasis>
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</taxonomicName>
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, however it has genitalia with unique aedeagus composed of two pieces typical for
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<taxonomicName id="4C004D2147486E6EFE8C1F7BFEDDFB2A" authorityName="Kieffer" authorityYear="1917" box="[273,381,1129,1154]" class="Insecta" family="Ceratopogonidae" genus="Allohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EAB047486E6EFE8C1F7BFEDDFB2A" box="[273,381,1129,1154]" italics="true" pageId="3" pageNumber="554">Allohelea</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EF914B5347486E6EFE171F7BFD25FB2A" author="Wirth" box="[394,645,1129,1154]" pageId="3" pageNumber="554" refString="Wirth, W. W. & Grogan, W. L. (1988) The predaceous midges of the World. (Diptera: Ceratopogonidae; Tribe Ceratopogonini). Flora and Fauna Handbook 4. E. J. Brill. New York, 160 pp." type="book" year="1988">Wirth & Grogan 1988</bibRefCitation>
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, de
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<bibRefCitation id="EF914B5347486E6EFD2C1F7BFC10FB2A" author="Meillon" box="[689,944,1129,1154]" pageId="3" pageNumber="554" refString="de Meillon, B. & Wirth, W. W. (1991) The genera and subgenera (excluding Culicoides) of the Afrotropical Biting Midges (Diptera: Ceratopogonidae). Annals of the Natal Museum, 32, 27 - 147." type="journal article" year="1991">Meillon & Wirth 1991</bibRefCitation>
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). This suggests that the genus
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<taxonomicName id="4C004D2147486E6EFA9E1F7BFA23FB2A" authorityName="Clastrier & Delecolle" authorityYear="1990" box="[1283,1411,1129,1154]" class="Insecta" family="Ceratopogonidae" genus="Boreohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EAB047486E6EFA9E1F7BFA23FB2A" box="[1283,1411,1129,1154]" italics="true" pageId="3" pageNumber="554">Boreohelea</emphasis>
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</taxonomicName>
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is not necessary, when we expand limits of the genus
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<taxonomicName id="4C004D2147486E6EFD491F9FFC9FFB0E" authorityName="Kieffer" authorityYear="1917" box="[724,831,1165,1190]" class="Insecta" family="Ceratopogonidae" genus="Allohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EAB047486E6EFD491F9FFC9FFB0E" box="[724,831,1165,1190]" italics="true" pageId="3" pageNumber="554">Allohelea</emphasis>
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</taxonomicName>
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including there species without patterned wings.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |