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<document id="85E71A9EE10A34267C0972435AB14C89" ID="10.11646/zootaxa.4225.1.1" ID-CLB-Dataset="37263" ID-DOI="10.5281/zenodo.259087" ID-GBIF-Dataset="05be601b-f9f1-4952-ac09-7012ef31aab0" ID-ISSN="1175-5326" ID-Zenodo-Dep="259087" ID-ZooBank="4676EE59-569B-4197-B108-9B32FD88E68E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1485243787505" checkinUser="plazi" docAuthor="Skartveit, John &amp; Nel, André" docDate="2017" docId="03A9879CFFC6F40545D335DA7E823CA3" docLanguage="en" docName="zootaxa.4225.1.1.pdf" docOrigin="Zootaxa 4225 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Plecia rhenana Heyden &amp; Heyden 1865" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="FF90FFE4FFFEF43C4544344E7D193E27" masterDocTitle="Revision of fossil Bibionidae (Insecta: Diptera) from French Oligocene deposits" masterLastPageNumber="83" masterPageNumber="1" pageNumber="57" updateTime="1698436270113" updateUser="ExternalLinkService">
<mods:mods id="B247A2F0368698F13CC790F160D90889" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BF5564AE90B7336EDC621C837984B096">
<mods:title id="ADB5FF046D47C69FE86EE59199B3B6DA">Revision of fossil Bibionidae (Insecta: Diptera) from French Oligocene deposits</mods:title>
</mods:titleInfo>
<mods:name id="96AE0A26930CA4AB105EC4442D78B976" type="personal">
<mods:role id="768C9F4F54B98005D1CA5404E6EB3ED4">
<mods:roleTerm id="C03445B8B5D9204C8B923E7B06F857DC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F6AC20BE1CEDC776293A20489EA492F">Skartveit, John</mods:namePart>
</mods:name>
<mods:name id="4B78AD1BDAD49532629EBB113BD98A0A" type="personal">
<mods:role id="1023C5ED454BAB7E0ED8BB0F5B57FC5C">
<mods:roleTerm id="BF2BAA5CB9D418EE24CFA2C81B013F2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="50BB00E616EE6023ADD7096804D8618C">Nel, André</mods:namePart>
</mods:name>
<mods:typeOfResource id="15802560E12F85549A7FF48EFC7A0953">text</mods:typeOfResource>
<mods:relatedItem id="416119A8DF41A00B0C2C490B5FB056CB" type="host">
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<mods:title id="E870F8CFF1051368902F398AF9417B2E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="29F0DBF6EFE7CB7B9EB4D526A329CCA9">
<mods:date id="3F71A6C9C35152BC09AAD3EFFE72C212">2017</mods:date>
<mods:detail id="383BD8F5A19776F44363B529D7FE69B9" type="volume">
<mods:number id="A820AB6E3D430F7705C3E9850EFD9A76">4225</mods:number>
</mods:detail>
<mods:detail id="F5D34B0573FED6FA9603A879C536BD6F" type="issue">
<mods:number id="F27DC44AA465D45B40335099B99CD761">1</mods:number>
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<mods:start id="3CD906A11EACFAC70ABF0431A21BC661">1</mods:start>
<mods:end id="730194E748F5872B594F5F02D6D887C9">83</mods:end>
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<mods:classification id="36E9C221F91EB1C4C3F4DBB49254211F">journal article</mods:classification>
<mods:identifier id="69A37EDC29DF856F1C71619EEB25FF74" type="CLB-Dataset">37263</mods:identifier>
<mods:identifier id="90DEB949241B33F1B628BF88F703C701" type="DOI">10.5281/zenodo.259087</mods:identifier>
<mods:identifier id="DE186834CD09178656B268C50191EE5A" type="GBIF-Dataset">05be601b-f9f1-4952-ac09-7012ef31aab0</mods:identifier>
<mods:identifier id="9F7F848B96508941F7BD7FA5898B6BE8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="31C6BDD687973A91728F342732066FC1" type="Zenodo-Dep">259087</mods:identifier>
<mods:identifier id="7E4E35138744718825ABB1162796A8F9" type="ZooBank">4676EE59-569B-4197-B108-9B32FD88E68E</mods:identifier>
</mods:mods>
<treatment id="03A9879CFFC6F40545D335DA7E823CA3" ID-DOI="http://doi.org/10.5281/zenodo.6039654" ID-GBIF-Taxon="127624367" ID-Zenodo-Dep="6039654" LSID="urn:lsid:plazi:treatment:03A9879CFFC6F40545D335DA7E823CA3" httpUri="http://treatment.plazi.org/id/03A9879CFFC6F40545D335DA7E823CA3" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<subSubSection id="C31A6501FFC6F40445D335DA7C703FE8" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="8BBF368AFFC6F40445D335DA7F913F89" blockId="56.[151,648,404,463]" box="[151,648,404,430]" pageId="56" pageNumber="57">
<heading id="D0F781E6FFC6F40445D335DA7F913F89" bold="true" box="[151,648,404,430]" fontSize="11" level="1" pageId="56" pageNumber="57" reason="1">
<taxonomicName id="4C004D09FFC6F40445D335DA7F913F89" ID-CoL="8DFJ2" authority="Heyden &amp; Heyden, 1865" authorityName="Heyden &amp; Heyden" authorityYear="1865" box="[151,648,404,430]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="rhenana">
<emphasis id="B974EA98FFC6F40445D335DA7F913F89" bold="true" box="[151,648,404,430]" pageId="56" pageNumber="57">
<emphasis id="B974EA98FFC6F40445D335DA7C493F89" bold="true" box="[151,336,404,430]" italics="true" pageId="56" pageNumber="57">Plecia rhenana</emphasis>
<bibRefCitation id="EF914B7BFFC6F404441335DA7F913F89" author="Heyden" box="[343,648,404,430]" pageId="56" pageNumber="57" refString="Heyden, C. von &amp; Heyden, L. von (1865) Bibioniden aus der Rheinischen Braunkohle von Rott. Palaeontographica, 14, 19 - 30." type="journal article" year="1865">Heyden &amp; Heyden, 1865</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBF368AFFC6F40445D335F87C703FE8" blockId="56.[151,648,404,463]" box="[151,361,438,463]" pageId="56" pageNumber="57">
(
<figureCitation id="133B2A0FFFC6F40445DB35F87DEB3FE8" box="[159,242,438,463]" captionStart="FIGURE 49" captionStartId="18.[151,250,1817,1839]" captionTargetBox="[188,1388,203,1786]" captionTargetId="figure@18.[184,1403,193,1796]" captionTargetPageId="18" captionText="FIGURE 49. Plecia rhenana, female. Syntype of Plecia rhenana Heyden &amp; Heyden, NHMUK In 38798. Image reproduced with permission from NHMUK." httpUri="https://zenodo.org/record/259096/files/figure.png" pageId="56" pageNumber="57">Fig. 49</figureCitation>
,
<figureCitation id="133B2A0FFFC6F40445B935F87C793FE8" box="[253,352,438,463]" captionStart="FIGURES 160 – 161" captionStartId="56.[151,264,1947,1970]" captionTargetBox="[223,1361,729,1917]" captionTargetId="figure@56.[213,1373,716,1927]" captionTargetPageId="56" captionText="FIGURES 160 – 161. Plecia rhenana Heyden &amp; Heyden, 1865 Fig. 160. Female, wing, NHMUK In 38798." httpUri="https://zenodo.org/record/259112/files/figure.png" pageId="56" pageNumber="57">
160
<emphasis id="B974EA98FFC6F404446335F97C2C3FE9" box="[295,309,439,462]" italics="true" pageId="56" pageNumber="57">–</emphasis>
161
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFC6F40445D335B07DF43C75" pageId="56" pageNumber="57" type="reference_group">
<paragraph id="8BBF368AFFC6F40445D335B07F243C33" blockId="56.[151,1436,510,594]" box="[151,573,510,532]" pageId="56" pageNumber="57">
<taxonomicName id="4C004D09FFC6F40445D335B07F213C33" ID-CoL="8DFJ2" authority="Heyden &amp; Heyden, 1865" authorityName="Heyden &amp; Heyden" authorityYear="1865" box="[151,568,510,532]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="rhenana">
<emphasis id="B974EA98FFC6F40445D335B07C2B3C33" box="[151,306,510,532]" italics="true" pageId="56" pageNumber="57">Plecia rhenana</emphasis>
<bibRefCitation id="EF914B7BFFC6F404447C35B07F213C33" author="Heyden" box="[312,568,510,532]" pageId="56" pageNumber="57" refString="Heyden, C. von &amp; Heyden, L. von (1865) Bibioniden aus der Rheinischen Braunkohle von Rott. Palaeontographica, 14, 19 - 30." type="journal article" year="1865">Heyden &amp; Heyden, 1865</bibRefCitation>
</taxonomicName>
:
</paragraph>
<paragraph id="8BBF368AFFC6F40445D336537DF43C75" blockId="56.[151,1436,510,594]" pageId="56" pageNumber="57">
<taxonomicName id="4C004D09FFC6F40445D336537CF83C14" ID-CoL="4K2NL" authority="Brongniart, 1879" authorityName="Brongniart" authorityYear="1879" box="[151,481,541,563]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B974EA98FFC6F40445D336537C353C14" box="[151,300,541,563]" italics="true" pageId="56" pageNumber="57">Plecia robusta</emphasis>
Brongniart, 1879
</taxonomicName>
(date taken from specimen label)—presumably an unpublished cabinet name in collection of EM
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFC6F40545D336307C333F67" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="materials_examined">
<paragraph id="8BBF368AFFC6F40445D3363079133CB0" blockId="56.[151,1034,638,663]" box="[151,1034,638,663]" pageId="56" pageNumber="57">
<emphasis id="B974EA98FFC6F40445D336307C993CB0" bold="true" box="[151,384,638,663]" pageId="56" pageNumber="57">Material examined.</emphasis>
Holotype of
<taxonomicName id="4C004D09FFC6F404475036307FD93CB1" box="[532,704,638,662]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="rhenana">
<emphasis id="B974EA98FFC6F404475036307FD93CB1" box="[532,704,638,662]" italics="true" pageId="56" pageNumber="57">Plecia rhenana</emphasis>
</taxonomicName>
(female) NHMUK In 38798.
</paragraph>
<caption id="DF7F6602FFC6F40445D333D57F4F39F6" httpUri="https://zenodo.org/record/259112/files/figure.png" pageId="56" pageNumber="57" targetBox="[223,1361,729,1917]" targetPageId="56">
<paragraph id="8BBF368AFFC6F40445D333D57F4F39F6" blockId="56.[151,785,1947,2031]" pageId="56" pageNumber="57">
<emphasis id="B974EA98FFC6F40445D333D57C743996" bold="true" box="[151,365,1947,1970]" pageId="56" pageNumber="57">FIGURES 160–161.</emphasis>
<taxonomicName id="4C004D09FFC6F404443733D27E083995" authority="Heyden &amp; Heyden, 1865" authorityName="Heyden &amp; Heyden" authorityYear="1865" box="[371,785,1948,1970]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="rhenana">
<emphasis id="B974EA98FFC6F404443733D27F173995" box="[371,526,1948,1970]" italics="true" pageId="56" pageNumber="57">Plecia rhenana</emphasis>
Heyden &amp; Heyden, 1865
</taxonomicName>
Fig. 160. Female, wing, NHMUK In 38798.
</paragraph>
</caption>
<paragraph id="8BBF368AFFC6F40445D333977FDC39C8" blockId="56.[151,785,1947,2031]" box="[151,709,2009,2031]" pageId="56" pageNumber="57">
<figureCitation id="133B2A0FFFC6F40445D333977DF739C8" box="[151,238,2009,2031]" captionStart="FIGURES 160 – 161" captionStartId="56.[151,264,1947,1970]" captionTargetBox="[223,1361,729,1917]" captionTargetId="figure@56.[213,1373,716,1927]" captionTargetPageId="56" captionText="FIGURES 160 – 161. Plecia rhenana Heyden &amp; Heyden, 1865 Fig. 160. Female, wing, NHMUK In 38798." httpUri="https://zenodo.org/record/259112/files/figure.png" pageId="56" pageNumber="57">Fig. 161</figureCitation>
. Female, head and thorax. NHMUK In 38798.
</paragraph>
<paragraph id="8BBF368AFFC7F405458334D9781A3E97" blockId="57.[151,1436,151,645]" box="[199,1283,151,176]" pageId="57" pageNumber="58">Additional material: 4 females, Coll. Coquand, EM: EM 70007, 70016, 70018, 70206 (A 10.14).</paragraph>
<paragraph id="8BBF368AFFC7F405458334F27F483EF3" blockId="57.[151,1436,151,645]" box="[199,593,188,213]" pageId="57" pageNumber="58">
<emphasis id="B974EA98FFC7F405458334F27C583EF2" bold="true" box="[199,321,188,213]" pageId="57" pageNumber="58">Localities.</emphasis>
Aix-en-Provence (Rott)
</paragraph>
<paragraph id="8BBF368AFFC7F405458334917C333F67" blockId="57.[151,1436,151,645]" pageId="57" pageNumber="58">
<materialsCitation id="3B683CD7FFC7F405458334917C333F67" ID-GBIF-Occurrence="1425980291" collectionCode="MNHN, FSL" collectorName="Coll. Coquand, EM" country="Germany" county="Aix" location="The" municipality="It" pageId="57" pageNumber="58" specimenCount="1" typeStatus="holotype">
<location id="8EDF6051FFC7F405458334917DEA3EDF" LSID="urn:lsid:plazi:treatment:03A9879CFFC6F40545D335DA7E823CA3:8EDF6051FFC7F405458334917DEA3EDF" box="[199,243,223,248]" country="Germany" county="Aix" municipality="It" name="The" pageId="57" pageNumber="58">The</location>
species was described from
<location id="8EDF6051FFC7F405477534AE7F7D3ED0" LSID="urn:lsid:plazi:treatment:03A9879CFFC6F40545D335DA7E823CA3:8EDF6051FFC7F405477534AE7F7D3ED0" box="[561,612,224,247]" country="Germany" county="Aix" municipality="It" name="Rott" pageId="57" pageNumber="58">Rott</location>
,
<collectingCountry id="F317761AFFC7F405472B34AE7FC03EDF" box="[623,729,224,248]" name="Germany" pageId="57" pageNumber="58">Germany</collectingCountry>
.
<collectingMunicipality id="6BDBACF0FFC7F40547A734AE7FED3ED0" box="[739,756,224,247]" pageId="57" pageNumber="58">It</collectingMunicipality>
is seemingly not present in the material from
<collectingCounty id="62DE4E06FFC7F40541B7349178043EDF" box="[1267,1309,223,248]" pageId="57" pageNumber="58">Aix</collectingCounty>
in
<collectionCode id="ED11AE4FFFC7F405400434AE78813ED0" box="[1344,1432,224,247]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="57" pageNumber="58">MNHN</collectionCode>
, but there are four females in
<collectorName id="26F5535CFFC7F405449B354A7FA53F3B" box="[479,700,260,285]" pageId="57" pageNumber="58">Coll. Coquand, EM</collectorName>
(temporarily stored at
<collectionCode id="ED11AE4FFFC7F4054684354A7EEB3F3B" box="[960,1010,260,284]" country="France" httpUri="http://grbio.org/cool/p8j1-ms8j" name="Collections de la Faculte des Sciences de Lyon" pageId="57" pageNumber="58">FSL</collectionCode>
) which are probably conspecific with the
<typeStatus id="54BB8828FFC7F405458435697C3F3F67" box="[192,294,295,320]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFC7F405458335027C583F42" box="[199,321,332,357]" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BBF368AFFC7F405458335027C583F42" blockId="57.[151,1436,151,645]" box="[199,321,332,357]" pageId="57" pageNumber="58">
<emphasis id="B974EA98FFC7F405458335027C583F42" bold="true" box="[199,321,332,357]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFC7F405458335217E823CA3" pageId="57" pageNumber="58" type="description">
<paragraph id="8BBF368AFFC7F405458335217F323FAF" blockId="57.[151,1436,151,645]" box="[199,555,367,392]" pageId="57" pageNumber="58">
<emphasis id="B974EA98FFC7F405458335217C6D3FAF" bold="true" box="[199,372,367,392]" pageId="57" pageNumber="58">Redescription.</emphasis>
Male unknown.
</paragraph>
<paragraph id="8BBF368AFFC7F405458335DA7FB23F8B" blockId="57.[151,1436,151,645]" box="[199,683,404,429]" pageId="57" pageNumber="58">
Female: Total length
<quantity id="4CF89B6FFFC7F40544F135DA7F413F8B" box="[437,600,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.23" metricValueMin="1.05" pageId="57" pageNumber="58" unit="mm" value="11.4" valueMax="12.3" valueMin="10.5">
10.5
<emphasis id="B974EA98FFC7F40544A235DB7CED3F8B" box="[486,500,405,428]" italics="true" pageId="57" pageNumber="58">–</emphasis>
12.3 mm
</quantity>
(N=4).
</paragraph>
<paragraph id="8BBF368AFFC7F405458335F9788A3FF7" blockId="57.[151,1436,151,645]" box="[199,1427,439,464]" pageId="57" pageNumber="58">
Head (fig. 161): Length
<quantity id="4CF89B6FFFC7F405449C35F67F623FE8" box="[472,635,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" metricValueMax="10.1" metricValueMin="9.1" pageId="57" pageNumber="58" unit="mm" value="0.96" valueMax="1.01" valueMin="0.91">
0.91
<emphasis id="B974EA98FFC7F405474D35F77F0E3FF7" box="[521,535,441,464]" italics="true" pageId="57" pageNumber="58">–</emphasis>
1.01 mm
</quantity>
(N=3), width
<quantity id="4CF89B6FFFC7F405465F35F67EA73FE8" box="[795,958,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.035" metricValueMax="1.1" metricValueMin="0.97" pageId="57" pageNumber="58" unit="mm" value="1.035" valueMax="1.1" valueMin="0.97">
0.97
<emphasis id="B974EA98FFC7F405460835F77E433FF7" box="[844,858,441,464]" italics="true" pageId="57" pageNumber="58">–</emphasis>
1.10 mm
</quantity>
(N=3). Flagellum length
<quantity id="4CF89B6FFFC7F405419935F978583FE8" box="[1245,1345,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="57" pageNumber="58" unit="mm" value="0.68">0.68 mm</quantity>
(N=1).
</paragraph>
<paragraph id="8BBF368AFFC7F405458335927E653FD3" blockId="57.[151,1436,151,645]" box="[199,892,476,501]" pageId="57" pageNumber="58">
Thorax: Length 2.8
<emphasis id="B974EA98FFC7F40544E435937CB73FD3" box="[416,430,477,500]" italics="true" pageId="57" pageNumber="58">–</emphasis>
3.0 mm (N=4), width
<quantity id="4CF89B6FFFC7F40547E035927E333FD3" box="[676,810,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="57" pageNumber="58" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">
2.3
<emphasis id="B974EA98FFC7F405478335937FCC3FD3" box="[711,725,477,500]" italics="true" pageId="57" pageNumber="58">–</emphasis>
2.5 mm
</quantity>
(N=3).
</paragraph>
<paragraph id="8BBF368AFFC7F405458335B17CAC3C1B" blockId="57.[151,1436,151,645]" pageId="57" pageNumber="58">
Wing (fig. 160): Length
<quantity id="4CF89B6FFFC7F405449B364E7F7E3C30" box="[479,615,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.5" metricValueMin="8.7" pageId="57" pageNumber="58" unit="mm" value="9.1" valueMax="9.5" valueMin="8.7">
8.7
<emphasis id="B974EA98FFC7F4054746364F7F093C3F" box="[514,528,513,536]" italics="true" pageId="57" pageNumber="58">–</emphasis>
9.5 mm
</quantity>
(N=4), width 3.0
<emphasis id="B974EA98FFC7F405466A364F7E253C3F" box="[814,828,513,536]" italics="true" pageId="57" pageNumber="58">–</emphasis>
<quantity id="4CF89B6FFFC7F405467835B17E8A3C30" box="[828,915,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="57" pageNumber="58" unit="mm" value="3.6">3.6 mm</quantity>
(N=4), length/width = 2.63
<emphasis id="B974EA98FFC7F4054189364F79C23C3F" box="[1229,1243,513,536]" italics="true" pageId="57" pageNumber="58">–</emphasis>
2.93 (N=4). Vein measurements in
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.
</paragraph>
<paragraph id="8BBF368AFFC7F405458336067EED3C47" blockId="57.[151,1436,151,645]" box="[199,1012,583,608]" pageId="57" pageNumber="58">
Legs: Length of fore femur
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(N=1), fore tibia
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1.7
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2.5 mm
</quantity>
(N=3).
</paragraph>
<paragraph id="8BBF368AFFC7F405458336227E823CA3" blockId="57.[151,1436,151,645]" box="[199,923,620,645]" pageId="57" pageNumber="58">
Abdomen: Length
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7.6
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9.4 mm
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(N=4), width 2.5
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3.0 mm (N=3).
</paragraph>
</subSubSection>
</treatment>
</document>
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