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<document id="9731E450DA8646649E140302DB4B4079" ID-DOI="10.1653/024.098.0116" ID-ISSN="1938-5102" ID-Zenodo-Dep="12769502" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721258582094" checkinUser="felipe" docAuthor="Gasca-Álvarez, Héctor Jaime &amp; Deloya, Cuauhtémoc" docDate="2015" docId="03A98784FFDEFFB161E4FEFAF9C63ED7" docLanguage="en" docName="FlaEntomol.98.1.100-103.pdf" docOrigin="Florida Entomologist 98 (1)" docSource="http://www.bioone.org/doi/10.1653/024.098.0116" docStyle="DocumentStyle:C0CD22107794FB65089F701CBA118AA5.3:FlaEntomol.2015-.journal_article" docStyleId="C0CD22107794FB65089F701CBA118AA5" docStyleName="FlaEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Hologymnetis reyesi Gasca-Alvarez and Deloya 2015, new species" docType="treatment" docVersion="2" masterDocId="FF90FFFCFFDCFFB56174FFF1FFC63D38" masterDocTitle="A new Mexican species of Hologymnetis Martínez (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini)" masterLastPageNumber="100" masterPageNumber="100" pageNumber="100" updateTime="1721267252050" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2FD7737BC95E98B61F763A82C090F9D7">A new Mexican species of Hologymnetis Martínez (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini)</mods:title>
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<mods:namePart id="25AAADBCDAADE614EC9136EB607ECA77">Gasca-Álvarez, Héctor Jaime</mods:namePart>
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<mods:namePart id="6AF735A2899141D80E324BBD43936DAD">Deloya, Cuauhtémoc</mods:namePart>
</mods:name>
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<mods:date id="D592EAD7A8721CC630178AE3664F40C6">2015</mods:date>
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<mods:number id="542683BC3AD3DDFFFBBD4D92AFFF7808">2015-03-31</mods:number>
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<mods:number id="BEF4D828D7185EB49E705B5CAEA81D61">98</mods:number>
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<mods:identifier id="D73FAE6AE877A79264C9C488FEEBF503" type="DOI">10.1653/024.098.0116</mods:identifier>
<mods:identifier id="EDBF978E6D8D6236EE3A5FAEACB4DE5D" type="ISSN">1938-5102</mods:identifier>
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<treatment id="03A98784FFDEFFB161E4FEFAF9C63ED7" LSID="urn:lsid:plazi:treatment:03A98784FFDEFFB161E4FEFAF9C63ED7" httpUri="http://treatment.plazi.org/id/03A98784FFDEFFB161E4FEFAF9C63ED7" lastPageId="4" pageId="2">
<subSubSection id="C31A6519FFDEFFB761E4FEFAFEF83C7E" pageId="2" type="nomenclature">
<paragraph id="8BBF3692FFDEFFB761E4FEFAFEF83C7E" blockId="2.[144,816,267,326]" pageId="2">
<heading id="D0F781FEFFDEFFB761E4FEFAFEF83C7E" centered="true" fontSize="10" level="2" pageId="2" reason="3">
<taxonomicName id="4C004D11FFDEFFB761E4FEFAFD7D3C1C" ID-CoL="6M9YN" authority="Gasca-Alvarez and Deloya" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[144,699,267,292]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="reyesi" status="sp. nov.">
<emphasis id="B974EA80FFDEFFB761E4FEFAFEB93C1B" box="[144,383,267,291]" italics="true" pageId="2">Hologymnetis reyesi</emphasis>
Gasca-Álvarez and Deloya
</taxonomicName>
,
<taxonomicNameLabel id="A24757FBFFDEFFB763B1FEFAFF7B3C7E" pageId="2" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="133B2A17FFDEFFB761BEFEDCFEF33C7E" box="[202,309,301,326]" captionStart-0="Figs" captionStart-1="Fig" captionStart-2="Figs" captionStartId-0="2.[156,193,1966,1984]" captionStartId-1="2.[868,896,1991,2009]" captionStartId-2="3.[108,145,1293,1311]" captionTargetBox-0="[150,821,1202,1924]" captionTargetBox-1="[861,1530,1052,1961]" captionTargetBox-2="[103,771,197,1244]" captionTargetId-0="figure-461@2.[144,824,1198,1928]" captionTargetId-1="figure-487@2.[856,1536,1049,1963]" captionTargetId-2="figure-422@3.[96,776,192,1248]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figs. 14. Hologymnetis reyesi, 1, habitus, dorsal aspect; 2, habitus, lateral aspect; 3, pygidium, 4a, parameres, frontal view; 4b, parameres, lateral view." captionText-1="Fig. 5. Hologymnetis reyesi, mesometasternal process (scale bar: 0.5 mm)." captionText-2="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi-0="http://doi.org/10.5281/zenodo.12769504" figureDoi-1="http://doi.org/10.5281/zenodo.12769506" figureDoi-2="http://doi.org/10.5281/zenodo.12769508" httpUri-0="https://zenodo.org/record/12769504/files/figure.png" httpUri-1="https://zenodo.org/record/12769506/files/figure.png" httpUri-2="https://zenodo.org/record/12769508/files/figure.png" pageId="2">Figs. 16</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A6519FFDEFFB761E4FE87FEE33F35" pageId="2" type="materials_examined">
<paragraph id="8BBF3692FFDEFFB761E4FE87FEFF3CB4" blockId="2.[144,313,374,396]" box="[144,313,374,396]" pageId="2">
<heading id="D0F781FEFFDEFFB761E4FE87FEFF3CB4" box="[144,313,374,396]" fontSize="10" level="3" pageId="2" reason="9">
<collectionCode id="ED11AE57FFDEFFB761E4FE87FEFF3CB4" box="[144,313,374,396]" pageId="2">TYPE MATERIAL</collectionCode>
</heading>
</paragraph>
<paragraph id="8BBF3692FFDEFFB761C0FE50FEE33F35" blockId="2.[144,825,417,525]" pageId="2">
<materialsCitation id="3B683CCFFFDEFFB761C0FE50FEE33F35" collectorName="B. K. Dozier" country="Mexico" location="Division of Plant Industry" municipality="Collection of Arthropods" pageId="2" specimenCount="1" stateProvince="Colima" typeStatus="holotype">
<typeStatus id="54BB8830FFDEFFB761C0FE50FECB3C8D" box="[180,269,417,437]" pageId="2" type="holotype">Holotype</typeStatus>
male labeled “
<collectingCountry id="F3177602FFDEFFB760C0FE50FDCD3C8D" box="[436,523,417,437]" name="Mexico" pageId="2">MEXICO</collectingCountry>
,
<collectingRegion id="49C4F870FFDEFFB7636EFE50FDA73C8D" box="[538,609,417,437]" country="Mexico" name="Colima" pageId="2">Colima</collectingRegion>
,
<quantity id="4CF89B77FFDEFFB76305FE50FD053C8D" box="[625,707,417,437]" metricMagnitude="4" metricUnit="m" metricValue="1.81855872" pageId="2" unit="mi" value="11.3">11.3 mi</quantity>
S.
<collectingRegion id="49C4F870FFDEFFB7639BFE50FCF03C8D" box="[751,822,417,437]" country="Mexico" name="Colima" pageId="2">Colima</collectingRegion>
. vi.27.1983.
<collectorName id="26F55344FFDEFFB76077FE4FFEBD3CEA" box="[259,379,446,466]" pageId="2">B. K. Dozier</collectorName>
” and deposited in Florida State
<collectingMunicipality id="6BDBACE8FFDEFFB763CFFE4FFEC43CD7" pageId="2">Collection of Arthropods</collectingMunicipality>
,
<location id="8EDF6049FFDEFFB7607FFE2AFDC43CD7" LSID="urn:lsid:plazi:treatment:03A98784FFDEFFB161E4FEFAF9C63ED7:8EDF6049FFDEFFB7607FFE2AFDC43CD7" box="[267,514,475,495]" country="Mexico" municipality="Collection of Arthropods" name="Division of Plant Industry" pageId="2" stateProvince="Colima">Division of Plant Industry</location>
- Entomology, Gainesville, Florida 32608 USA.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A6519FFDEFFB761E4FDD7FC273E24" pageId="2" type="description">
<paragraph id="8BBF3692FFDEFFB761E4FDD7FEE73F04" blockId="2.[144,289,550,572]" box="[144,289,550,572]" pageId="2">
<heading id="D0F781FEFFDEFFB761E4FDD7FEE73F04" box="[144,289,550,572]" fontSize="10" level="3" pageId="2" reason="9">DESCRIPTION</heading>
</paragraph>
<paragraph id="8BBF3692FFDEFFB761C0FDA0FA303F94" blockId="2.[144,826,593,1141]" lastBlockId="2.[856,1537,195,684]" pageId="2">
<typeStatus id="54BB8830FFDEFFB761C0FDA0FEDB3F5D" box="[180,285,593,613]" pageId="2" type="holotype">HOLOTYPE</typeStatus>
. Male (
<figureCitation id="133B2A17FFDEFFB7602BFDA0FE703F5D" box="[351,438,593,613]" captionStart="Figs" captionStartId="2.[156,193,1966,1984]" captionTargetBox="[150,821,1202,1924]" captionTargetId="figure-461@2.[144,824,1198,1928]" captionTargetPageId="2" captionText="Figs. 14. Hologymnetis reyesi, 1, habitus, dorsal aspect; 2, habitus, lateral aspect; 3, pygidium, 4a, parameres, frontal view; 4b, parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12769504" httpUri="https://zenodo.org/record/12769504/files/figure.png" pageId="2">Figs. 12</figureCitation>
). Length
<quantity id="4CF89B77FFDEFFB7637FFDA0FDA83F5D" box="[523,622,593,613]" metricMagnitude="-2" metricUnit="m" metricValue="1.757" pageId="2" unit="mm" value="17.57">17.57 mm</quantity>
; width across humeri
<quantity id="4CF89B77FFDEFFB761E4FD9FFF2E3FBA" box="[144,232,622,642]" metricMagnitude="-3" metricUnit="m" metricValue="9.97" pageId="2" unit="mm" value="9.97">9.97 mm</quantity>
. Color dorsally varying from opaque light to smokey gray; pronotum usually with light brown, longitudinal line in center; elytra usually with broad, transverse, diffuse light brown band just behind middle, with 2 black spots and usually with weakly raised costae (terminating at apical umbone) darker than background color; venter weakly glossy, darker smoky gray, occasionally with faint metallic copper reflection; punctures and other sculpturing black. Head: Surface densely punctate; punctures small to large, round to irregular in shape;setae minute,straw colored. Clypeus feebly tumescent at center; apex strongly reflexed, emarginate at center, apices usually distinctly angulate either side of center. Interocular width is 4.05.0 eye diameters as seen from above. Antenna with basal antennomere at apex with 10 or fewer pale, slender setae on dorsal edge. Pronotum. Surface moderately punctate; punctures small to large (larger and denser laterally), round to kidney-shaped, disk strongly punctate. Anterior margin without tubercle. Bead of lateral margin incomplete; margin just anterior of basolateral angle distinctly emarginate. Mesepimera densely punctate; punctures round to kidney-shaped,setigerous; setae pale, ofen dense, long. Elytra.Surface densely punctate (much less so basomedially); punctures varying from small to mostly large, round kidney-shaped, U-shaped near to apex. Weakly elevated sutural costa and 2 discal costae terminating at apical umbone. Pygidium. Surface with coarsely, transverse,vermiform punctures;punctures strongly impressed, kidney-shaped to elongate, setigerous (
<figureCitation id="133B2A17FFDEFFB764A9FEC9FCA13C51" captionStart="Figs" captionStartId="2.[156,193,1966,1984]" captionTargetBox="[150,821,1202,1924]" captionTargetId="figure-461@2.[144,824,1198,1928]" captionTargetPageId="2" captionText="Figs. 14. Hologymnetis reyesi, 1, habitus, dorsal aspect; 2, habitus, lateral aspect; 3, pygidium, 4a, parameres, frontal view; 4b, parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12769504" httpUri="https://zenodo.org/record/12769504/files/figure.png" pageId="2">Fig. 3</figureCitation>
); setae very short, pale. Venter: Setae testaceous. Mesometasternal protrusion short, broad, extending anteriorly beyond mesocoxae but not reaching procoxae; surface piceus, glossy, apex rounded, anterior surface with long, slender setae that curl around apex of protrusion. In ventral view, sides of mesometasternal protrusion slightly diverging to rounded apex (
<figureCitation id="133B2A17FFDEFFB76293FE19FBD83CC4" box="[999,1054,488,508]" captionStart="Fig" captionStartId="2.[868,896,1991,2009]" captionTargetBox="[861,1530,1052,1961]" captionTargetId="figure-487@2.[856,1536,1049,1963]" captionTargetPageId="2" captionText="Fig. 5. Hologymnetis reyesi, mesometasternal process (scale bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.12769506" httpUri="https://zenodo.org/record/12769506/files/figure.png" pageId="2">Fig. 5</figureCitation>
). Last sternite at apex with fringe of slender, short, testaceous setae. Sternites completely smokey gray and with small piceous spot basomedially; punctures moderate to large (larger laterally), kidney-shaped. Legs.Femora colored similarly to venter, tibiae predomi- nantly piceous. Protibia with 3 subequally spaced teeth, middle tooth slightly closer to anterior tooth than basal. Parameres: Rounded at base in caudal view, apices with strong, laterally projecting tooth (
<figureCitation id="133B2A17FFDEFFB764E0FD69FA123F94" box="[1428,1492,664,684]" captionStart="Figs" captionStartId="2.[156,193,1966,1984]" captionTargetBox="[150,821,1202,1924]" captionTargetId="figure-461@2.[144,824,1198,1928]" captionTargetPageId="2" captionText="Figs. 14. Hologymnetis reyesi, 1, habitus, dorsal aspect; 2, habitus, lateral aspect; 3, pygidium, 4a, parameres, frontal view; 4b, parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12769504" httpUri="https://zenodo.org/record/12769504/files/figure.png" pageId="2">Figs. 4</figureCitation>
,
<figureCitation id="133B2A17FFDEFFB764A8FD69FA2D3F94" box="[1500,1515,664,684]" captionStart="Figs" captionStartId="3.[108,145,1293,1311]" captionTargetBox="[103,771,197,1244]" captionTargetId="figure-422@3.[96,776,192,1248]" captionTargetPageId="3" captionText="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi="http://doi.org/10.5281/zenodo.12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="2">6</figureCitation>
).
</paragraph>
<paragraph id="8BBF3692FFDEFFB7622CFD2CFC763FCB" blockId="2.[856,944,733,755]" box="[856,944,733,755]" pageId="2">
<heading id="D0F781FEFFDEFFB7622CFD2CFC763FCB" box="[856,944,733,755]" fontSize="10" level="3" pageId="2" reason="9">FEMALE</heading>
</paragraph>
<paragraph id="8BBF3692FFDEFFB76208FCF9FC273E24" blockId="2.[892,993,776,796]" box="[892,993,776,796]" pageId="2">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31A6519FFDEFFB6622CFCBCFC1E3A55" lastPageId="3" pageId="2" type="distribution">
<paragraph id="8BBF3692FFDEFFB7622CFCBCFC323E5B" blockId="2.[856,1012,845,867]" box="[856,1012,845,867]" pageId="2">
<heading id="D0F781FEFFDEFFB7622CFCBCFC323E5B" box="[856,1012,845,867]" fontSize="10" level="3" pageId="2" reason="9">DISTRIBUTION</heading>
</paragraph>
<paragraph id="8BBF3692FFDEFFB76208FC89FA0C3EFF" blockId="2.[856,1536,888,967]" pageId="2">
<taxonomicName id="4C004D11FFDEFFB76208FC89FBFC3EB4" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[892,1082,888,908]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B974EA80FFDEFFB76208FC89FBFC3EB4" box="[892,1082,888,908]" italics="true" pageId="2">Hologymnetis reyesi</emphasis>
</taxonomicName>
<materialsCitation id="3B683CCFFFDEFFB76536FC89FA003EFF" country="Mexico" location="Colima" pageId="2" specimenCount="1" stateProvince="Colima">
is known only from the
<typeStatus id="54BB8830FFDEFFB7645EFC89FA923EB4" box="[1322,1364,888,908]" pageId="2">type</typeStatus>
locality at
<quantity id="4CF89B77FFDEFFB764CBFC89FCB03E91" metricMagnitude="4" metricUnit="m" metricValue="1.8181" pageId="2" unit="km" value="18.181">18.181 km</quantity>
south of
<collectingRegion id="49C4F870FFDEFFB762B8FC64FBD63E91" box="[972,1040,917,937]" country="Mexico" name="Colima" pageId="2">Colima</collectingRegion>
,
<collectingRegion id="49C4F870FFDEFFB76563FC64FB493E91" box="[1047,1167,917,937]" country="Mexico" name="Colima" pageId="2">Colima State</collectingRegion>
,
<collectingCountry id="F3177602FFDEFFB765E2FC64FB183E91" box="[1174,1246,917,937]" name="Mexico" pageId="2">Mexico</collectingCountry>
; probably corresponding to the municipality of Ixtlahuacan. The habitat is tropical deciduous forest
</materialsCitation>
.
</paragraph>
<caption id="DF7F661AFFDEFFB761E8F85FFCEC3AE0" ID-DOI="http://doi.org/10.5281/zenodo.12769504" ID-Zenodo-Dep="12769504" httpUri="https://zenodo.org/record/12769504/files/figure.png" pageId="2" startId="2.[156,193,1966,1984]" targetBox="[150,821,1202,1924]" targetPageId="2" targetType="figure">
<paragraph id="8BBF3692FFDEFFB761E8F85FFCEC3AE0" blockId="2.[144,824,1966,2008]" pageId="2">
<emphasis id="B974EA80FFDEFFB761E8F85FFF373AF8" bold="true" box="[156,241,1966,1984]" pageId="2">Figs. 14.</emphasis>
<emphasis id="B974EA80FFDEFFB7618CF85FFE693AF8" box="[248,431,1966,1984]" italics="true" pageId="2">
<taxonomicName id="4C004D11FFDEFFB7618CF85FFE6D3AF8" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[248,427,1966,1984]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="reyesi">Hologymnetis reyesi</taxonomicName>
,
</emphasis>
1, habitus, dorsal aspect; 2, habitus, lateral aspect; 3, pygidium, 4a, parameres, frontal view; 4b, parameres, lateral view.
</paragraph>
</caption>
<caption id="DF7F661AFFDEFFB76210F836FA233AE1" ID-DOI="http://doi.org/10.5281/zenodo.12769506" ID-Zenodo-Dep="12769506" box="[868,1509,1991,2009]" httpUri="https://zenodo.org/record/12769506/files/figure.png" pageId="2" startId="2.[868,896,1991,2009]" targetBox="[861,1530,1052,1961]" targetPageId="2" targetType="figure">
<paragraph id="8BBF3692FFDEFFB76210F836FA233AE1" blockId="2.[868,1509,1991,2009]" box="[868,1509,1991,2009]" pageId="2">
<emphasis id="B974EA80FFDEFFB76210F836FC5E3AE1" bold="true" box="[868,920,1991,2009]" pageId="2">Fig. 5.</emphasis>
<taxonomicName id="4C004D11FFDEFFB762E9F836FB8D3AE1" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[925,1099,1991,2009]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B974EA80FFDEFFB762E9F836FB8D3AE1" box="[925,1099,1991,2009]" italics="true" pageId="2">Hologymnetis reyesi</emphasis>
</taxonomicName>
, mesometasternal process (scale bar: 0.5 mm).
</paragraph>
</caption>
<caption id="DF7F661AFFDFFFB66118FAFCFD1F385F" ID-DOI="http://doi.org/10.5281/zenodo.12769508" ID-Zenodo-Dep="12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="3" startId="3.[108,145,1293,1311]" targetBox="[103,771,197,1244]" targetPageId="3" targetType="figure">
<paragraph id="8BBF3692FFDFFFB66118FAFCFCCE3827" blockId="3.[96,776,1293,1383]" box="[108,776,1293,1311]" pageId="3">
<emphasis id="B974EA80FFDFFFB66118FAFCFF073827" bold="true" box="[108,193,1293,1311]" pageId="3">Figs. 68.</emphasis>
Parameres, 6a,
<taxonomicName id="4C004D11FFDFFFB66026FAFCFDC73827" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[338,513,1293,1311]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B974EA80FFDFFFB66026FAFCFDC73827" box="[338,513,1293,1311]" italics="true" pageId="3">Hologymnetis reyesi</emphasis>
</taxonomicName>
, frontal view (scale bar = 0.3
</paragraph>
<paragraph id="8BBF3692FFDFFFB66114FAD4FD1F385F" blockId="3.[96,776,1293,1383]" pageId="3">
mm) and 6b, lateral view (scale bar = 0.3 mm); 7a,
<taxonomicName id="4C004D11FFDFFFB6636BFAD4FD53380F" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[543,661,1317,1335]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="argenteola">
<emphasis id="B974EA80FFDFFFB6636BFAD4FD53380F" box="[543,661,1317,1335]" italics="true" pageId="3">H. argenteola</emphasis>
</taxonomicName>
, frontal view and 7b, lateral view [modified from
<bibRefCitation id="EF914B63FFDFFFB660E9FACCFDAF3877" author="Ratcliffe BC &amp; Deloya, C." box="[413,617,1341,1359]" pageId="3" pagination="161 - 202" refId="ref3534" refString="Ratcliffe BC, Deloya, C. 1992. The biogeography and phylogeny of Hologymnetis (Coleoptera: Scarabaeidae: Cetoniinae) with a revision of the genus. The Coleopterists Bulletin 46 (2): 161 - 202." type="journal article" year="1992">Ratcliffe &amp;Deloya 1992</bibRefCitation>
]; 8a,
<taxonomicName id="4C004D11FFDFFFB663E2FACCFCC13877" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[662,775,1341,1359]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="margaritis">
<emphasis id="B974EA80FFDFFFB663E2FACCFCC13877" box="[662,775,1341,1359]" italics="true" pageId="3">H. margaritis</emphasis>
</taxonomicName>
frontal view and 8b, lateral view [modified from
<bibRefCitation id="EF914B63FFDFFFB66376FAA4FD16385F" author="Ratcliffe BC &amp; Deloya, C." box="[514,720,1365,1383]" pageId="3" pagination="161 - 202" refId="ref3534" refString="Ratcliffe BC, Deloya, C. 1992. The biogeography and phylogeny of Hologymnetis (Coleoptera: Scarabaeidae: Cetoniinae) with a revision of the genus. The Coleopterists Bulletin 46 (2): 161 - 202." type="journal article" year="1992">Ratcliffe &amp; Deloya 1992</bibRefCitation>
].
</paragraph>
</caption>
<paragraph id="8BBF3692FFDFFFB66240FA76FC2538C1" blockId="3.[96,1489,1415,1901]" pageId="3">
<emphasis id="B974EA80FFDFFFB66240FA76FC5438A1" bold="true" box="[820,914,1415,1433]" pageId="3">Figs. 913.</emphasis>
Parameres, 9a,
<taxonomicName id="4C004D11FFDFFFB6656AFA76FB1F38A1" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[1054,1241,1415,1433]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B974EA80FFDFFFB6656AFA76FB1F38A1" box="[1054,1241,1415,1433]" italics="true" pageId="3">Hologymnetis cinerea</emphasis>
</taxonomicName>
, frontal view and 9b, lateral ETYMOLOGY view;10a,
<taxonomicName id="4C004D11FFDFFFB662F7FA6EFC213889" baseAuthorityName="Vigors" baseAuthorityYear="1825" box="[899,999,1439,1457]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B974EA80FFDFFFB662F7FA6EFC213889" box="[899,999,1439,1457]" italics="true" pageId="3">H.undulata</emphasis>
</taxonomicName>
, frontal view and 10b,lateral view; 11a,
<taxonomicName id="4C004D11FFDFFFB6644AFA6EFA543889" authorityName="Ratcliffe &amp; Deloya" authorityYear="1992" box="[1342,1426,1439,1457]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B974EA80FFDFFFB6644AFA6EFA543889" box="[1342,1426,1439,1457]" italics="true" pageId="3">H.moroni</emphasis>
</taxonomicName>
frontal This species is named in honor of Dr. Pedro Reyes Castillo in rec- view and 11b, lateral view; 12a,
<taxonomicName id="4C004D11FFDFFFB66538FA46FB0A38F1" authorityName="Ratcliffe &amp; Deloya" authorityYear="1992" box="[1100,1228,1463,1481]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="vulcanorum">
<emphasis id="B974EA80FFDFFFB66538FA46FB0A38F1" box="[1100,1228,1463,1481]" italics="true" pageId="3">H. vulcanorum</emphasis>
</taxonomicName>
frontal view and 12b, lateral ognition of his important contribution to the
<taxonomicName id="4C004D11FFDFFFB66362FA23FDBD38DE" authorityName="Leach" authorityYear="1815" box="[534,635,1490,1510]" class="Insecta" family="Passalidae" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
of America. view; 13a,
<taxonomicName id="4C004D11FFDFFFB662F1FA3EFC3D38D9" authorityName="Ratcliffe &amp; Deloya" authorityYear="1992" box="[901,1019,1487,1505]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="kinichahau">
<emphasis id="B974EA80FFDFFFB662F1FA3EFC3D38D9" box="[901,1019,1487,1505]" italics="true" pageId="3">H. kinichahau</emphasis>
</taxonomicName>
frontal view and 13b, lateral view [modified from
<bibRefCitation id="EF914B63FFDFFFB664D9FA3EFC1C38C1" author="Ratcliffe BC &amp; Deloya, C." pageId="3" pagination="161 - 202" refId="ref3534" refString="Ratcliffe BC, Deloya, C. 1992. The biogeography and phylogeny of Hologymnetis (Coleoptera: Scarabaeidae: Cetoniinae) with a revision of the genus. The Coleopterists Bulletin 46 (2): 161 - 202." type="journal article" year="1992">Ratcliffe &amp; Deloya 1992</bibRefCitation>
].
</paragraph>
<paragraph id="8BBF3692FFDFFFB66114FA0EFE9E3B2D" blockId="3.[96,1489,1415,1901]" box="[96,344,1535,1557]" pageId="3">TAXONOMIC REMARKS</paragraph>
<paragraph id="8BBF3692FFDFFFB661F0F9DBFCCE3B41" blockId="3.[96,1489,1415,1901]" pageId="3">
<taxonomicName id="4C004D11FFDFFFB661F0F9DBFE8D3B06" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[132,331,1578,1598]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B974EA80FFDFFFB661F0F9DBFE8D3B06" box="[132,331,1578,1598]" italics="true" pageId="3">Hologymnetis reyesi</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C004D11FFDFFFB6636AF9DBFD663B06" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[542,672,1578,1598]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="margaritis">
<emphasis id="B974EA80FFDFFFB6636AF9DBFD663B06" box="[542,672,1578,1598]" italics="true" pageId="3">H. margaritis</emphasis>
</taxonomicName>
. Adults of 7.— Parameres thin at middle, internal margin sinuate, latero-apithese species are externally similar in color, punctures of the frons, cal region rounded and short (
<figureCitation id="133B2A17FFDFFFB665E9F9B2FB153B6F" box="[1181,1235,1603,1623]" captionStart="Figs" captionStartId="3.[108,145,1293,1311]" captionTargetBox="[103,771,197,1244]" captionTargetId="figure-422@3.[96,776,192,1248]" captionTargetPageId="3" captionText="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi="http://doi.org/10.5281/zenodo.12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="3">Fig. 8</figureCitation>
)....
<taxonomicName id="4C004D11FFDFFFB66473F9B2FA093B6F" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[1287,1487,1603,1623]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="margaritis">
<emphasis id="B974EA80FFDFFFB66473F9B2FA413B6F" box="[1287,1415,1603,1623]" italics="true" pageId="3">H. margaritis</emphasis>
(Bates)
</taxonomicName>
pronotum, elytra and pygidium. However, the lighter longitudinal nar-
</paragraph>
<paragraph id="8BBF3692FFDFFFB6625CF982FCCE3BAE" blockId="3.[96,1489,1415,1901]" pageId="3">
7´.— Parameres wide at middle, internal margin straight, latero-apirow band in the pronotum is more strongly delineated in
<taxonomicName id="4C004D11FFDFFFB663F3F973FD1F3BAE" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[647,729,1666,1686]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B974EA80FFDFFFB663F3F973FD1F3BAE" box="[647,729,1666,1686]" italics="true" pageId="3">H. reyesi</emphasis>
</taxonomicName>
. The
</paragraph>
<paragraph id="8BBF3692FFDFFFB6620DF97CFA953A7E" blockId="3.[96,1489,1415,1901]" pageId="3">
cal region with curved, narrow, elongate tooth (
<figureCitation id="133B2A17FFDFFFB6644EF97CFABC3B99" box="[1338,1402,1677,1697]" captionStart="Figs" captionStartId="2.[156,193,1966,1984]" captionTargetBox="[150,821,1202,1924]" captionTargetId="figure-461@2.[144,824,1198,1928]" captionTargetPageId="2" captionText="Figs. 14. Hologymnetis reyesi, 1, habitus, dorsal aspect; 2, habitus, lateral aspect; 3, pygidium, 4a, parameres, frontal view; 4b, parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12769504" httpUri="https://zenodo.org/record/12769504/files/figure.png" pageId="3">Figs. 4</figureCitation>
,
<figureCitation id="133B2A17FFDFFFB664F6F97CFA493B99" box="[1410,1423,1677,1697]" captionStart="Figs" captionStartId="3.[108,145,1293,1311]" captionTargetBox="[103,771,197,1244]" captionTargetId="figure-422@3.[96,776,192,1248]" captionTargetPageId="3" captionText="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi="http://doi.org/10.5281/zenodo.12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="3">6</figureCitation>
)..... parameres of
<taxonomicName id="4C004D11FFDFFFB6619DF96EFEFB3B8B" authorityName="Gasca-Alvarez and Deloya" authorityYear="2015" box="[233,317,1695,1715]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B974EA80FFDFFFB6619DF96EFEFB3B8B" box="[233,317,1695,1715]" italics="true" pageId="3">H. reyesi</emphasis>
</taxonomicName>
are unique among
<taxonomicName id="4C004D11FFDFFFB66089F96EFD123B8B" box="[509,724,1695,1715]" pageId="3">
<emphasis id="B974EA80FFDFFFB66089F96EFD453B8B" box="[509,643,1695,1715]" italics="true" pageId="3">Hologymnetis</emphasis>
species
</taxonomicName>
; they....................
<taxonomicName id="4C004D11FFDFFFB66514F959FA503B84" authority="Gasca-Alvarez &amp; Deloya" authorityName="Gasca-Alvarez &amp; Deloya" authorityYear="2015" box="[1120,1430,1704,1724]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="reyesi" status="n. sp.">
<emphasis id="B974EA80FFDFFFB66514F959FB773B84" box="[1120,1201,1704,1724]" italics="true" pageId="3">H. reyesi</emphasis>
Gasca-Álvarez &amp; Deloya
</taxonomicName>
<emphasis id="B974EA80FFDFFFB664EFF959FA163B84" bold="true" box="[1435,1488,1704,1724]" pageId="3">
<taxonomicNameLabel id="A24757FBFFDFFFB664EFF959FA163B84" box="[1435,1488,1704,1724]" pageId="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
resemble most closely
<taxonomicName id="4C004D11FFDFFFB6604FF94CFE793BE9" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[315,447,1725,1745]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="argenteola">
<emphasis id="B974EA80FFDFFFB6604FF94CFE793BE9" box="[315,447,1725,1745]" italics="true" pageId="3">H. argenteola</emphasis>
</taxonomicName>
but the apicolateral tooth is much more developed and elongated (
<figureCitation id="133B2A17FFDFFFB660E8F92BFE153BD6" box="[412,467,1754,1774]" captionStart="Figs" captionStartId="3.[108,145,1293,1311]" captionTargetBox="[103,771,197,1244]" captionTargetId="figure-422@3.[96,776,192,1248]" captionTargetPageId="3" captionText="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi="http://doi.org/10.5281/zenodo.12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="3">Fig. 7</figureCitation>
). This key is mainly made with coloration characters of the body, In the
<bibRefCitation id="EF914B63FFDFFFB661B3F906FE793A33" author="Ratcliffe BC &amp; Deloya, C." box="[199,447,1783,1803]" pageId="3" pagination="161 - 202" refId="ref3534" refString="Ratcliffe BC, Deloya, C. 1992. The biogeography and phylogeny of Hologymnetis (Coleoptera: Scarabaeidae: Cetoniinae) with a revision of the genus. The Coleopterists Bulletin 46 (2): 161 - 202." type="journal article" year="1992">Ratcliffe &amp; Deloya (1992)</bibRefCitation>
key,
<taxonomicName id="4C004D11FFDFFFB66081F906FD8C3A33" authorityName="Gasca-Alvarez &amp; Deloya" authorityYear="2015" box="[501,586,1783,1803]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="reyesi">
<emphasis id="B974EA80FFDFFFB66081F906FD8C3A33" box="[501,586,1783,1803]" italics="true" pageId="3">H. reyesi</emphasis>
</taxonomicName>
will key out in the which could contain intraspecific variation. We propose a new identi- final couplet, where it is compared with
<taxonomicName id="4C004D11FFDFFFB66085F8E5FDB43A10" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[497,626,1812,1832]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="margaritis">
<emphasis id="B974EA80FFDFFFB66085F8E5FDB43A10" box="[497,626,1812,1832]" italics="true" pageId="3">H. margaritis</emphasis>
</taxonomicName>
by the form of fication key for current
<taxonomicName id="4C004D11FFDFFFB66578F8E4FB193A11" box="[1036,1247,1813,1833]" pageId="3">
<emphasis id="B974EA80FFDFFFB66578F8E4FB543A11" box="[1036,1170,1813,1833]" italics="true" pageId="3">Hologymnetis</emphasis>
species
</taxonomicName>
based on morphological parameres: characters, with emphasis on the form of the parameres:
</paragraph>
<paragraph id="8BBF3692FFDFFFB6632CF8A8FC1E3A55" blockId="3.[96,1489,1415,1901]" box="[600,984,1881,1901]" pageId="3">
Key to adults of
<taxonomicName id="4C004D11FFDFFFB66381F8A8FC1E3A55" authority="Martinez" authorityName="Martinez" authorityYear="1949" box="[757,984,1881,1901]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA80FFDFFFB66381F8A8FCBD3A55" box="[757,891,1881,1901]" italics="true" pageId="3">Hologymnetis</emphasis>
Martínez
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C31A6519FFDFFFB16114F878F9C63ED7" lastPageId="4" pageId="3" type="key">
<key id="FF9161A3FFDFFFB16114F878F9C63ED7" lastPageId="4" pageId="3">
<keyStep id="30F42E37FFDFFFB66114F878FA163AE6" pageId="3">
<paragraph id="8BBF3692FFDFFFB66114F878FA163AA5" blockId="3.[96,1488,1929,2014]" box="[96,1488,1929,1949]" pageId="3">
<keyLead id="30F195A7FFDFFFB66114F878FA163AA5" box="[96,1488,1929,1949]" pageId="3">1.— Venter largely glossy black in both sexes, sternites laterally ofen with cretaceous (chalky white) marking........................ 2</keyLead>
</paragraph>
<paragraph id="8BBF3692FFDFFFB66114F85EFA163AE6" blockId="3.[96,1488,1929,2014]" pageId="3">
<keyLead id="30F195A7FFDFFFB66114F85EFA163AE6" pageId="3">1.— Venter brown to grayish brown or pale brassy green or blue, weakly glossy or weakly to strongly metallic; females with central third of sternites bare, piceous............................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F42E37FFD8FFB161E4FF32F9C63C6F" pageId="4">
<paragraph id="8BBF3692FFD8FFB161E4FF32F9C63DC9" blockId="4.[144,1536,195,241]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FF32F9C63DC9" pageId="4">
2.— Dorsum unicolorous or black, rarely strongly punctuate. Parameres elongate, apex rounded, lateral lobes formed into teeth directed posteriorly (Fig. 9)..................................................................................
<taxonomicName id="4C004D11FFD8FFB1643EFF2CFA6A3DC9" ID-CoL="3M8WQ" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[1354,1452,221,241]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B974EA80FFD8FFB1643EFF2CFA6A3DC9" box="[1354,1452,221,241]" italics="true" pageId="4">H. cinerea</emphasis>
</taxonomicName>
(G. &amp; P.)
</keyLead>
</paragraph>
<paragraph id="8BBF3692FFD8FFB161E4FEFCF9C63C6F" blockId="4.[144,1537,269,343]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FEFCF9C63C6F" pageId="4">
2.— Dorsum black or piceous patterned with cretaceous lines; lines present longitudinally on pronotum and radiating from midline on elytra; occasionally reduced or nearly absent. Parameres elongated, quadrangular, without lateral lobes, apex slightly curved (Fig. 10)........................................................................................................
<taxonomicName id="4C004D11FFD8FFB1644AFEB2F9C63C6F" ID-CoL="3M8WV" authority="(Vigors)" baseAuthorityName="Vigors" baseAuthorityYear="1825" box="[1342,1536,323,343]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B974EA80FFD8FFB1644AFEB2FA763C6F" box="[1342,1456,323,343]" italics="true" pageId="4">H. undulata</emphasis>
(Vigors)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F42E37FFD8FFB161E4FE83F9C63C8E" pageId="4">
<paragraph id="8BBF3692FFD8FFB161E4FE83F9C63CBE" blockId="4.[144,1536,370,390]" box="[144,1536,370,390]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FE83F9C63CBE" box="[144,1536,370,390]" pageId="4">3.— Pronotum with a lighter, longitudinal, narrow band at center............................................................. 4</keyLead>
</paragraph>
<paragraph id="8BBF3692FFD8FFB161E4FE53F9C63C8E" blockId="4.[144,1536,418,438]" box="[144,1536,418,438]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FE53F9C63C8E" box="[144,1536,418,438]" pageId="4">3´.— Longitudinal, narrow band at center of pronotum absent................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F42E37FFD8FFB161E4FE23F9C63F08" pageId="4">
<paragraph id="8BBF3692FFD8FFB161E4FE23F9C63F39" blockId="4.[144,1536,466,513]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FE23F9C63F39" pageId="4">
4.— Bead of lateral margin of pronotum complete. Elytra unicolorous. Protibia with median tooth distinctly closer to apical tooth. Parameres elongate, thin at middle; apicolateral teeth developed (Fig. 11)....................................
<taxonomicName id="4C004D11FFD8FFB1659AFE1CF9C63F39" ID-CoL="3M8WT" authority="Ratcliffe &amp; Deloya" authorityName="Ratcliffe &amp; Deloya" authorityYear="1992" box="[1262,1536,493,513]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B974EA80FFD8FFB1659AFE1CFA883F39" box="[1262,1358,493,513]" italics="true" pageId="4">H. moroni</emphasis>
Ratcliffe &amp; Deloya
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF3692FFD8FFB161E4FDEDF9C63F08" blockId="4.[144,1536,540,560]" box="[144,1536,540,560]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FDEDF9C63F08" box="[144,1536,540,560]" pageId="4">4.— Bead of lateral margin of pronotum incomplete. Elytra pattern with darkening of costae. Protibia with teeth subequally spaced..... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F42E37FFD8FFB161E4FDBDF9C63FFD" pageId="4">
<paragraph id="8BBF3692FFD8FFB161E4FDBDF9C63F43" blockId="4.[144,1536,588,635]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FDBDF9C63F43" pageId="4">
5.— Mesometasternal process with glossy apex usually sharply and transversely delineated from shaf. Mesotibial carina present at middle of tibia. Parameres with apical, median ridges that taper to a point (Fig. 12)......................
<taxonomicName id="4C004D11FFD8FFB165B5FD96F9C63F43" ID-CoL="3M8WW" authority="Ratcliffe &amp; Deloya" authorityName="Ratcliffe &amp; Deloya" authorityYear="1992" box="[1217,1536,615,635]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="vulcanorum">
<emphasis id="B974EA80FFD8FFB165B5FD96FA883F43" box="[1217,1358,615,635]" italics="true" pageId="4">H. vulcanorum</emphasis>
Ratcliffe &amp; Deloya
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF3692FFD8FFB161E4FD66F9C63FFD" blockId="4.[144,1536,663,709]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FD66F9C63FFD" pageId="4">5.— Mesometasternal process with glossy apex and shaf usually broadly joined by a broad, piceous region. Mesotibial carina absent. Parameres not as above............................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F42E37FFD8FFB161E4FD10F9C73E62" pageId="4">
<paragraph id="8BBF3692FFD8FFB161E4FD10F9C63E28" blockId="4.[144,1536,737,784]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FD10F9C63E28" pageId="4">
6.— Pygidium with moderate to dense C-shaped to elongate punctures. Parameres thin at middle, internal margin sinuated, latero-apical region rounded, short (
<figureCitation id="133B2A17FFD8FFB160B5FD0DFE313E28" box="[449,503,764,784]" captionStart="Figs" captionStartId="3.[108,145,1293,1311]" captionTargetBox="[103,771,197,1244]" captionTargetId="figure-422@3.[96,776,192,1248]" captionTargetPageId="3" captionText="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi="http://doi.org/10.5281/zenodo.12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="4">Fig. 8</figureCitation>
).......................................................................
<taxonomicName id="4C004D11FFD8FFB1644CFD0DF9C63E28" ID-CoL="3M8WS" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[1336,1536,764,784]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="margaritis">
<emphasis id="B974EA80FFD8FFB1644CFD0DFA713E28" box="[1336,1463,764,784]" italics="true" pageId="4">H. margaritis</emphasis>
(Bates)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF3692FFD8FFB161E4FCDDF9C73E62" blockId="4.[144,1537,812,858]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FCDDF9C73E62" pageId="4">
6´.— Pygidium with punctures kidney-shaped to elongate. Parameres wide at middle, internal margin straight, latero-apical region with narrow, elongateed tooth (
<figureCitation id="133B2A17FFD8FFB160B7FCB7FDC23E62" box="[451,516,838,858]" captionStart="Figs" captionStartId="2.[156,193,1966,1984]" captionTargetBox="[150,821,1202,1924]" captionTargetId="figure-461@2.[144,824,1198,1928]" captionTargetPageId="2" captionText="Figs. 14. Hologymnetis reyesi, 1, habitus, dorsal aspect; 2, habitus, lateral aspect; 3, pygidium, 4a, parameres, frontal view; 4b, parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12769504" httpUri="https://zenodo.org/record/12769504/files/figure.png" pageId="4">Figs. 4</figureCitation>
,
<figureCitation id="133B2A17FFD8FFB16379FCB7FDDC3E62" box="[525,538,838,858]" captionStart="Figs" captionStartId="3.[108,145,1293,1311]" captionTargetBox="[103,771,197,1244]" captionTargetId="figure-422@3.[96,776,192,1248]" captionTargetPageId="3" captionText="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi="http://doi.org/10.5281/zenodo.12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="4">6</figureCitation>
).....................................................
<taxonomicName id="4C004D11FFD8FFB165F3FCB7FA033E62" ID-CoL="6M9YN" authority="Gasca-Alvarez &amp; Deloya" authorityName="Gasca-Alvarez &amp; Deloya" authorityYear="2015" box="[1159,1477,838,858]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="reyesi" status="n. sp.">
<emphasis id="B974EA80FFD8FFB165F3FCB7FB1D3E62" box="[1159,1243,838,858]" italics="true" pageId="4">H. reyesi</emphasis>
Gasca-Álvarez &amp; Deloya
</taxonomicName>
<emphasis id="B974EA80FFD8FFB164BEFCB7F9C73E62" bold="true" box="[1482,1537,838,858]" pageId="4">
<taxonomicNameLabel id="A24757FBFFD8FFB164BEFCB7F9C73E62" box="[1482,1537,838,858]" pageId="4" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F42E37FFD8FFB161E4FC87F9C63ED7" pageId="4">
<paragraph id="8BBF3692FFD8FFB161E4FC87F9C63E9D" blockId="4.[144,1536,886,933]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FC87F9C63E9D" pageId="4">
7.— Clypeus slightly convex on disc; apex broadly reflexed, distinctly emarginate at center. Elytral punctation relatively large, punctures usually U-shaped. Parameres short, stout, apex quadrangular (Fig. 13)..............................
<taxonomicName id="4C004D11FFD8FFB165BEFC60F9C63E9D" ID-CoL="3M8WR" authority="Ratcliffe &amp; Deloya" authorityName="Ratcliffe &amp; Deloya" authorityYear="1992" box="[1226,1536,913,933]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="kinichahau">
<emphasis id="B974EA80FFD8FFB165BEFC60FA883E9D" box="[1226,1358,913,933]" italics="true" pageId="4">H. kinichahau</emphasis>
Ratcliffe &amp; Deloya
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF3692FFD8FFB161E4FC31F9C63ED7" blockId="4.[144,1536,960,1007]" pageId="4">
<keyLead id="30F195A7FFD8FFB161E4FC31F9C63ED7" pageId="4">
7´.— Clypeus weakly tumescent at center; apex strongly reflexed, usually strongly arcuate. Elytral punctation greatly reduced. Parameres curved, apicolateral teeth elongated and strongly narrowed (
<figureCitation id="133B2A17FFD8FFB16259FC2AFCA53ED7" box="[813,867,987,1007]" captionStart="Figs" captionStartId="3.[108,145,1293,1311]" captionTargetBox="[103,771,197,1244]" captionTargetId="figure-422@3.[96,776,192,1248]" captionTargetPageId="3" captionText="Figs. 68. Parameres, 6a, Hologymnetis reyesi, frontal view (scale bar = 0.3 mm) and 6b, lateral view (scale bar = 0.3 mm); 7a, H. argenteola, frontal view and 7b, lateral view [modified from Ratcliffe &amp;Deloya 1992]; 8a, H. margaritis frontal view and 8b, lateral view [modified from Ratcliffe &amp; Deloya 1992]." figureDoi="http://doi.org/10.5281/zenodo.12769508" httpUri="https://zenodo.org/record/12769508/files/figure.png" pageId="4">Fig. 7</figureCitation>
).......................................
<taxonomicName id="4C004D11FFD8FFB16446FC2AF9C63ED7" ID-CoL="3M8WP" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1889" box="[1330,1536,987,1007]" class="Insecta" family="Scarabaeidae" genus="Hologymnetis" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="argenteola">
<emphasis id="B974EA80FFD8FFB16446FC2AFA713ED7" box="[1330,1463,987,1007]" italics="true" pageId="4">H. argenteola</emphasis>
(Bates)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>