134 lines
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134 lines
17 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.132.1508" ID-GBIF-Dataset="4e1ed198-07d4-4297-acfb-574a27c10709" ID-PMC="PMC3208432" ID-Pensoft-Pub="1313-2970-132-33" ID-PubMed="22140332" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-132-33" ModsDocOrigin="ZooKeys 132" ModsDocTitle="Taxonomic remarks about Semiclivina (Kult, 1947) new status, with description of Uroclivina subgen. n., and of two new species from South America (Coleoptera, Carabidae, Scaritinae, Clivinini)" checkinTime="1451249827898" checkinUser="pensoft" docAuthor="Dostal, Alexander" docDate="2011" docId="DBBA8CEB4C8CC0771367CAF57D066A20" docLanguage="en" docName="ZooKeys 132: 33-50" docOrigin="ZooKeys 132" docSource="http://dx.doi.org/10.3897/zookeys.132.1508" docTitle="Semiclivina Kult 1947, stat. n." docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="FFE3FFACFFB0FF82FFAC0B32C709FFA3" masterDocTitle="Taxonomic remarks about Semiclivina (Kult, 1947) new status, with description of Uroclivina subgen. n., and of two new species from South America (Coleoptera, Carabidae, Scaritinae, Clivinini)" masterLastPageNumber="50" masterPageNumber="33" pageNumber="34" updateTime="1668152314760" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic remarks about Semiclivina (Kult, 1947) new status, with description of Uroclivina subgen. n., and of two new species from South America (Coleoptera, Carabidae, Scaritinae, Clivinini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dostal, Alexander</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>132</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>50</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.132.1508</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.132.1508</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-132-33</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152031188" LSID="urn:lsid:zoobank.org:act:581B1ABC-14ED-4D8B-B216-7119CF90DCDC" httpUri="http://treatment.plazi.org/id/DBBA8CEB4C8CC0771367CAF57D066A20" lastPageId="3" lastPageNumber="35" pageId="1" pageNumber="34">
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<subSubSection pageId="1" pageNumber="34" type="nomenclature">
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<paragraph pageId="1" pageNumber="34">
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Genus
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<taxonomicName LSID="http://species-id.net/wiki/Semiclivina" authority="Kult, 1947" authorityName="Kult" authorityYear="1947" class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Semiclivina Kult, 1947</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="34">stat. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="34" type="reference_group">
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<paragraph pageId="1" pageNumber="34">
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<taxonomicName class="Insecta" family="Carabidae" genus="Clivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Clivina</taxonomicName>
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Subgenus
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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<bibRefCitation author="Kult, K" journalOrPublisher="Spolecnosti Entomologicke)" pageId="11" pageNumber="44" pagination="26 - 37" title="Treti studie o strevlicich tribu Clivini (Col.). The 3 rd study to the knowledge of tribus Clivinini (Col., Carab.). (18 th Contribution to the knowledge of Carabidae.) Acta Societatis Entomologicae Cechoslovakiae (Casopis Csl." volume="44" year="1947">Kult 1947</bibRefCitation>
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: 31-32; Reichardt 1977: 391;
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<bibRefCitation pageId="1" pageNumber="34">Nichols 1988a</bibRefCitation>
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: 154;
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<bibRefCitation pageId="1" pageNumber="34">1988b</bibRefCitation>
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: 91;
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<bibRefCitation author="Ball, GE" journalOrPublisher="Osaka" pageId="10" pageNumber="43" pagination="129 - 156" title="The subgenera of Clivina Latreille in the Western Hemisphere, and a Revision of Subgenus Antroforceps Barr (new status), with Notes about Evolutionary Aspects (Coleoptera: Carabidae: Clivinini). Spec. Publ. Japan Coleopt. Soc." volume="1" year="2001">Ball 2001</bibRefCitation>
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: 136;
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae), Second edition, typeset and printed by the author, Tutzing" pageId="11" pageNumber="44" title="Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
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: 145;
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="10" pageNumber="43" pagination="1 - 220" title="The Australian Clivinini 1. The genera Ancus Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara Sloane, Syleter Andrewes, the subgenera Paraclivina Kult, Semiclivina Kult, and the atrata-, biplagiata-, brevicornis-, coronata-, coryzoides-, cribrosa-, debilis-, denticollis-, grandiceps-, incerta-, lobata-, obliquata-, obsoleta-, orbitalis-, planiceps-, sulcaticeps-, tranquebarica-, and wurargae-groups of the genus Clivina Latreille. With a note on a record of the genus Parathlibops Basilewsky (Scapterini) (Carabidae, Scaritinae)." volume="12" year="2008">Baehr 2008</bibRefCitation>
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: 23;
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 41.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="36" pageId="1" pageNumber="34" type="type species">
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<paragraph pageId="1" pageNumber="34">Type Species.</paragraph>
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<paragraph pageId="1" pageNumber="34">
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<taxonomicName class="Insecta" family="Carabidae" genus="Clivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clivina dentipes" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="dentipes">Clivina dentipes</taxonomicName>
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Dejaen, 1825, by original designation:
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<bibRefCitation author="Kult, K" journalOrPublisher="Spolecnosti Entomologicke)" pageId="11" pageNumber="44" pagination="26 - 37" title="Treti studie o strevlicich tribu Clivini (Col.). The 3 rd study to the knowledge of tribus Clivinini (Col., Carab.). (18 th Contribution to the knowledge of Carabidae.) Acta Societatis Entomologicae Cechoslovakiae (Casopis Csl." volume="44" year="1947">Kult 1947</bibRefCitation>
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: 31.
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</paragraph>
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<paragraph lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
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The genus
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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(Kult, 1947) is readily recognized by the sculptured band of the proepisternum, extended more or less parallel to the proepisternal margin in the basal part of the proepisternum, curved inward in the anterior part, ended at the anterior end of the proepisternal-prosternal suture (Fig. 1). This structure was differently interpreted by diverse authors (
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 38) as furrow, ridge or carina or elongate striole (
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="10" pageNumber="43" pagination="1 - 220" title="The Australian Clivinini 1. The genera Ancus Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara Sloane, Syleter Andrewes, the subgenera Paraclivina Kult, Semiclivina Kult, and the atrata-, biplagiata-, brevicornis-, coronata-, coryzoides-, cribrosa-, debilis-, denticollis-, grandiceps-, incerta-, lobata-, obliquata-, obsoleta-, orbitalis-, planiceps-, sulcaticeps-, tranquebarica-, and wurargae-groups of the genus Clivina Latreille. With a note on a record of the genus Parathlibops Basilewsky (Scapterini) (Carabidae, Scaritinae)." volume="12" year="2008">Baehr 2008</bibRefCitation>
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: 23). In fact it is a slightly elevated band like structure consisting of
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<pageBreakToken pageId="2" pageNumber="35" start="start">very</pageBreakToken>
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fine, regular transverse ridges. This sculptured band (
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 38) is part of a stridulation organ in the sense of a pars stridens, whereas the corresponding plectrum is a fine, sharp longitudinal ridge, just above the distal part of the lower inner edge of the profemur (Fig. 2). In some species the outer edge of the profemur is produced to a remarkable denticle, but this structure does not interact with the stridulation organ in any way. The function of this stridulation organ has not been observed so far.
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</paragraph>
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<caption pageId="2" pageNumber="35">
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<paragraph pageId="2" pageNumber="35">
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Figure 1. Photograph of prothorax and base of metathorax of
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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species; right lateral view, showing the stridulation ridge (arrow); Scale bar: 0,5 mm.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="35">
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<paragraph pageId="2" pageNumber="35">
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Figure 2. Photograph of right leg, posterior view, of base of trochanter, femur, and base of tibia of
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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species, showing stridulation ridge on inner side of profemur (arrow); Scale bar: 0,5 mm.
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</paragraph>
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</caption>
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<paragraph lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
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This organ is not unique within the Tribe
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<taxonomicName lsidName="" pageId="2" pageNumber="35" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
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: exactly the same structures (the stridulation ridge on proepisterna and the ridge on the distal part of the inner edge of the profemur) together with other features separates the ardistomine genus
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<taxonomicName class="Insecta" family="Carabidae" genus="Ardistomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ardistomis" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Ardistomis</taxonomicName>
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Putzeys, 1846 from
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiardistomis" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiardistomis</taxonomicName>
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Kult, 1950, but it is unique within
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<taxonomicName class="Insecta" family="Carabidae" genus="Clivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Clivina</taxonomicName>
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(sensu lato), forming a very isolated, distinct group, restricted to the western hemisphere and to a single species from Australia (New South Wales) described by
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="10" pageNumber="43" pagination="1 - 220" title="The Australian Clivinini 1. The genera Ancus Putzeys, Aspidoglossa Putzeys, Clivinarchus Sloane, Platysphyrus Sloane, Pseudoclivina Kult, Rhysocara Sloane, Syleter Andrewes, the subgenera Paraclivina Kult, Semiclivina Kult, and the atrata-, biplagiata-, brevicornis-, coronata-, coryzoides-, cribrosa-, debilis-, denticollis-, grandiceps-, incerta-, lobata-, obliquata-, obsoleta-, orbitalis-, planiceps-, sulcaticeps-, tranquebarica-, and wurargae-groups of the genus Clivina Latreille. With a note on a record of the genus Parathlibops Basilewsky (Scapterini) (Carabidae, Scaritinae)." volume="12" year="2008">Baehr 2008</bibRefCitation>
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: 23-25. Within the tribe
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<taxonomicName lsidName="" pageId="2" pageNumber="35" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
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no other genera than
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<taxonomicName class="Insecta" family="Carabidae" genus="Ardistomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ardistomis" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Ardistomis</taxonomicName>
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Putzeys, 1846 and
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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(Kult, 1947) carry a stridulation organ on the proepisternum. Beside this,
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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is characterized by a distal spine-like projection at the outer end of the mesotibia, which is small in some taxa, but clearly identifiable by the apical seta
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<pageBreakToken pageId="3" pageNumber="36" start="start">excentrically</pageBreakToken>
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inserted, in contradiction to a tubercle, which carries the seta at the top of the tubercle (see also
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<bibRefCitation author="Ball, GE" journalOrPublisher="Osaka" pageId="10" pageNumber="43" pagination="129 - 156" title="The subgenera of Clivina Latreille in the Western Hemisphere, and a Revision of Subgenus Antroforceps Barr (new status), with Notes about Evolutionary Aspects (Coleoptera: Carabidae: Clivinini). Spec. Publ. Japan Coleopt. Soc." volume="1" year="2001">Ball 2001</bibRefCitation>
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: 136, 140;
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="10" pageNumber="43" pagination="37 - 48" title="Rediscovery of Clivina morio Dejaen with the description of Leucocara, a subgen. n. of Clivina Latreille (Coleoptera, Carabidae, Clivinini)." url="10.3897/zookeys.25.276" volume="25" year="2009">Bousquet 2009</bibRefCitation>
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: 41, 43); usually five setae in elytral interval 3 (including preapical puncture); elytral striae 1 to 5 free at base (between humeral channel and suture, the basal keel of some species and the flat tubercles at the end of intervals 2 to 5 do not limit this definition); profemur with a sometimes small dentiform projection at the outer edge toward apex; anal ventrite (abdominal sternum VII) with two marginal punctures at each side relatively close together. The species included in
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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are listed in Bousquet (2009: 39-41), where the assignment of some species remains uncertain. It is most likely, that species not included in the subgen. n.
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<taxonomicName class="Insecta" family="Carabidae" genus="Uroclivina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Uroclivina" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Uroclivina</taxonomicName>
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comprise a more or less heterogeneous group which form a probably paraphyletic group, defined as
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<taxonomicName class="Insecta" family="Carabidae" genus="Semiclivina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Semiclivina" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="genus">Semiclivina</taxonomicName>
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(s. str.). This subgenus will be the objective of further investigations in the future.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |