treatments-xml/data/55/97/5E/55975EEF91FB5DAE81D55462BA44C00E.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="55975EEF91FB5DAE81D55462BA44C00E" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana parvula Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="28F7FF73-3770-449C-90E5-AF6D321B0348" docUuidSource="ZooBank" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
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<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-20</mods:number>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e81961</mods:url>
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<taxonomicName LSID="https://zoobank.org/28F7FF73-3770-449C-90E5-AF6D321B0348" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Coarctana parvula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="parvula" status="sp. nov.">Coarctana parvula</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">Figs 25</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31M, N</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Pygofer (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25G</figureCitation>
) strongly tapered apically, subacute. Subgenital plate (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25H</figureCitation>
) with filiform setae near outer margin. Aedeagus (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25L, M</figureCitation>
) apodemal process slender and tapered, external surface bearing a small process near mid-length; shaft with pair of subapical processes short, directed dorsally and pair of apical processes curved ventrally, with half length of the subapical processes.
</paragraph>
<caption doi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79" start="Figure 25" startId="F25">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 25.</emphasis>
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana parvula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="parvula">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana parvula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male. 
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing. 
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 8.8 mm; paratypes, males (n = 5) 8.5-8.8 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in frontal view (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25B</figureCitation>
), frons 1.1 
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wider than long; frontogenal suture distant from eye margin by approximately the width of scape; clypeus very wide, 1.1 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins straight and parallel near base and apically strongly convergent; apex straight. Head, in lateral view (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25C</figureCitation>
), crown-face transition thin, with 3 transverse carinae; clypeus slightly inflated. Profemur AV row with 5-6 setae; PV row with 2 setae. Protibia PD row with 5 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 24-26, 13 and 13-15 macrosetae respectively. Metatarsomere I elongated 4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 7-8 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25A, B</figureCitation>
, 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31M, N</figureCitation>
) reddish-brown. Head (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocellus. Face (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25B</figureCitation>
) yellowish without maculae. Pronotum (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25A</figureCitation>
) with black punctures; pair of circular black spots, near anterior margin, behind eye; proepimeron (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25C</figureCitation>
), with narrow black band below pronotal carina. Forewing (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25D</figureCitation>
) with black maculae on apex of anal veins, cross veins of discal and apical cells. Metatibia (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25E</figureCitation>
) 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin slightly rounded. Valve (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25F</figureCitation>
) 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterior margin tapered and rounded. Pygofer, in lateral view (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25G</figureCitation>
), 2 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process thin and long, extending ventrally to almost half height of lateral lobe; anteroventral margin rounded; posterodorsal margin straight; posteroventral margin excavated; apex strongly tapered, subacute. Subgenital plate, in lateral view (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25G</figureCitation>
), short, not reaching pygofer apex; in ventral view (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25H</figureCitation>
), elongated, 3.7 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near half length; dorsal surface with filiform setae near outer margin; lateral margins slightly rounded; apex rounded. Connective (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25I</figureCitation>
) wider than long; anterior margin moderately excavated; dorsal keel and stem reduced. Style, in dorsal view (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25J</figureCitation>
), with outer lobe small and truncated; in lateral view (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25K</figureCitation>
), with blade slightly wider near base and tapered toward apex; dorsal and ventral margins slightly sinuate; ventral margin with apical third serrated; apex truncated, forming a small dorsal process. Aedeagus (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25L, M</figureCitation>
) preatrium not developed; dorsal apodeme with dorsal margin slightly excavated, lateral margins produced ventrolaterally; apodemal process slender and tapered toward apex, curved in right angle near base, external surface bearing small process near mid-length; apex acute, curved anterad; shaft cylindrical, very long and thin, strongly curved dorsally near base and slightly curved anterad at apical third; apex with pair of subapical processes short, directed dorsally and pair of apical processes curved ventrally, with half length of the subapical processes. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female">
<paragraph pageId="0" pageNumber="79">Female.</paragraph>
<paragraph pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2012-08-23" collectorName="R. R. Cavichioli" country="Peru" elevation="985" latitude="-13.3550005" location="Estrada" longLatPrecision="21" longitude="-70.88944" municipality="Peru" specimenCount="1" stateProvince="Cuzco" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male: 
<collectingCountry name="Peru">Peru</collectingCountry>
</emphasis>
: &quot; 
<collectingMunicipality>Peru</collectingMunicipality>
, 
<collectingRegion country="Peru" name="Cusco">Cuzco</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:BB1F8DE72D1C60C4F80A7D0CC6300286" country="Peru" latitude="-13.3550005" longLatPrecision="21" longitude="-70.88944" municipality="Peru" name="Estrada" stateProvince="Cuzco">Estrada</location>
30 C\ 
<locationDeviation location="Quincemil">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.6" unit="km" value="26.0">26km</quantity>
W Quincemil
</locationDeviation>
, 
<collectingDate value="2012-08-23">23.viii.2012</collectingDate>
, (light)\ 
<geoCoordinate degrees="13" direction="south" minutes="21" orientation="latitude" precision="15" seconds="18" value="-13.3550005">13°21′18″S</geoCoordinate>
<geoCoordinate degrees="70" direction="west" minutes="53" orientation="longitude" precision="15" seconds="22" value="-70.88944">70°53′22″W</geoCoordinate>
,\ 
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">985m</elevation>
</quantity>
<collectorName>R.R. Cavichioli</collectorName>
leg.&quot; (MUSM)
</materialsCitation>
. 
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="R. R. Cavichioli" country="Peru" elevation="985" latitude="-13.3550005" location="Estrada" longLatPrecision="21" longitude="-70.88944" municipality="Peru" specimenCount="4" specimenCount-male="4" stateProvince="Cusco" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Paratypes</typeStatus>
</emphasis>
: 
<specimenCount type="male">4♂</specimenCount>
, same data as holotype (DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="2009-02-06" collectorName="ME Irwin, GA" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Taporoto Rd" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<collectingCounty>Peru</collectingCounty>
: Dept. 
<collectingRegion country="Peru" name="Amazonas">Amazonas</collectingRegion>
,\ Distr. 
<normalizedToken originalValue="Águas">Aguas</normalizedToken>
Verdes,\ 
<collectingMunicipality>Bagua</collectingMunicipality>
\ 
<location LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:A0B9D47CDA8F2938CF50A1AF908300F3" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Taporoto Rd" stateProvince="Amazonas">Taporoto Rd</location>
(5N)\ at 
<location LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:C19532E6EF3599AEA66C88961DCD0B5E" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Km" stateProvince="Amazonas">Km</location>
403; 
<location LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:820782B2246A3D5B7B68690F2B3E76D7" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Malaise" stateProvince="Amazonas">Malaise</location>
\ 30.I-\ 
<collectingDate value="2009-02-06">6.II.2009</collectingDate>
; 
<collectorName>ME Irwin, GA</collectorName>
\ 
<location LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:720241BEFE040AF1B417F2F4D7CD8A32" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Amaya" stateProvince="Amazonas">Amaya</location>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">1125m</elevation>
</quantity>
;\ 
<geoCoordinate degrees="05" direction="south" minutes="41" orientation="latitude" precision="15" seconds="23" value="-5.689722">05°41′23″S</geoCoordinate>
<geoCoordinate degrees="77" direction="west" minutes="38" orientation="longitude" precision="15" seconds="13" value="-77.63694">77°38′13″W</geoCoordinate>
(DZRJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The new species name refers to the aedeagus with apodemal processes bearing very small projections at the mid-length of external.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana parvula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="parvula">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana parvula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
(Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
) was strongly supported (SR = 92) as sister group to 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. apena" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="apena">
<emphasis italics="true" pageId="0" pageNumber="79">C. apena</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
+ 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. occultata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="occultata">
<emphasis italics="true" pageId="0" pageNumber="79">C. occultata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
This clade was supported by three synapomorphies: 26:0, clypeus wider than long; 27:2, clypeus with lateral margins strongly convergent apically; 29:1, clypeus tumid in lateral view; and three homoplastic characters: 9:1, crown-face transition with three carinae; 31:1 absence of black irregular maculae near anterior margin of pronotum; and 94:1, pygofer with basodorsal processes strongly developed. 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana oricula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="oricula">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana oricula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
can be differentiated from 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. apena" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="apena">
<emphasis italics="true" pageId="0" pageNumber="79">C. apena</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
and 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. occultata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="occultata">
<emphasis italics="true" pageId="0" pageNumber="79">C. occultata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
by the apodemal processes of aedeagus (Fig. 
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25L, M</figureCitation>
) with a small process near mid-length of external surface, and the shaft with a pair of subapical processes short, directed dorsally and a pair of apical processes curved ventrally.
</paragraph>
</subSubSection>
</treatment>
</document>