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<mods:title id="656E92A6B65C5054677BE29FD966AE48">Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species.</mods:title>
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<taxonomicName id="808789A83C30FFA1FC37FCA3317DFC99" ID-CoL="9939M" authority="(Holthuis, 1979)" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[919,1383,829,854]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tenebrosa">
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<emphasis id="75F32E393C30FFA1FC37FCA33082FC99" bold="true" box="[919,1176,829,854]" italics="true" pageId="2" pageNumber="3">Borneosa tenebrosa</emphasis>
(
<bibRefCitation id="23168FDA3C30FFA1FB08FCA3317AFC99" author="Holthuis LB" box="[1192,1376,829,854]" pageId="2" pageNumber="3" pagination="3 - 47" refId="ref17606" refString="Holthuis LB. 1979. Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zool Verh 171: 3 - 47, pls. 1 - 8." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
)
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(
<figureCitation id="DFBCEEAE3C30FFA1FC45FCC3302AFCBA" box="[997,1072,861,885]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="2" pageNumber="3">Figs. 2</figureCitation>
, 
<figureCitation id="DFBCEEAE3C30FFA1FB9BFCC33050FCBA" box="[1083,1098,861,885]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="2" pageNumber="3">3</figureCitation>
, 
<figureCitation id="DFBCEEAE3C30FFA1FBF6FCC3309DFCBA" box="[1110,1159,861,885]" captionStart="Fig" captionStartId="25.[136,168,1870,1889]" captionTargetBox="[136,1413,729,1841]" captionTargetId="figure-349@25.[136,1413,729,1841]" captionTargetPageId="25" captionText="Fig. 19. Male sternopleonal cavity and pleonal locking tubercle on sternite 5. A, Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; C, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; D, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074568" httpUri="https://zenodo.org/record/8074568/files/figure.png" pageId="2" pageNumber="3">19A</figureCitation>
, 
<figureCitation id="DFBCEEAE3C30FFA1FB31FCC330FAFCBA" box="[1169,1248,861,885]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng &amp; Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="2" pageNumber="3">20A–L</figureCitation>
, 
<figureCitation id="DFBCEEAE3C30FFA1FB4AFCC33103FCBA" box="[1258,1305,861,885]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="2" pageNumber="3">23A</figureCitation>
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<emphasis id="75F32E393C30FFA1FCF2FC06305CFC64" box="[850,1094,920,939]" italics="true" pageId="2" pageNumber="3">Sundathelphusa tenebrosa</emphasis>
<bibRefCitation id="23168FDA3C30FFA1FBEFFC06311BFC64" author="Holthuis LB" box="[1103,1281,920,939]" pageId="2" pageNumber="3" pagination="3 - 47" refId="ref17606" refString="Holthuis LB. 1979. Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zool Verh 171: 3 - 47, pls. 1 - 8." type="journal article" year="1979">Holthuis, 1979: 39</bibRefCitation>
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, pl. 7, text-fig. 8; 
<treatmentCitation id="C626D43A3C30FFA1FC20FC2D303DFC09" author="Holthuis LB" box="[896,1063,947,966]" page="594" pageId="2" pageNumber="3" year="1986">
<bibRefCitation id="23168FDA3C30FFA1FC20FC2D303DFC09" author="Holthuis LB" box="[896,1063,947,966]" pageId="2" pageNumber="3" pagination="589 - 615" refId="ref17636" refString="Holthuis LB. 1986. Decapoda. In: Botosaneanu L (ed.) Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna Inhabiting Subterranean Waters (Including the Marine Interstitial). E. J. Brill, Leiden, pp. 589 - 615." type="book chapter" year="1986">Holthuis 1986: 594</bibRefCitation>
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; 
<treatmentCitation id="C626D43A3C30FFA1FB8FFC2D30A7FC09" author="Guinot D." box="[1071,1213,947,966]" page="13" pageId="2" pageNumber="3" year="1988">
<bibRefCitation id="23168FDA3C30FFA1FB8FFC2D30A7FC09" author="Guinot D." box="[1071,1213,947,966]" pageId="2" pageNumber="3" pagination="3 - 40" refId="ref17537" refString="Guinot D. 1988. Les crabes cavernicoles du monde. Mem Biospeol 15: 3 - 40." type="journal article" year="1988">Guinot 1988: 13</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="C626D43A3C30FFA1FB66FC2D3144FC09" author="Guinot D." box="[1222,1374,947,966]" page="171" pageId="2" pageNumber="3" year="1994">
<bibRefCitation id="23168FDA3C30FFA1FB66FC2D3144FC09" author="Guinot D." box="[1222,1374,947,966]" pageId="2" pageNumber="3" pagination="165 - 179" refId="ref17556" refString="Guinot D. 1994. Decapoda Brachyura. In: Juberthie C, Decou V (eds) Encyclopaedia Biospeologica. Tome 1. Societe de Biospeologie, CNRS, Moulis, Ariege, &amp; Academie Roumaine, Bucarest, pp. 165 - 179, pl. 3." type="journal article" year="1994">Guinot 1994: 171</bibRefCitation>
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; 
<treatmentCitation id="C626D43A3C30FFA0FAC8FC2D355BFEDF" author="Fransen CHJM &amp; Holthuis LB &amp; Adema JPHM" lastPageId="3" lastPageNumber="4" page="136" pageId="2" pageNumber="3" year="1997">
<bibRefCitation id="23168FDA3C30FFA0FAC8FC2D355BFEDF" author="Fransen CHJM &amp; Holthuis LB &amp; Adema JPHM" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" pagination="1 - 344" refId="ref17326" refString="Fransen CHJM, Holthuis LB, Adema JPHM. 1997. Type-catalogue of the Decapod Crustacea in the collections of the Nationaal Naturhistorisch Museum, with appendices of pre- 1900 collectors and material. Zool Verh 311: i - xvi, 1 - 344." type="journal article" year="1997">Fransen et al. 1997: 136</bibRefCitation>
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; Ng 2004: 329; 
<treatmentCitation id="C626D43A3C31FFA0FE72FF63368AFEDF" author="Ng PKL &amp; Yeo DCJ" box="[466,656,253,272]" page="12" pageId="3" pageNumber="4" year="2007">
<bibRefCitation id="23168FDA3C31FFA0FE72FF63368AFEDF" author="Ng PKL &amp; Yeo DCJ" box="[466,656,253,272]" pageId="3" pageNumber="4" pagination="95 - 120" refId="ref19171" refString="Ng PKL, Yeo DCJ. 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua L (ed.) Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia. 28 - 30 June 2005, Forest Research Institute Malaysia, Kepong, pp. 95 - 120." type="book chapter" year="2007">Ng and Yeo 2007: 12</bibRefCitation>
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; 
<treatmentCitation id="C626D43A3C31FFA0FD3BFF63356BFEE5" author="Grinang J &amp; Nyanti L." page="2" pageId="3" pageNumber="4" year="2007">
<bibRefCitation id="23168FDA3C31FFA0FD3BFF63356BFEE5" author="Grinang J &amp; Nyanti L." pageId="3" pageNumber="4" pagination="2" refId="ref17492" refString="Grinang J, Nyanti L. 2007 b. Distribution range of montane crab, Sundathelphusa tenebrosa Holthuis (Decapoda: Brachyura: Parathelphusidae) in Borneo. Research Bulletin, Faculty of Resource Science &amp; Technology, Universiti Malaysia Sarawak 4: 2." type="journal article" year="2007">Grinang and Nyanti 2007b: 2</bibRefCitation>
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(part); 
<treatmentCitation id="C626D43A3C31FFA0FE18FE863678FEE5" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[440,610,279,299]" page="73" pageId="3" pageNumber="4" year="2008">
<bibRefCitation id="23168FDA3C31FFA0FE18FE863678FEE5" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[440,610,279,299]" pageId="3" pageNumber="4" pagination="1 - 286" refId="ref18968" refString="Ng PKL, Guinot D, Davie PJF. 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull Zool Suppl 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008: 73</bibRefCitation>
</treatmentCitation>
; 
<bibRefCitation id="23168FDA3C31FFA0FDCEFE893504FE8A" author="Cumberlidge N &amp; Ng PKL &amp; Yeo DCJ &amp; Magalhaes C &amp; Campos MR &amp; Alvarez F &amp; Naruse T &amp; Daniels SR &amp; Esser LJ &amp; Attipoe FYK &amp; Clotilde-Ba F-L &amp; Darwall W &amp; Mcivor A &amp; Collen B &amp; Ram M." pageId="3" pageNumber="4" pagination="1665 - 1673" refId="ref17167" refString="Cumberlidge N, Ng PKL, Yeo DCJ, Magalhaes C, Campos MR, Alvarez F, Naruse T, Daniels SR, Esser LJ, Attipoe FYK, Clotilde-Ba F-L, Darwall W, Mcivor A, Collen B, Ram M. 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biol Conserv 142: 1665 - 1673. doi: 10.1016 / j. biocon. 2009.02.038." type="journal article" year="2009">Cumberlidge et al. 2009</bibRefCitation>
: unpaginated appendix; 
<treatmentCitation id="C626D43A3C31FFA0FDA6FEAC36C7FE8A" author="Klaus S &amp; Brandis D &amp; Ng PKL &amp; Yeo DCJ &amp; Schubart CD" box="[518,733,306,325]" page="514" pageId="3" pageNumber="4" year="2009">
<bibRefCitation id="23168FDA3C31FFA0FDA6FEAC36C7FE8A" author="Klaus S &amp; Brandis D &amp; Ng PKL &amp; Yeo DCJ &amp; Schubart CD" box="[518,733,306,325]" pageId="3" pageNumber="4" pagination="509 - 531" refId="ref18088" refString="Klaus S, Brandis D, Ng PKL, Yeo DCJ, Schubart CD. 2009. Phylogeny and biogeography of Asian freshwater crabs of the family Gecarcinucidae (Brachyura: Potamoidea). In: Martin JW, Crandall KA, Felder DL (eds) Crustacean Issues 18: Decapod Crustacean Phylogenetics, CRC Press, England, pp. 509 - 531. doi: 10.1201 / 9781420092592 - c 26." type="book chapter" year="2009">Klaus et al. 2009: 514</bibRefCitation>
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; 
<treatmentCitation id="C626D43A3C31FFA0FF15FED33585FEAF" author="McFarlane D &amp; Lundberg J &amp; Christenson K." box="[181,415,333,352]" page="12" pageId="3" pageNumber="4" year="2011">
<bibRefCitation id="23168FDA3C31FFA0FF15FED33585FEAF" author="McFarlane D &amp; Lundberg J &amp; Christenson K." box="[181,415,333,352]" pageId="3" pageNumber="4" pagination="11 - 18" refId="ref18233" refString="McFarlane D, Lundberg J, Christenson K. 2011. New records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of northern Sarawak. Speleobiology Notes 3: 11 - 18." type="journal article" year="2011">McFarlane et al. 2011: 12</bibRefCitation>
</treatmentCitation>
, figs. 2 (unnumbered), 4; 
<treatmentCitation id="C626D43A3C31FFA0FD2DFED334CDFEB5" author="Ng PKL" page="494" pageId="3" pageNumber="4" year="2015">
<bibRefCitation id="23168FDA3C31FFA0FD2DFED334CDFEB5" author="Ng PKL" pageId="3" pageNumber="4" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng 2015: 494</bibRefCitation>
</treatmentCitation>
, figs. 3, 7F–H.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="13F8A2A33C30FFA1FF0EF8F436BDF879" ID-DOI="http://doi.org/10.5281/zenodo.8074522" ID-Zenodo-Dep="8074522" httpUri="https://zenodo.org/record/8074522/files/figure.png" pageId="2" pageNumber="3" startId="2.[174,205,1898,1917]" targetBox="[191,1434,1032,1873]" targetPageId="2" targetType="figure">
<paragraph id="4738F22B3C30FFA1FF0EF8F436BDF879" blockId="2.[174,1451,1898,1974]" pageId="2" pageNumber="3">
<emphasis id="75F32E393C30FFA1FF0EF8F434FCF8B2" bold="true" box="[174,230,1898,1917]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
A, B, frontal margin and median triangle; C, D, male anterior thoracic sternum and sternopleonal cavity.A, C, 
<taxonomicName id="808789A83C30FFA1FB3BF8F5352EF855" authority="(von Martens, 1868)" baseAuthorityName="von Martens" baseAuthorityYear="1868" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="75F32E393C30FFA1FB3BF8F53165F8B1" box="[1179,1407,1899,1918]" italics="true" pageId="2" pageNumber="3">Sundathelphusa philippina</emphasis>
(von Martens, 1868)
</taxonomicName>
, paralectotype male (51.4 × 42.8 mm) (ZMB 1050), Samar, Philippines; B, D, 
<taxonomicName id="808789A83C30FFA1FC4FF819314CF855" authority="(Ng &amp; Stuebing), 1989" baseAuthorityName="Ng &amp; Stuebing" baseAuthorityYear="1989" box="[1007,1366,1927,1946]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="aspera">
<emphasis id="75F32E393C30FFA1FC4FF8193098F856" box="[1007,1154,1927,1945]" italics="true" pageId="2" pageNumber="3">Borneosa aspera</emphasis>
(Ng &amp; Stuebing), 1989
</taxonomicName>
, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah.
</paragraph>
</caption>
<footnote id="249CEE253C30FFA1FB3CF87B31B1F838" box="[1180,1451,2020,2039]" pageId="2" pageNumber="3">
<paragraph id="4738F22B3C30FFA1FB3CF87B31B1F838" blockId="2.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="2" pageNumber="3">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<subSubSection id="0F9DA1A03C31FFA5FF65FE3D3072FDD9" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="4738F22B3C31FFA0FF65FE3D3663FB14" blockId="3.[136,738,419,1979]" pageId="3" pageNumber="4">
<emphasis id="75F32E393C31FFA0FF65FE3D35B4FE74" box="[197,430,419,443]" italics="true" pageId="3" pageNumber="4">Material examined</emphasis>
: 
<materialsCitation id="F7EFF8763C31FFA0FE64FE3D3669FD94" ID-GBIF-Occurrence="4134795307" collectingDate="1978-03-18" collectionCode="NHM" collectorName="J. Dring" country="Malaysia" elevation="2" latitude="4.05" location="Gunong Mulu National Park" longLatPrecision="1307" longitude="114.88333" pageId="3" pageNumber="4" specimenCode="NHM 1979.135" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="holotype">
<typeStatus id="983C4C893C31FFA0FE64FE3D362EFE74" box="[452,564,419,443]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
<specimenCount id="518139A23C31FFA0FDE1FE3D3660FE74" box="[577,634,419,443]" pageId="3" pageNumber="4" type="male">male</specimenCount>
(23.5 × 
<quantity id="807F5FCE3C31FFA0FF28FE5D34F0FE14" box="[136,234,451,475]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="3" pageNumber="4" unit="mm" value="19.1">19.1 mm</quantity>
) (
<specimenCode id="17215A503C31FFA0FEA2FE5D35A8FE14" box="[258,434,451,475]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="4" type="Herbarium">NHM 1979.135</specimenCode>
), Camp 2, in 2 × 2 m pool, 
<quantity id="807F5FCE3C31FFA0FF28FE7D34D0FE35" box="[136,202,483,507]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="cm" value="10.0">10 cm</quantity>
deep, clear cold mountain stream with pebble and gravel bottom, hill dipterocarp forest, 
<geoCoordinate id="22B394EC3C31FFA0FD86FD9D366AFDD4" box="[550,624,515,539]" degrees="4" direction="north" minutes="03" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="4.05">4°03'N</geoCoordinate>
<geoCoordinate id="22B394EC3C31FFA0FDD9FD9D36C7FDD4" box="[633,733,515,539]" degrees="114" direction="east" minutes="53" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="114.88333">114°53'E</geoCoordinate>
, 
<elevation id="CCAA15183C31FFA0FF28FDBD351AFDF4" box="[136,256,547,571]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="m" value="500.0">
<quantity id="807F5FCE3C31FFA0FF28FDBD34C9FDF5" box="[136,211,547,571]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="m" value="500.0">500 m</quantity>
asl
</elevation>
, 
<location id="4258A4F03C31FFA0FEB0FDBD367DFDF4" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FEB0FDBD367DFDF4" box="[272,615,547,571]" country="Malaysia" latitude="4.05" longLatPrecision="1307" longitude="114.88333" name="Gunong Mulu National Park" pageId="3" pageNumber="4" stateProvince="Sarawak">Gunong Mulu National Park</location>
, 
<collectingRegion id="85433CC93C31FFA0FDD6FDBD36C1FDF4" box="[630,731,547,571]" pageId="3" pageNumber="4">Sarawak</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C31FFA0FF28FDDD34EFFD94" box="[136,245,579,603]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
, coll. 
<collectorName id="EA7297FD3C31FFA0FEE2FDDD35BCFD94" box="[322,422,579,603]" pageId="3" pageNumber="4">J. Dring</collectorName>
, 
<date id="3339D4EB3C31FFA0FE15FDDD3674FD94" box="[437,622,579,603]" pageId="3" pageNumber="4" value="1978-03-18">
<collectingDate id="237D2D033C31FFA0FE15FDDD3674FD94" box="[437,622,579,603]" pageId="3" pageNumber="4" value="1978-03-18">18 March 1978</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="F7EFF8763C31FFA0FDDDFDDD3515FD54" ID-GBIF-Occurrence="4134795304" collectingDate="1978-03-18" collectionCode="NHM" collectorName="J. Dring" country="Malaysia" elevation="2" latitude="4.05" location="Gunong Mulu National Park" longLatPrecision="1307" longitude="114.88333" pageId="3" pageNumber="4" specimenCode="NHM 1979.136" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="paratype">
<typeStatus id="983C4C893C31FFA0FDDDFDDD36C5FD94" box="[637,735,579,603]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
<specimenCount id="518139A23C31FFA0FF28FDFD34A7FDB4" box="[136,189,611,635]" pageId="3" pageNumber="4" type="male">male</specimenCount>
(21.9 × 
<quantity id="807F5FCE3C31FFA0FE84FDFD3590FDB4" box="[292,394,611,635]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="3" pageNumber="4" unit="mm" value="17.8">17.8 mm</quantity>
) (
<specimenCode id="17215A503C31FFA0FE03FDFD3640FDB4" box="[419,602,611,635]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="4" type="Herbarium">NHM 1979.136</specimenCode>
), same data as holotype;
</materialsCitation>
<materialsCitation id="F7EFF8763C31FFA0FEB9FD1D35A2FD34" ID-GBIF-Occurrence="4134795311" collectingDate="1978-03-10" collectionCode="NNM" collectorName="Gunong Mulu National &amp; P. Chapman" country="Malaysia" location="Deer Cave" pageId="3" pageNumber="4" specimenCode="D31972" specimenCount="1" specimenCount-female="1" stateProvince="Sarawak" typeStatus="paratype">
<specimenCount id="518139A23C31FFA0FEB9FD1D353CFD54" box="[281,294,643,667]" pageId="3" pageNumber="4" type="female">1</specimenCount>
<typeStatus id="983C4C893C31FFA0FE8FFD1D3597FD54" box="[303,397,643,667]" pageId="3" pageNumber="4">paratype</typeStatus>
<specimenType id="172979273C31FFA0FE37FD1D35FBFD54" box="[407,481,643,667]" pageId="3" pageNumber="4">female</specimenType>
(
<collectionCode id="21966AEE3C31FFA0FE51FD1D3628FD54" box="[497,562,643,667]" collectionName="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">NNM</collectionCode>
<specimenCode id="17215A503C31FFA0FD9BFD1D368DFD54" box="[571,663,643,667]" collectionCode="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">D31972</specimenCode>
), 
<location id="4258A4F03C31FFA0FD0CFD1D34DEFD74" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FD0CFD1D34DEFD74" country="Malaysia" county="Gunong Mulu National Park" latitude="4.05" longLatPrecision="1307" longitude="114.88333" municipality="Gunong Mulu National Park" name="Deer Cave" pageId="3" pageNumber="4" stateProvince="Sarawak">Deer Cave</location>
, 
<collectorName id="EA7297FD3C31FFA0FF72FD3D35C0FD74" box="[210,474,675,699]" pageId="3" pageNumber="4">Gunong Mulu National</collectorName>
stream near southwest entrance, under rock, 
<collectingRegion id="85433CC93C31FFA0FE36FD5D35E4FD14" box="[406,510,707,731]" country="Malaysia" name="Sarawak" pageId="3" pageNumber="4">Sarawak</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C31FFA0FDAEFD5D3667FD14" box="[526,637,707,731]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
, coll. 
<collectorName id="EA7297FD3C31FFA0FD6DFD5D34EFFD34" pageId="3" pageNumber="4">P. Chapman</collectorName>
, 
<date id="3339D4EB3C31FFA0FEA3FD7D35A9FD34" box="[259,435,739,763]" pageId="3" pageNumber="4" value="1978-03-10">
<collectingDate id="237D2D033C31FFA0FEA3FD7D35A9FD34" box="[259,435,739,763]" pageId="3" pageNumber="4" value="1978-03-10">10 March 1978</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="F7EFF8763C31FFA0FE62FD7D367FFC94" ID-GBIF-Occurrence="4134795314" collectingDate="1978-04-25" collectionCode="NNM" collectorName="P. Chapman" country="Malaysia" elevation="100" location="Deer Water Cave" pageId="3" pageNumber="4" specimenCode="D31979" specimenCount="1" stateProvince="Sarawak" typeStatus="paratype">
1 
<typeStatus id="983C4C893C31FFA0FE7AFD7D3620FD34" box="[474,570,739,763]" pageId="3" pageNumber="4">paratype</typeStatus>
<specimenType id="172979273C31FFA0FDE3FD7D3694FD34" box="[579,654,739,763]" pageId="3" pageNumber="4">female</specimenType>
(
<collectionCode id="21966AEE3C31FFA0FD3FFD7D36FAFD34" box="[671,736,739,763]" collectionName="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">NNM</collectionCode>
<specimenCode id="17215A503C31FFA0FF28FC9D34FDFCD4" box="[136,231,771,795]" collectionCode="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">D31979</specimenCode>
), 
<location id="4258A4F03C31FFA0FF5DFC9D35DCFCD4" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FF5DFC9D35DCFCD4" box="[253,454,771,795]" country="Malaysia" county="Gunong Mulu National Park" latitude="4.05" longLatPrecision="1307" longitude="114.88333" municipality="Gunong Mulu National Park" name="Deer Water Cave" pageId="3" pageNumber="4" stateProvince="Sarawak">Deer Water Cave</location>
, 
<location id="4258A4F03C31FFA0FE75FC9D34A0FCF4" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FE75FC9D34A0FCF4" country="Malaysia" county="Gunong Mulu National Park" latitude="4.05" longLatPrecision="1307" longitude="114.88333" municipality="Gunong Mulu National Park" name="Gunong Mulu National Park" pageId="3" pageNumber="4" stateProvince="Sarawak">Gunong Mulu National Park</location>
in stream about 
<quantity id="807F5FCE3C31FFA0FEDDFCBD35DFFCF5" box="[381,453,803,827]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="m" value="100.0">
<elevation id="CCAA15183C31FFA0FEDDFCBD35DFFCF5" box="[381,453,803,827]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="m" value="100.0">100 m</elevation>
</quantity>
from entrance, 
<collectingRegion id="85433CC93C31FFA0FDDBFCBD36C6FCF4" box="[635,732,803,827]" country="Malaysia" name="Sarawak" pageId="3" pageNumber="4">Sarawak</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C31FFA0FF28FCDD34F7FC94" box="[136,237,835,859]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
, coll. 
<collectorName id="EA7297FD3C31FFA0FE92FCDD35A2FC94" box="[306,440,835,859]" pageId="3" pageNumber="4">P. Chapman</collectorName>
, 
<date id="3339D4EB3C31FFA0FE64FCDD367BFC94" box="[452,609,835,859]" pageId="3" pageNumber="4" value="1978-04-25">
<collectingDate id="237D2D033C31FFA0FE64FCDD367BFC94" box="[452,609,835,859]" pageId="3" pageNumber="4" value="1978-04-25">25 April 1978</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="F7EFF8763C31FFA0FDCEFCDD36D3FC74" ID-GBIF-Occurrence="4134795313" collectingDate="1978-04-13" collectionCode="NNM" collectorName="P. Chapman" location="Gunong Mulu National Park" municipality="Clearwater Cave" pageId="3" pageNumber="4" specimenCode="D31972" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="518139A23C31FFA0FDCEFCDD3661FC94" box="[622,635,835,859]" pageId="3" pageNumber="4" type="female">2</specimenCount>
<typeStatus id="983C4C893C31FFA0FD25FCDD36FBFC94" box="[645,737,835,859]" pageId="3" pageNumber="4">paratype</typeStatus>
<specimenType id="172979273C31FFA0FF28FCFD34FAFCB4" box="[136,224,867,891]" pageId="3" pageNumber="4">females</specimenType>
(
<collectionCode id="21966AEE3C31FFA0FF52FCFD352EFCB4" box="[242,308,867,891]" collectionName="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">NNM</collectionCode>
<specimenCode id="17215A503C31FFA0FE9FFCFD3584FCB4" box="[319,414,867,891]" collectionCode="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="4" type="Museum">D31972</specimenCode>
), 
<collectingMunicipality id="A75C68513C31FFA0FE14FCFD366DFCB4" box="[436,631,867,891]" country="Malaysia" county="Gunong Mulu National Park" latitude="4.05" longLatPrecision="1307" longitude="114.88333" municipality="Gunong Mulu National Park" name="Clearwater Cave" pageId="3" pageNumber="4">Clearwater Cave</collectingMunicipality>
, 
<location id="4258A4F03C31FFA0FD27FCFD3563FC54" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FD27FCFD3563FC54" country="Malaysia" county="Gunong Mulu National Park" latitude="4.05" longLatPrecision="1307" longitude="114.88333" municipality="Clearwater Cave" name="Gunong Mulu National Park" pageId="3" pageNumber="4">Gunong Mulu National Park</location>
in main stream passage near river junction, coll. 
<collectorName id="EA7297FD3C31FFA0FEDBFC3D3611FC74" box="[379,523,931,955]" pageId="3" pageNumber="4">P. Chapman</collectorName>
, 
<date id="3339D4EB3C31FFA0FDBAFC3D36DEFC74" box="[538,708,931,955]" pageId="3" pageNumber="4" value="1978-04-13">
<collectingDate id="237D2D033C31FFA0FDBAFC3D36DEFC74" box="[538,708,931,955]" pageId="3" pageNumber="4" value="1978-04-13">13 April 1978</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="F7EFF8763C31FFA0FD74FC3D34DDFBF4" ID-GBIF-Occurrence="4134795309" collectingDate="1978-04-16" collectionCode="ZRC" collectorName="R. P. Lim &amp; G. Liew" country="Malaysia" location="Sungei Melinau" pageId="3" pageNumber="4" specimenCode="ZRC 1989.3420" specimenCount="1" specimenCount-female="1" stateProvince="Sarawak">
<specimenCount id="518139A23C31FFA0FD74FC3D34CEFC14" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
(30.8 × 
<quantity id="807F5FCE3C31FFA0FE9DFC5D35BEFC14" box="[317,420,963,987]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="3" pageNumber="4" unit="mm" value="24.6">24.6 mm</quantity>
) (
<specimenCode id="17215A503C31FFA0FE1EFC5D3666FC14" box="[446,636,963,987]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="4">ZRC 1989.3420</specimenCode>
), 
<location id="4258A4F03C31FFA0FD33FC5D34FEFC34" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FD33FC5D34FEFC34" country="Malaysia" name="Sungei Melinau" pageId="3" pageNumber="4" stateProvince="Sarawak">Sungei Melinau</location>
, 
<location id="4258A4F03C31FFA0FF4FFC7D3587FC34" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FF4FFC7D3587FC34" box="[239,413,995,1019]" country="Malaysia" name="Merimau Gorge" pageId="3" pageNumber="4" stateProvince="Sarawak">Merimau Gorge</location>
, 
<location id="4258A4F03C31FFA0FE08FC7D36C7FC34" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FE08FC7D36C7FC34" box="[424,733,995,1019]" country="Malaysia" name="Gunong Mulu National Park" pageId="3" pageNumber="4" stateProvince="Sarawak">Gunong Mulu National Park</location>
, 
<collectingRegion id="85433CC93C31FFA0FF28FB9D34FDFBD4" box="[136,231,1027,1051]" country="Malaysia" name="Sarawak" pageId="3" pageNumber="4">Sarawak</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C31FFA0FF53FB9D354DFBD4" box="[243,343,1027,1051]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
, coll. 
<collectorName id="EA7297FD3C31FFA0FE3BFB9D35E0FBD4" box="[411,506,1027,1051]" pageId="3" pageNumber="4">R.P. Lim</collectorName>
&amp; 
<collectorName id="EA7297FD3C31FFA0FD80FB9D3661FBD4" box="[544,635,1027,1051]" pageId="3" pageNumber="4">G. Liew</collectorName>
, 
<date id="3339D4EB3C31FFA0FD27FB9D34D9FBF4" pageId="3" pageNumber="4" value="1978-04-16">
<collectingDate id="237D2D033C31FFA0FD27FB9D34D9FBF4" pageId="3" pageNumber="4" value="1978-04-16">16 April 1978</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="F7EFF8763C31FFA0FF71FBBD3650FBB4" ID-GBIF-Occurrence="4134795310" collectingDate="1978-04-17" collectionCode="ZRC" collectorName="R. P. Lim" country="Malaysia" location="Sungei Melinau" pageId="3" pageNumber="4" specimenCode="ZRC 1989.3682" specimenCount="1" specimenCount-juvenile="1" stateProvince="Sarawak">
<specimenCount id="518139A23C31FFA0FF71FBBD3559FBF4" box="[209,323,1059,1083]" pageId="3" pageNumber="4" type="juvenile">1 juvenile</specimenCount>
(
<specimenCode id="17215A503C31FFA0FEF4FBBD360AFBF4" box="[340,528,1059,1083]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="4">ZRC 1989.3682</specimenCode>
), 
<location id="4258A4F03C31FFA0FD86FBBD36C6FBF4" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FD86FBBD36C6FBF4" box="[550,732,1059,1083]" country="Malaysia" name="Sungei Melinau" pageId="3" pageNumber="4" stateProvince="Sarawak">Sungei Melinau</location>
, 
<location id="4258A4F03C31FFA0FF28FBDD352CFB94" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FF28FBDD352CFB94" box="[136,310,1091,1115]" country="Malaysia" name="Merimau Gorge" pageId="3" pageNumber="4" stateProvince="Sarawak">Merimau Gorge</location>
, 
<location id="4258A4F03C31FFA0FEE0FBDD366EFB94" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FEE0FBDD366EFB94" box="[320,628,1091,1115]" country="Malaysia" name="Gunong Mulu National Park" pageId="3" pageNumber="4" stateProvince="Sarawak">Gunong Mulu National Park</location>
, 
<collectingRegion id="85433CC93C31FFA0FDDFFBDD36C4FB94" box="[639,734,1091,1115]" country="Malaysia" name="Sarawak" pageId="3" pageNumber="4">Sarawak</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C31FFA0FF28FBFD34F4FBB4" box="[136,238,1123,1147]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
, coll. 
<collectorName id="EA7297FD3C31FFA0FE95FBFD3580FBB4" box="[309,410,1123,1147]" pageId="3" pageNumber="4">R.P. Lim</collectorName>
, 
<date id="3339D4EB3C31FFA0FE07FBFD365CFBB4" box="[423,582,1123,1147]" pageId="3" pageNumber="4" value="1978-04-17">
<collectingDate id="237D2D033C31FFA0FE07FBFD365CFBB4" box="[423,582,1123,1147]" pageId="3" pageNumber="4" value="1978-04-17">17 April 1978</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="F7EFF8763C31FFA0FDF4FBFD3663FB14" ID-GBIF-Occurrence="4134795302" collectingDate="2006-11-04" collectionCode="ZRC" collectorName="A. D. Tranh" country="Malaysia" location="Mulu Summit trail at Camp 2" pageId="3" pageNumber="4" specimenCode="ZRC 2013.1293" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak">
<specimenCount id="518139A23C31FFA0FDF4FBFD3685FBB4" box="[596,671,1123,1147]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
(16.1 × 13.0 mm) (
<specimenCode id="17215A503C31FFA0FE87FB1D35F1FB54" box="[295,491,1155,1179]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="4">ZRC 2013.1293</specimenCode>
), 
<location id="4258A4F03C31FFA0FDA3FB1D351FFB74" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FDA3FB1D351FFB74" country="Malaysia" name="Mulu Summit trail at Camp 2" pageId="3" pageNumber="4" stateProvince="Sarawak">Mulu Summit trail at Camp 2</location>
, 
<location id="4258A4F03C31FFA0FEB5FB3D3673FB74" LSID="urn:lsid:plazi:treatment:CF2E433D3C30FFA5FC37FCA33072FDD9:4258A4F03C31FFA0FEB5FB3D3673FB74" box="[277,617,1187,1211]" country="Malaysia" name="Gunong Mulu National Park" pageId="3" pageNumber="4" stateProvince="Sarawak">Gunong Mulu National Park</location>
, 
<collectingRegion id="85433CC93C31FFA0FDD7FB3D36C1FB74" box="[631,731,1187,1211]" country="Malaysia" name="Sarawak" pageId="3" pageNumber="4">Sarawak</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C31FFA0FF28FB5D34F6FB14" box="[136,236,1219,1243]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
, coll. 
<collectorName id="EA7297FD3C31FFA0FE8BFB5D35BDFB14" box="[299,423,1219,1243]" pageId="3" pageNumber="4">A.D. Tranh</collectorName>
, 
<date id="3339D4EB3C31FFA0FE11FB5D366FFB14" box="[433,629,1219,1243]" pageId="3" pageNumber="4" value="2006-11-04">
<collectingDate id="237D2D033C31FFA0FE11FB5D366FFB14" box="[433,629,1219,1243]" pageId="3" pageNumber="4" value="2006-11-04">4 November 2006</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="4738F22B3C31FFA0FF65FB7A308BFE99" blockId="3.[136,738,419,1979]" lastBlockId="3.[813,1414,253,1973]" pageId="3" pageNumber="4">
<emphasis id="75F32E393C31FFA0FF65FB7A3550FB34" box="[197,330,1251,1275]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
: Adult carapace transversely subquadrate, width to length ratio 1.23–1.25 (
<figureCitation id="DFBCEEAE3C31FFA0FD07FA9D34D7FAF5" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2A, B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FF7BFABD34E7FAF4" box="[219,253,1315,1339]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); striae on lateral parts of carapace distinct (
<figureCitation id="DFBCEEAE3C31FFA0FF31FADD34E3FA94" box="[145,249,1347,1371]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FEA9FADD3531FA94" box="[265,299,1347,1371]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); branchial regions gently convex in frontal view (
<figureCitation id="DFBCEEAE3C31FFA0FEBCFAFD3568FAB4" box="[284,370,1379,1403]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Fig. 2C</figureCitation>
); dorsal margin of frontal median triangle adjacent to but not contiguous with lateral margins, separated by a gap or connected by small flattened granules (
<figureCitation id="DFBCEEAE3C31FFA0FEC9FA5D35FEFA14" box="[361,484,1475,1499]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Fig. 2C–F</figureCitation>
); exorbital tooth with outer margin almost entire (
<figureCitation id="DFBCEEAE3C31FFA0FE19FA7D3603FA34" box="[441,537,1507,1531]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FD86FA7D365DFA34" box="[550,583,1507,1531]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); epibranchial tooth very low, just visible, separated from rest of margin by shallow low cleft (
<figureCitation id="DFBCEEAE3C31FFA0FE40F9BD365FF9F4" box="[480,581,1571,1595]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FDF5F9BD366DF9F4" box="[597,631,1571,1595]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); median lobe of posterior margin of epistome acutely triangular (
<figureCitation id="DFBCEEAE3C31FFA0FF30F9FD3512F9B4" box="[144,264,1635,1659]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Fig. 2C–F</figureCitation>
); ischium of third maxilliped subquadrate (
<figureCitation id="DFBCEEAE3C31FFA0FF31F91D34EEF954" box="[145,244,1667,1691]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
); fingers of adult male cheliped closing along entire cutting margins (
<figureCitation id="DFBCEEAE3C31FFA0FE44F93D3659F974" box="[484,579,1699,1723]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">Fig. 3D</figureCitation>
); P3 dactylus longer than P4 dactylus (
<figureCitation id="DFBCEEAE3C31FFA0FE10F95D362EF914" box="[432,564,1731,1755]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">Fig. 3G, H</figureCitation>
); male pleonal somite 6 rectangular, longer than broad (
<figureCitation id="DFBCEEAE3C31FFA0FDD4F97D36C8F934" box="[628,722,1763,1787]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">Fig. 3C</figureCitation>
); male sternopleonal cavity not prominently anterior, distance between tip of cavity and suture of thoracic sternites 2 and 3 subequal to length of thoracic sternite 2 (
<figureCitation id="DFBCEEAE3C31FFA0FF08F8FD3518F8B4" box="[168,258,1891,1915]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">Fig. 3B</figureCitation>
); adult Gl subterminal segment relatively slender; terminal segment subcylindrical, distal half gently curved outwards, tapering gradually to truncate tip, 0.25–0.28 times length of subterminal segment, distal opening large, ovate (
<figureCitation id="DFBCEEAE3C31FFA0FBC6FE83313DFEF9" box="[1126,1319,285,310]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng &amp; Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="3" pageNumber="4">Fig. 20F, G, I–K</figureCitation>
); vulvae transversely subovate (
<figureCitation id="DFBCEEAE3C31FFA0FB82FEA3309CFE99" box="[1058,1158,317,342]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="3" pageNumber="4">Fig. 23A</figureCitation>
).
</paragraph>
<paragraph id="4738F22B3C31FFA7FCC9FEC036BAFEFA" blockId="3.[813,1414,253,1973]" lastBlockId="4.[174,775,253,438]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="75F32E393C31FFA0FCC9FEC037F1FEBA" box="[873,1003,350,373]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
: Carapace transversely subquadrate, width to length ratio 1.23–1.25 (
<figureCitation id="DFBCEEAE3C31FFA0FB15FEE3315EFE5A" box="[1205,1348,381,406]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2A, B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FAF4FEE0316DFE59" box="[1364,1399,382,406]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); regions separated by distinct grooves; dorsal surface gently convex, glabrous, pitted, rugose, especially along margins and cristae, striae on lateral parts of carapace most distinct (
<figureCitation id="DFBCEEAE3C31FFA0FC70FE63302EFDD9" box="[976,1076,509,534]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FBE3FE60307FFDD9" box="[1091,1125,510,534]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); frontal region with low granules and rugae especially nearer epigastric region, orbital regions finely pitted to smooth; cervical groove deep, broad, extending from lateral edge of postorbital cristae to deep H-shaped gastric groove; branchial regions gently convex, not distinctly inflated in frontal view (
<figureCitation id="DFBCEEAE3C31FFA0FCD2FD23300AFD1A" box="[882,1040,701,726]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B, C, F</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FBBEFD203025FD19" box="[1054,1087,702,726]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); postorbital cristae relatively sharp, distinct, gradually becoming lower, less distinct towards cervical groove, with separate short, low rugose crista just before anterolateral margin; postorbital crista separated from epigastric crista by shallow oblique groove; epigastric cristae anterior of postorbital cristae, distinct, not sharp, prominently rugose, margins sloping anteriorly, separated medially by deep longitudinal groove which extends to just before mesogastric area (
<figureCitation id="DFBCEEAE3C31FFA0FC95FC4337A5FC3A" box="[821,959,989,1014]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B, F</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FC6FFC4037E9FC39" box="[975,1011,990,1014]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
). Frontal margin slightly sinuous to almost straight, slightly deflexed; frontal median triangle transversely narrow, median in position, edges of dorsal margin adjacent to but not contiguous with lateral margins, separated by a gap or connected by small flattened granules (
<figureCitation id="DFBCEEAE3C31FFA0FBEEFBE330DCFB59" box="[1102,1222,1149,1174]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Fig. 2C–F</figureCitation>
). Exorbital tooth low, broadly triangular, outer margin length ca. 4 times longer than inner margin, outer margin almost straight, lined with small granules, almost entire, curving to meet anterolateral margin, forming very small triangular notch (
<figureCitation id="DFBCEEAE3C31FFA0FCDAFA8337C1FAF9" box="[890,987,1309,1334]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FC48FA803010FAF9" box="[1000,1034,1310,1334]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
). epibranchial tooth very low, just visible, separated from rest of margin by shallow low cleft (
<figureCitation id="DFBCEEAE3C31FFA0FCCBFAC337D3FAB9" box="[875,969,1373,1398]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FC75FAC037ECFAB9" box="[981,1014,1374,1398]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
); anterolateral margin gently convex, lined with small granules, adjacent surfaces covered with numerous rows of strong oblique striae; not clearly demarcated from posterolateral margin; posterolateral margin gently concave, adjacent surfaces lined with strong oblique striae, converging to almost straight or gently convex posterior carapace margin (
<figureCitation id="DFBCEEAE3C31FFA0FABAF983319AF9F9" box="[1306,1408,1565,1590]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C31FFA0FC8DF9A03754F999" box="[813,846,1598,1622]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="3" pageNumber="4">3E</figureCitation>
). Pterygostomial, suborbital, sub-hepatic and subbranchial regions distinctly rugose or lined with oblique granules (
<figureCitation id="DFBCEEAE3C31FFA0FC36F9E3301DF959" box="[918,1031,1661,1686]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Fig. 2C–F</figureCitation>
). Posterior margin of epistome with median lobe acutely triangular, tip sharp; lateral margin with low, broadly triangular lobe, separated by deep fissures (
<figureCitation id="DFBCEEAE3C31FFA0FC2BF94337E6F939" box="[907,1020,1757,1782]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="3" pageNumber="4">Fig. 2C–F</figureCitation>
). Third maxilliped quadrate; ischium subquadrate, longer than broad, sulcus deep, oblique, closer to inner margin; merus squarish, lateral and distal margins cristate, medially depressed; exopod slender, reaching to about half length of merus, with distinct triangular projection on inner margin of distal part, flagellum long, extending across width of merus (
<figureCitation id="DFBCEEAE3C31FFA7FAF9F80334C9FED9" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
). Terminal segment of mandibular palp bilobed, inner lobe twice length of outer lobe, flap-like.
</paragraph>
<footnote id="249CEE253C31FFA0FF28F87B3582F838" box="[136,408,2020,2039]" pageId="3" pageNumber="4">
<paragraph id="4738F22B3C31FFA0FF28F87B3582F838" blockId="3.[136,408,2020,2039]" box="[136,408,2020,2039]" pageId="3" pageNumber="4">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4738F22B3C36FFA6FF4AFEA336C3FEF9" blockId="4.[174,775,253,438]" lastBlockId="5.[136,737,253,437]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Chelipeds subequal with one chela larger but not inflated; outer surfaces of articles rugose (
<figureCitation id="DFBCEEAE3C36FFA7FDCDFEC336D1FEB9" box="[621,715,349,374]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="4" pageNumber="5">Figs. 2A</figureCitation>
, 
<figureCitation id="DFBCEEAE3C36FFA7FD76FEC036E0FEB9" box="[726,762,350,374]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="4" pageNumber="5">3D</figureCitation>
). Basis-ischium fused, with low tooth on anterior margin (
<figureCitation id="DFBCEEAE3C36FFA7FF16FE033509FE79" box="[182,275,413,438]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="4" pageNumber="5">Fig. 3B</figureCitation>
). Merus trigonal in cross-section, margins uneven, covered with striae and low granules; anterior margin with low subterminal tooth. Upper surfaces of carpus with broad, shallow longitudinal depression, inner angle with prominent sharp spine, base with low tubercle (
<figureCitation id="DFBCEEAE3C36FFA7FC1BFEE3300FFE59" box="[955,1045,381,406]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
). Fingers of chela longer than palm; cutting edges with numerous sharp teeth and denticles, tips of fingers strongly curved; fingers of adult male cheliped closing along entire cutting margins (
<figureCitation id="DFBCEEAE3C37FFA6FDD7FE8336D7FEF9" box="[631,717,285,310]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="5" pageNumber="6">Fig. 3D</figureCitation>
).
</paragraph>
<caption id="13F8A2A33C36FFA7FF0EF8F7365FF87B" ID-DOI="http://doi.org/10.5281/zenodo.8074524" ID-Zenodo-Dep="8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="4" pageNumber="5" startId="4.[174,205,1897,1916]" targetBox="[174,1451,484,1872]" targetPageId="4" targetType="figure">
<paragraph id="4738F22B3C36FFA7FF0EF8F7365FF87B" blockId="4.[174,1452,1897,1972]" pageId="4" pageNumber="5">
<emphasis id="75F32E393C36FFA7FF0EF8F734FDF8B3" bold="true" box="[174,231,1897,1916]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="808789A83C36FFA7FF53F8F4362FF8B3" authority="(Holthuis, 1979)" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[243,565,1897,1916]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="75F32E393C36FFA7FF53F8F43584F8B3" box="[243,414,1897,1916]" italics="true" pageId="4" pageNumber="5">Borneosa tenebrosa</emphasis>
(
<bibRefCitation id="23168FDA3C36FFA7FE0BF8F73634F8B3" author="Holthuis LB" box="[427,558,1897,1916]" pageId="4" pageNumber="5" pagination="3 - 47" refId="ref17606" refString="Holthuis LB. 1979. Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zool Verh 171: 3 - 47, pls. 1 - 8." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
)
</taxonomicName>
, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax.
</paragraph>
</caption>
<footnote id="249CEE253C36FFA7FB3CF87B31B1F838" box="[1180,1451,2020,2039]" pageId="4" pageNumber="5">
<paragraph id="4738F22B3C36FFA7FB3CF87B31B1F838" blockId="4.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="4" pageNumber="5">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4738F22B3C37FFA6FF65FEA33110FE59" blockId="5.[136,737,253,437]" lastBlockId="5.[813,1414,253,437]" pageId="5" pageNumber="6">
Ambulatory legs not elongated, P3 longest; outer surfaces gently rugose (
<figureCitation id="DFBCEEAE3C37FFA6FE7AFEC33625FEB9" box="[474,575,349,374]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="5" pageNumber="6">Figs. 2A</figureCitation>
, 
<figureCitation id="DFBCEEAE3C37FFA6FDEDFEC0369CFEB9" box="[589,646,350,374]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="5" pageNumber="6">3F–I</figureCitation>
). Merus stout, anterior margin slightly crested, uneven, appears weakly serrated, dorsal subterminal angle just visible but without spine or tooth; propodus with 2 rows of downward pointing spines on posterior margin; dactylus almost straight or gently curved, margins with numerous well developed spines pointing obliquely distally; P3 dactylus longer than P4 dactylus (
<figureCitation id="DFBCEEAE3C37FFA6FB36FEE330E4FE59" box="[1174,1278,381,406]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="5" pageNumber="6">Fig. 3F–I</figureCitation>
).
</paragraph>
<paragraph id="4738F22B3C37FFA5FCC9FE03364CFD99" blockId="5.[813,1414,253,437]" lastBlockId="6.[174,776,253,1974]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Surface of thoracic sternum punctate; sternites 1 and 2 completely fused, forming triangular plate; separated from sternite 3 by gently concave suture; sternites 3 and 4 proportionately wide, completely fused except for lateral clefts and shallow median groove; sternopleonal cavity deep, almost reaching imaginary line connecting anterior edges of coxae of chelipeds, cavity not prominently anterior, distance between tip of cavity and suture of thoracic sternites 2 and 3 subequal to length of thoracic sternite 2 (
<figureCitation id="DFBCEEAE3C34FFA5FD96FE63368CFDD9" box="[566,662,509,534]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="6" pageNumber="7">Fig. 3B</figureCitation>
); pleonal locking mechanism composed of rounded tubercle on submedian part of sternite 5 (
<figureCitation id="DFBCEEAE3C34FFA5FE46FDA33653FD99" box="[486,585,573,598]" captionStart="Fig" captionStartId="25.[136,168,1870,1889]" captionTargetBox="[136,1413,729,1841]" captionTargetId="figure-349@25.[136,1413,729,1841]" captionTargetPageId="25" captionText="Fig. 19. Male sternopleonal cavity and pleonal locking tubercle on sternite 5. A, Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; C, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; D, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074568" httpUri="https://zenodo.org/record/8074568/files/figure.png" pageId="6" pageNumber="7">Fig. 19A</figureCitation>
).
</paragraph>
<caption id="13F8A2A33C37FFA6FF28F8D336F7F87B" ID-DOI="http://doi.org/10.5281/zenodo.8074526" ID-Zenodo-Dep="8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="5" pageNumber="6" startId="5.[136,167,1869,1888]" targetBox="[136,1413,500,1842]" targetPageId="5" targetType="figure">
<paragraph id="4738F22B3C37FFA6FF28F8D336F7F87B" blockId="5.[136,1414,1869,1973]" pageId="5" pageNumber="6">
<emphasis id="75F32E393C37FFA6FF28F8D334D8F8AF" bold="true" box="[136,194,1869,1888]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="808789A83C37FFA6FF70F8D03609F8AF" authority="(Holthuis, 1979)" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[208,531,1869,1888]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="75F32E393C37FFA6FF70F8D03561F8AF" box="[208,379,1869,1888]" italics="true" pageId="5" pageNumber="6">Borneosa tenebrosa</emphasis>
(
<bibRefCitation id="23168FDA3C37FFA6FE28F8D33617F8AF" author="Holthuis LB" box="[392,525,1869,1888]" pageId="5" pageNumber="6" pagination="3 - 47" refId="ref17606" refString="Holthuis LB. 1979. Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zool Verh 171: 3 - 47, pls. 1 - 8." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
)
</taxonomicName>
. A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale.
</paragraph>
</caption>
<footnote id="249CEE253C37FFA6FF28F87B3582F838" box="[136,408,2020,2039]" pageId="5" pageNumber="6">
<paragraph id="4738F22B3C37FFA6FF28F87B3582F838" blockId="5.[136,408,2020,2039]" box="[136,408,2020,2039]" pageId="5" pageNumber="6">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4738F22B3C34FFA5FF4AFDC334E6FC59" blockId="6.[174,776,253,1974]" pageId="6" pageNumber="7">
Pleon distinctly T-shaped; all somites and telson free; somite 1 longitudinally narrow, extending across width of posterior carapace margin, reaching bases of coxae of P5; somite 2 as wide as somite 1; somite 3 subrectangular, with convex lateral margins; somites 4 and 5 trapezoidal with gently concave to straight lateral margins; male pleonal somite 6 rectangular, longer than broad, distal and proximal margins subequal; telson triangular with concave lateral margins, rounded tip (
<figureCitation id="DFBCEEAE3C34FFA5FD7DFCC334F5FC59" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="6" pageNumber="7">Fig. 3B, C</figureCitation>
).
</paragraph>
<paragraph id="4738F22B3C34FFA5FF4AFC0335FAFA99" blockId="6.[174,776,253,1974]" pageId="6" pageNumber="7">
Adult Gl directed obliquely outwards; subterminal segment relatively slender, tapering gradually from broad base, sinuous on proximal part of outer margin, rest of margin gently concave; terminal segment clearly demarcated from subterminal segment but grooves shallow; terminal segment subcylindrical, dorsal and ventral surfaces with numerous long setae, distal half gently curved outwards, tapering gradually to truncate tip, 0.25–0.28 times length of subterminal segment, groove for G2 marginal or slightly dorsal in position, dorsal and ventral folds slightly overlapping, distal opening large, ovate (
<figureCitation id="DFBCEEAE3C34FFA5FE39FB63364BFAD9" box="[409,593,1277,1302]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng &amp; Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="6" pageNumber="7">Fig. 20F, G, I–K</figureCitation>
). G2 longer than Gl, flagellum well developed, about one-third length of basal segment (
<figureCitation id="DFBCEEAE3C34FFA5FEF3FAA335CEFA99" box="[339,468,1341,1366]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng &amp; Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="6" pageNumber="7">Fig. 20H, L</figureCitation>
).
</paragraph>
<paragraph id="4738F22B3C34FFA5FF4AFAC33595F999" blockId="6.[174,776,253,1974]" pageId="6" pageNumber="7">
<emphasis id="75F32E393C34FFA5FF4AFAC33525FABA" box="[234,319,1373,1397]" italics="true" pageId="6" pageNumber="7">Female</emphasis>
: Adult female chelipeds are relatively slenderer compared to those of males. The female pleon is ovate, has all the somites and telson free, and covers most of the thoracic sternal surface. The vulvae are large, transversely subovate, without a clear sternal cover with the posterior margin slightly rimmed, and are positioned on the submedian part or anterior third of sternite 6 (
<figureCitation id="DFBCEEAE3C34FFA5FEBFF9A33599F999" box="[287,387,1597,1622]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="6" pageNumber="7">Fig. 23A</figureCitation>
).
</paragraph>
<paragraph id="4738F22B3C34FFA5FF4AF9C33605F8D9" blockId="6.[174,776,253,1974]" pageId="6" pageNumber="7">
<emphasis id="75F32E393C34FFA5FF4AF9C3352CF9BA" box="[234,310,1629,1653]" italics="true" pageId="6" pageNumber="7">Colour</emphasis>
: 
<bibRefCitation id="23168FDA3C34FFA5FEE7F9C3363AF9B9" author="Holthuis LB" box="[327,544,1629,1654]" pageId="6" pageNumber="7" pagination="3 - 47" refId="ref17606" refString="Holthuis LB. 1979. Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zool Verh 171: 3 - 47, pls. 1 - 8." type="journal article" year="1979">Holthuis (1979: 42)</bibRefCitation>
described the colour as follows: “brown with orange-brown edges to the carapace… Dorsally dark brown with a slight purplish tinge. Venter pale brown, mouthparts dark and purple tinged.” This is similar to colour photographs of the species in 
<bibRefCitation id="23168FDA3C34FFA5FEBDF9633601F8D9" author="McFarlane D &amp; Lundberg J &amp; Christenson K." box="[285,539,1789,1814]" pageId="6" pageNumber="7" pagination="11 - 18" refId="ref18233" refString="McFarlane D, Lundberg J, Christenson K. 2011. New records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of northern Sarawak. Speleobiology Notes 3: 11 - 18." type="journal article" year="2011">McFarlane et al. (2011)</bibRefCitation>
.
</paragraph>
<paragraph id="4738F22B3C34FFA5FF4AF8833581F8BA" blockId="6.[174,776,253,1974]" pageId="6" pageNumber="7">
<emphasis id="75F32E393C34FFA5FF4AF8833552F8FA" box="[234,328,1821,1845]" italics="true" pageId="6" pageNumber="7">Remarks</emphasis>
: The taxonomy of this species has been discussed at length by 
<bibRefCitation id="23168FDA3C34FFA5FE00F8A03615F89A" author="Ng PKL" box="[416,527,1853,1878]" pageId="6" pageNumber="7" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng (2015)</bibRefCitation>
and there is no need to elaborate further here.
</paragraph>
<paragraph id="4738F22B3C34FFA5FF4AF8E03072FDD9" blockId="6.[174,776,253,1974]" lastBlockId="6.[850,1451,253,534]" pageId="6" pageNumber="7">
<emphasis id="75F32E393C34FFA5FF4AF8E03556F85A" box="[234,332,1917,1941]" italics="true" pageId="6" pageNumber="7">Biology</emphasis>
: The species was described from specimens collected in Deer Cave, Deer Water Cave and Clearwater Cave in Gunong Mulu National Park, the highest altitude being 
<quantity id="807F5FCE3C34FFA5FBC8FE8330B6FEFA" box="[1128,1196,285,309]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="m" value="500.0">500 m</quantity>
but they are clearly not troglobites (
<bibRefCitation id="23168FDA3C34FFA5FC78FEA33063FE99" author="Holthuis LB" box="[984,1145,317,342]" pageId="6" pageNumber="7" pagination="3 - 47" refId="ref17606" refString="Holthuis LB. 1979. Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zool Verh 171: 3 - 47, pls. 1 - 8." type="journal article" year="1979">Holthuis 1979</bibRefCitation>
) (see also 
<bibRefCitation id="23168FDA3C34FFA5FB55FEA33791FEB9" author="McFarlane D &amp; Lundberg J &amp; Christenson K." pageId="6" pageNumber="7" pagination="11 - 18" refId="ref18233" refString="McFarlane D, Lundberg J, Christenson K. 2011. New records of crabs from the caves of Gunung Mulu National Park, Sarawak, with a field key to the cavernicolous taxa of northern Sarawak. Speleobiology Notes 3: 11 - 18." type="journal article" year="2011">McFarlane et al. 2011</bibRefCitation>
). It has also been found in streams outside caves (
<collectionCode id="21966AEE3C34FFA5FCFAFEE03788FE59" box="[858,914,382,406]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC</collectionCode>
2013.1293). Although most of the specimens have been found in caves, they should be considered as facultative cavernicoles at best as they have none of the adaptations associated with troglobites (cf. 
<bibRefCitation id="23168FDA3C34FFA5FAEFFE433791FDDA" author="Holthuis LB" pageId="6" pageNumber="7" pagination="589 - 615" refId="ref17636" refString="Holthuis LB. 1986. Decapoda. In: Botosaneanu L (ed.) Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna Inhabiting Subterranean Waters (Including the Marine Interstitial). E. J. Brill, Leiden, pp. 589 - 615." type="book chapter" year="1986">Holthuis 1986</bibRefCitation>
; 
<bibRefCitation id="23168FDA3C34FFA5FC36FE633006FDD9" author="Guinot D." box="[918,1052,509,534]" pageId="6" pageNumber="7" pagination="3 - 40" refId="ref17537" refString="Guinot D. 1988. Les crabes cavernicoles du monde. Mem Biospeol 15: 3 - 40." type="journal article" year="1988">Guinot 1988</bibRefCitation>
<bibRefCitation id="23168FDA3C34FFA5FB83FE603047FDD9" author="Guinot D." box="[1059,1117,510,534]" pageId="6" pageNumber="7" pagination="165 - 179" refId="ref17556" refString="Guinot D. 1994. Decapoda Brachyura. In: Juberthie C, Decou V (eds) Encyclopaedia Biospeologica. Tome 1. Societe de Biospeologie, CNRS, Moulis, Ariege, &amp; Academie Roumaine, Bucarest, pp. 165 - 179, pl. 3." type="journal article" year="1994">1994</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>