284 lines
54 KiB
XML
284 lines
54 KiB
XML
<document id="6BE22D790C1A3A8D6873CCF0CAD7AADE" ID-CLB-Dataset="54577" ID-DOI="10.6620/ZS.2022.61-13" ID-GBIF-Dataset="854b8390-f85f-4b0a-8c7a-2bfc1da39225" ID-PMC="PMC9579960" ID-PubMed="36330021" ID-Zenodo-Dep="8074520" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687184145743" checkinUser="valdenar" docAuthor="Ng, Peter K L & Grinang, Jongkar" docDate="2022" docId="CF2E433D3C24FFB4FE98FB0334F8FEB9" docLanguage="en" docName="ZoolStud.61.13.1-35.pdf" docOrigin="Zoological studies 61 (13)" docSource="http://dx.doi.org/10.6620/ZS.2022.61-13" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Borneosa serrata Ng & Grinang 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="24" masterDocId="33173B453C32FFA3FFA0FF9E341AFFCF" masterDocTitle="Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species." masterLastPageNumber="35" masterPageNumber="1" pageNumber="23" updateTime="1687519204558" updateUser="ExternalLinkService">
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<mods:title id="7DB697E13347FE6CC6BB3BFDD44953BF">Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species.</mods:title>
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<mods:namePart id="A0E355C54E6C327E1F96EBA5F94D5C49">Ng, Peter K L</mods:namePart>
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<mods:namePart id="813D64EDF16FBCBE07D6D1C8824B2A87">Grinang, Jongkar</mods:namePart>
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<mods:title id="FFB4B9AAB70AB30CB2E36B8756C1F31E">Zoological studies</mods:title>
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<mods:part id="B115EF6C9843ADF3304AB4F47D9C5F95">
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<mods:date id="4681C3D10BA8A793B192C2AA32BC7923">2022</mods:date>
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<mods:number id="D08294E66BE91226D049167B19B230A4">2022-05-05</mods:number>
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<mods:number id="3C1E3849535A3B76CB980B66D967D26B">61</mods:number>
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<mods:number id="6443ECD05B0E4F62B3E519FF9A2256F8">13</mods:number>
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<treatment id="CF2E433D3C24FFB4FE98FB0334F8FEB9" ID-DOI="http://doi.org/10.5281/zenodo.8055978" ID-Zenodo-Dep="8055978" LSID="urn:lsid:plazi:treatment:CF2E433D3C24FFB4FE98FB0334F8FEB9" httpUri="http://treatment.plazi.org/id/CF2E433D3C24FFB4FE98FB0334F8FEB9" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection id="0F9DA1A03C24FFB5FE98FB033667FB79" box="[312,637,1181,1206]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="4738F22B3C24FFB5FE98FB033667FB79" blockId="22.[204,745,1181,1290]" box="[312,637,1181,1206]" pageId="22" pageNumber="23">
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<heading id="1C7045473C24FFB5FE98FB033667FB79" bold="true" box="[312,637,1181,1206]" centered="true" fontSize="10" level="2" pageId="22" pageNumber="23" reason="2">
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<emphasis id="75F32E393C24FFB5FE98FB033667FB79" bold="true" box="[312,637,1181,1206]" pageId="22" pageNumber="23">
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<taxonomicName id="808789A83C24FFB5FE98FB033609FB79" ID-CoL="992XS" authority="Ng & Grinang, 2022" authorityName="Ng & Grinang" authorityYear="2022" box="[312,531,1181,1206]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="serrata" status="sp. nov.">
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<emphasis id="75F32E393C24FFB5FE98FB033609FB79" bold="true" box="[312,531,1181,1206]" italics="true" pageId="22" pageNumber="23">Borneosa serrata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6EC093423C24FFB5FDBAFB033667FB79" box="[538,637,1181,1206]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0F9DA1A03C24FFB5FEC5FB233630FAC5" pageId="22" pageNumber="23" type="description">
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<paragraph id="4738F22B3C24FFB5FEC5FB233630FAC5" blockId="22.[204,745,1181,1290]" pageId="22" pageNumber="23">
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(
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<figureCitation id="DFBCEEAE3C24FFB5FECDFB2335DCFB1A" box="[365,454,1213,1237]" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Figs. 17</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C24FFB5FE71FB2335F4FB1A" box="[465,494,1213,1237]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">18</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C24FFB5FE59FB233653FB1A" box="[505,585,1213,1237]" captionStart="Fig" captionStartId="28.[174,205,1869,1888]" captionTargetBox="[239,1395,251,1840]" captionTargetId="figure-5@28.[227,1398,240,1871]" captionTargetPageId="28" captionText="Fig. 22. A–D, Borneosa kapit sp. nov., paratype male (23.5 × 19.1 mm) (ZRC 2021.0518), Belaga, Sarawak; E–G, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak; H, B. kapit sp. nov., paratype male (35.8 × 28.0 mm) (ZRC 2021.0521), Kapit, Sarawak; I– L, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak; M–P, S. serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. Scale bars: A, D, E, H, I, L, M, P = 1.0 mm; B, C, F, G, J, K, N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074574" httpUri="https://zenodo.org/record/8074574/files/figure.png" pageId="22" pageNumber="23">22M–P</figureCitation>
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)
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<uri id="3316FE293C24FFB5FF6CFB463630FAC5" pageId="22" pageNumber="23">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="3321C8FE3C24FFB5FE19FB463630FAC5" pageId="22" pageNumber="23">B65416DA-4501-43C1-A0FB-31C9D46816A0</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="0F9DA1A03C24FFB5FF4AFAAE36C6FA67" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph id="4738F22B3C24FFB5FF4AFAAE36C6FA67" blockId="22.[174,775,1328,1960]" pageId="22" pageNumber="23">
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<emphasis id="75F32E393C24FFB5FF4AFAAE35ABFA87" box="[234,433,1328,1352]" italics="true" pageId="22" pageNumber="23">Material examined</emphasis>
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||
:
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<materialsCitation id="F7EFF8763C24FFB5FE1FFAAF36C6FA67" collectingDate="1995-08-20" collectorName="R. Diesel" country="Indonesia" latitude="2.7" location="Kalimantan Utara" longLatPrecision="7852" longitude="116.7" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Utara" typeStatus="holotype">
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||
<specimenCount id="518139A23C24FFB5FE1FFAAF361FFA87" box="[447,517,1328,1353]" pageId="22" pageNumber="23" type="male">1 male</specimenCount>
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||
(25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean (= Pengean), Long Yiu Hulu, Kabupaten Bulungan,
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<collectingRegion id="85433CC93C24FFB5FE16FAEE3693FA47" box="[438,649,1392,1416]" country="Indonesia" name="Kalimantan Utara" pageId="22" pageNumber="23">Kalimantan Utara</collectingRegion>
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, ca.
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||
<geoCoordinate id="22B394EC3C24FFB5FD63FAEF371FFA46" box="[707,773,1393,1417]" degrees="2.7" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="5555" value="2.7">2.7°N</geoCoordinate>
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||
<geoCoordinate id="22B394EC3C24FFB5FF0EFA0E351CFA67" box="[174,262,1424,1448]" degrees="116.7" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="5555" value="116.7">116.7°E</geoCoordinate>
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||
,
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||
<collectingCountry id="3F90B2BB3C24FFB5FEB1FA0E3561FA67" box="[273,379,1424,1448]" name="Indonesia" pageId="22" pageNumber="23">Indonesia</collectingCountry>
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||
, coll.
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||
<collectorName id="EA7297FD3C24FFB5FE1CFA0F3638FA67" box="[444,546,1424,1448]" pageId="22" pageNumber="23">R. Diesel</collectorName>
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||
,
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||
<date id="3339D4EB3C24FFB5FD8CFA0F36C3FA67" box="[556,729,1424,1449]" pageId="22" pageNumber="23" value="1995-08-20">
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||
<collectingDate id="237D2D033C24FFB5FD8CFA0F36C3FA67" box="[556,729,1424,1449]" pageId="22" pageNumber="23" value="1995-08-20">20 August 1995</collectingDate>
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||
</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0F9DA1A03C24FFB5FF4AFA2F302DFDF9" pageId="22" pageNumber="23" type="diagnosis">
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||
<paragraph id="4738F22B3C24FFB5FF4AFA2F31B2FE1A" blockId="22.[174,775,1328,1960]" lastBlockId="22.[850,1452,253,1974]" pageId="22" pageNumber="23">
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<emphasis id="75F32E393C24FFB5FF4AFA2F3575FA07" box="[234,367,1457,1480]" italics="true" pageId="22" pageNumber="23">Diagnosis</emphasis>
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: Adult carapace transversely subquadrate, width to length ratio 1.16 (
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<figureCitation id="DFBCEEAE3C24FFB5FD35FA4E34D9F9C6" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17A, B</figureCitation>
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); striae on lateral parts of carapace distinct (
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<figureCitation id="DFBCEEAE3C24FFB5FD78FA6E34F8F9E6" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17B</figureCitation>
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); branchial regions gently convex in frontal view (
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<figureCitation id="DFBCEEAE3C24FFB5FF56F9AE359AF986" box="[246,384,1584,1609]" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17C, F</figureCitation>
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); dorsal margin of frontal median triangle contiguous with lateral margins (
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<figureCitation id="DFBCEEAE3C24FFB5FDD7F9CE36E0F9A6" box="[631,762,1616,1641]" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17C–E</figureCitation>
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); exorbital tooth with outer margin serrated (
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<figureCitation id="DFBCEEAE3C24FFB5FD33F9EE36E3F946" box="[659,761,1648,1673]" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17B</figureCitation>
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); epibranchial tooth prominent, separated from rest of margin by distinct cleft (
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<figureCitation id="DFBCEEAE3C24FFB5FE73F92E3624F906" box="[467,574,1712,1737]" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17B</figureCitation>
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); median lobe of posterior margin of epistome acutely triangular (
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<figureCitation id="DFBCEEAE3C24FFB5FD7AF94E351AF8C6" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17C–E</figureCitation>
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); ischium of third maxilliped subrectangular (
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<figureCitation id="DFBCEEAE3C24FFB5FF16F88E3500F8E6" box="[182,282,1808,1833]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18A</figureCitation>
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); fingers of adult male cheliped closing along entire cutting margins (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FE1BF8AE3639F886" box="[443,547,1840,1865]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18E</figureCitation>
|
||
); P3 and P4 dactyli subequal in length (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FE29F8CE3609F8A6" box="[393,531,1872,1897]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18G, H</figureCitation>
|
||
); male pleonal somite 6 subquadrate, slightly longer than broad (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FD32F8EE36E0F846" box="[658,762,1904,1929]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18C</figureCitation>
|
||
); male sternopleonal cavity prominently anterior, distance between tip of cavity and suture of thoracic sternites 2 and 3 distinctly shorter than length of thoracic sternite 2 (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FCD1FEA337C2FE99" box="[881,984,317,342]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18B</figureCitation>
|
||
); adult Gl subterminal segment relatively slender; terminal segment subconical, distal half gently curved outwards, tapering gradually to subtruncate tip, 0.31 times length of subterminal segment, distal opening large, ovate (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FB99FE2330D8FE19" box="[1081,1218,445,470]" captionStart="Fig" captionStartId="28.[174,205,1869,1888]" captionTargetBox="[239,1395,251,1840]" captionTargetId="figure-5@28.[227,1398,240,1871]" captionTargetPageId="28" captionText="Fig. 22. A–D, Borneosa kapit sp. nov., paratype male (23.5 × 19.1 mm) (ZRC 2021.0518), Belaga, Sarawak; E–G, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak; H, B. kapit sp. nov., paratype male (35.8 × 28.0 mm) (ZRC 2021.0521), Kapit, Sarawak; I– L, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak; M–P, S. serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. Scale bars: A, D, E, H, I, L, M, P = 1.0 mm; B, C, F, G, J, K, N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074574" httpUri="https://zenodo.org/record/8074574/files/figure.png" pageId="22" pageNumber="23">Fig. 22M–O</figureCitation>
|
||
); females not known.
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C24FFB5FC2FFE43302DFDF9" blockId="22.[850,1452,253,1974]" pageId="22" pageNumber="23">
|
||
<emphasis id="75F32E393C24FFB5FC2FFE4337C5FE3A" box="[911,991,477,501]" italics="true" pageId="22" pageNumber="23">Colour</emphasis>
|
||
: When freshly collected, the specimen was a uniform dark brown on all the dorsal surfaces (R. Diesel, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F9DA1A03C24FFB5FC2FFDA3308AFD5A" pageId="22" pageNumber="23" type="etymology">
|
||
<paragraph id="4738F22B3C24FFB5FC2FFDA3308AFD5A" blockId="22.[850,1452,253,1974]" pageId="22" pageNumber="23">
|
||
<emphasis id="75F32E393C24FFB5FC2FFDA3301BFD9A" box="[911,1025,573,597]" italics="true" pageId="22" pageNumber="23">Etymology</emphasis>
|
||
: The species is named for the serrated appearance of the anterolateral margin, with the epibranchial tooth prominent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F9DA1A03C24FFB5FC2FFD033163F879" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="4738F22B3C24FFB5FC2FFD033004FAD9" blockId="22.[850,1452,253,1974]" pageId="22" pageNumber="23">
|
||
<emphasis id="75F32E393C24FFB5FC2FFD0337EBFD7A" box="[911,1009,669,693]" italics="true" pageId="22" pageNumber="23">Remarks</emphasis>
|
||
: Morphologically,
|
||
<taxonomicName id="808789A83C24FFB5FB73FD00315DFD7A" authority="Ng & Grinang, 2022" authorityName="Ng & Grinang" authorityYear="2022" box="[1235,1351,670,693]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="serrata" status="sp. nov.">
|
||
<emphasis id="75F32E393C24FFB5FB73FD00315DFD7A" box="[1235,1351,670,693]" italics="true" pageId="22" pageNumber="23">B. serrata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EC093423C24FFB5FAF1FD0031B1FD7A" box="[1361,1451,670,693]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
most closely resembles
|
||
<taxonomicName id="808789A83C24FFB5FBFEFD2030B5FD1A" authority="Ng & Grinang, 2022" authorityName="Ng & Grinang" authorityYear="2022" box="[1118,1199,701,725]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="niah" status="sp. nov.">
|
||
<emphasis id="75F32E393C24FFB5FBFEFD2030B5FD1A" box="[1118,1199,701,725]" italics="true" pageId="22" pageNumber="23">B. niah</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EC093423C24FFB5FB19FD203108FD1A" box="[1209,1298,702,725]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, especially in the presence of a more well-developed epibranchial tooth separated from the anterolateral margin by a deep cleft; and the male sternopleonal cavity projects deeper into thoracic sternite 3 with only a narrow space present between the tip of the cavity and the suture between sternites 1 and 3.
|
||
<taxonomicName id="808789A83C24FFB5FBBFFCE030F9FC5A" authorityName="Ng & Grinang" authorityYear="2022" box="[1055,1251,894,917]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="serrata">
|
||
<emphasis id="75F32E393C24FFB5FBBFFCE030F9FC5A" box="[1055,1251,894,917]" italics="true" pageId="22" pageNumber="23">Borneosa serrata</emphasis>
|
||
</taxonomicName>
|
||
, however, can be distinguished in having the outer margin of the exorbital tooth serrated (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC5CFC23307BFC19" box="[1020,1121,957,982]" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17A</figureCitation>
|
||
) (versus almost smooth in
|
||
<taxonomicName id="808789A83C24FFB5FA34FC20379BFC3A" authorityName="Ng & Grinang" authorityYear="2022" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="niah">
|
||
<emphasis id="75F32E393C24FFB5FA34FC20379BFC3A" italics="true" pageId="22" pageNumber="23">B. niah</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC30FC4337F9FC39" box="[912,995,989,1014]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="22" pageNumber="23">Fig. 8B</figureCitation>
|
||
); the median tooth on the posterior margin of the epistome is sharply triangular in shape (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FAE0FC63377EFBF9" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17C– E</figureCitation>
|
||
) (versus blunter in
|
||
<taxonomicName id="808789A83C24FFB5FBEFFB8030B9FBFA" authorityName="Ng & Grinang" authorityYear="2022" box="[1103,1187,1053,1077]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="niah">
|
||
<emphasis id="75F32E393C24FFB5FBEFFB8030B9FBFA" box="[1103,1187,1053,1077]" italics="true" pageId="22" pageNumber="23">B. niah</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FB16FB83312BFBF9" box="[1206,1329,1053,1078]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="22" pageNumber="23">Fig. 8C–E</figureCitation>
|
||
); the third maxilliped ischium is proportionately longer (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FAE2FBA331B9FB99" box="[1346,1443,1085,1110]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18A</figureCitation>
|
||
) (versus proportionately shorter in
|
||
<taxonomicName id="808789A83C24FFB5FB7DFBC0312AFBBA" authorityName="Ng & Grinang" authorityYear="2022" box="[1245,1328,1117,1141]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="niah">
|
||
<emphasis id="75F32E393C24FFB5FB7DFBC0312AFBBA" box="[1245,1328,1117,1141]" italics="true" pageId="22" pageNumber="23">B. niah</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FAE2FBC33184FBB9" box="[1346,1438,1117,1142]" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
); the male pleon (notably somites 3–5) is relatively more narrow (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC11FB033009FB79" box="[945,1043,1181,1206]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18C</figureCitation>
|
||
) (versus relatively wider in
|
||
<taxonomicName id="808789A83C24FFB5FAF3FB0031BEFB7A" authorityName="Ng & Grinang" authorityYear="2022" box="[1363,1444,1181,1205]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="niah">
|
||
<emphasis id="75F32E393C24FFB5FAF3FB0031BEFB7A" box="[1363,1444,1181,1205]" italics="true" pageId="22" pageNumber="23">B. niah</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FCF2FB2337B7FB19" box="[850,941,1213,1238]" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="22" pageNumber="23">Fig. 9C</figureCitation>
|
||
); and the G1 terminal segment is relatively longer (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC0CFB433021FB39" box="[940,1083,1245,1270]" captionStart="Fig" captionStartId="28.[174,205,1869,1888]" captionTargetBox="[239,1395,251,1840]" captionTargetId="figure-5@28.[227,1398,240,1871]" captionTargetPageId="28" captionText="Fig. 22. A–D, Borneosa kapit sp. nov., paratype male (23.5 × 19.1 mm) (ZRC 2021.0518), Belaga, Sarawak; E–G, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak; H, B. kapit sp. nov., paratype male (35.8 × 28.0 mm) (ZRC 2021.0521), Kapit, Sarawak; I– L, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak; M–P, S. serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. Scale bars: A, D, E, H, I, L, M, P = 1.0 mm; B, C, F, G, J, K, N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074574" httpUri="https://zenodo.org/record/8074574/files/figure.png" pageId="22" pageNumber="23">Fig. 22M–O</figureCitation>
|
||
) (versus relatively shorter in
|
||
<taxonomicName id="808789A83C24FFB5FA34FB40379BFADA" authorityName="Ng & Grinang" authorityYear="2022" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="niah">
|
||
<emphasis id="75F32E393C24FFB5FA34FB40379BFADA" italics="true" pageId="22" pageNumber="23">B. niah</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC30FB633009FAD9" box="[912,1043,1277,1302]" captionStart="Fig" captionStartId="27.[136,167,1926,1945]" captionTargetBox="[147,1405,1034,1897]" captionTargetId="figure-6@27.[136,1413,1023,1905]" captionTargetPageId="27" captionText="Fig. 21. A–D, Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak; E–G, B. bario sp. nov., holotype male (29.6 × 23.1 mm) (ZRC 2021.0692), Bario, Sarawak. Scale bars:A, D, E, H = 1.0 mm; B, C, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074572" httpUri="https://zenodo.org/record/8074572/files/figure.png" pageId="22" pageNumber="23">Fig. 21A–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C24FFB5FC2FFA833163F879" blockId="22.[850,1452,253,1974]" pageId="22" pageNumber="23">
|
||
The G1 structure of
|
||
<taxonomicName id="808789A83C24FFB5FB2CFA80311FFAFA" authorityName="Ng & Grinang" authorityYear="2022" box="[1164,1285,1310,1333]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="serrata">
|
||
<emphasis id="75F32E393C24FFB5FB2CFA80311FFAFA" box="[1164,1285,1310,1333]" italics="true" pageId="22" pageNumber="23">B. serrata</emphasis>
|
||
</taxonomicName>
|
||
is very close to that of
|
||
<taxonomicName id="808789A83C24FFB5FC6DFAA03082FA9A" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[973,1176,1341,1365]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brachyphallus">
|
||
<emphasis id="75F32E393C24FFB5FC6DFAA03082FA9A" box="[973,1176,1341,1365]" italics="true" pageId="22" pageNumber="23">B. brachyphallus</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName id="808789A83C24FFB5FB7BFAA03149FA9A" authorityName="Ng & Grinang" authorityYear="2022" box="[1243,1363,1342,1365]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="serrata">
|
||
<emphasis id="75F32E393C24FFB5FB7BFAA03149FA9A" box="[1243,1363,1342,1365]" italics="true" pageId="22" pageNumber="23">B. serrata</emphasis>
|
||
</taxonomicName>
|
||
can be separated in possessing a distinct epibranchial tooth (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FCFAFAE337A0FA59" box="[858,954,1405,1430]" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17B</figureCitation>
|
||
) (versus tooth very low to almost absent in
|
||
<taxonomicName id="808789A83C24FFB5FA34FAE037F7FA7A" baseAuthorityName="Ng" baseAuthorityYear="2015" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brachyphallus">
|
||
<emphasis id="75F32E393C24FFB5FA34FAE037F7FA7A" italics="true" pageId="22" pageNumber="23">B. brachyphallus</emphasis>
|
||
</taxonomicName>
|
||
with the anterolateral margin appearing almost entire;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC5AFA233044FA1A" box="[1018,1118,1469,1493]" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="22" pageNumber="23">Figs. 6B</figureCitation>
|
||
,
|
||
<figureCitation id="DFBCEEAE3C24FFB5FBCDFA20309DFA19" box="[1133,1159,1470,1494]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="22" pageNumber="23">7I</figureCitation>
|
||
); the outer margin of the exorbital tooth is tuberculate and appears serrated (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FA20FA433765F9D9" captionStart="Fig" captionStartId="23.[136,167,1898,1917]" captionTargetBox="[136,1413,434,1872]" captionTargetId="figure-97@23.[136,1413,434,1872]" captionTargetPageId="23" captionText="Fig. 17. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074564" httpUri="https://zenodo.org/record/8074564/files/figure.png" pageId="22" pageNumber="23">Fig. 17B</figureCitation>
|
||
) (versus almost smooth in
|
||
<taxonomicName id="808789A83C24FFB5FB0DFA60317CF9DA" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[1197,1382,1533,1557]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brachyphallus">
|
||
<emphasis id="75F32E393C24FFB5FB0DFA60317CF9DA" box="[1197,1382,1533,1557]" italics="true" pageId="22" pageNumber="23">B. brachyphallus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FAD7FA63376EF9FA" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="22" pageNumber="23">Figs. 6B</figureCitation>
|
||
,
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC21F9803780F9F9" box="[897,922,1566,1590]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="22" pageNumber="23">7I</figureCitation>
|
||
); the third maxilliped ischium is proportionately longer (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FC09F9A33017F999" box="[937,1037,1597,1622]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18A</figureCitation>
|
||
) (versus shorter in
|
||
<taxonomicName id="808789A83C24FFB5FB49F9A031BEF99A" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[1257,1444,1597,1621]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brachyphallus">
|
||
<emphasis id="75F32E393C24FFB5FB49F9A031BEF99A" box="[1257,1444,1597,1621]" italics="true" pageId="22" pageNumber="23">B. brachyphallus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FCF2F9C337B3F9B9" box="[850,937,1629,1654]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="22" pageNumber="23">Fig. 7A</figureCitation>
|
||
); the male sternopleonal cavity projects deeper into thoracic sternite 3, with only a narrow space present between the cavity and the suture between sternites 1 and 3 (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FBB3F9233068F919" box="[1043,1138,1725,1750]" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18B</figureCitation>
|
||
) (versus there is a wider gap between the tip of the male sternopleonal cavity and the suture between sternites 2 and
|
||
<quantity id="807F5FCE3C24FFB5FB10F96030C6F8DA" box="[1200,1244,1789,1814]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="22" pageNumber="23" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="808789A83C24FFB5FB46F96031BEF8DA" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[1254,1444,1789,1813]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brachyphallus">
|
||
<emphasis id="75F32E393C24FFB5FB46F96031BEF8DA" box="[1254,1444,1789,1813]" italics="true" pageId="22" pageNumber="23">B. brachyphallus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FCF2F88337BFF8F9" box="[850,933,1821,1846]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="22" pageNumber="23">Fig. 7B</figureCitation>
|
||
); male pleonal somite 6 is distinctly longer (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FA20F883379AF899" captionStart="Fig" captionStartId="24.[174,205,1898,1917]" captionTargetBox="[174,1451,650,1871]" captionTargetId="figure-268@24.[174,1451,650,1871]" captionTargetPageId="24" captionText="Fig. 18. Borneosa serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074566" httpUri="https://zenodo.org/record/8074566/files/figure.png" pageId="22" pageNumber="23">Fig. 18C</figureCitation>
|
||
) (versus proportionately shorter and more quadrate in
|
||
<taxonomicName id="808789A83C24FFB5FCD0F8C03031F8BA" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[880,1067,1885,1909]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brachyphallus">
|
||
<emphasis id="75F32E393C24FFB5FCD0F8C03031F8BA" box="[880,1067,1885,1909]" italics="true" pageId="22" pageNumber="23">B. brachyphallus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FB9CF8C33088F8B9" box="[1084,1170,1885,1910]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="22" pageNumber="23">Fig. 7C</figureCitation>
|
||
); and the G1 subterminal segment is slightly longer (
|
||
<figureCitation id="DFBCEEAE3C24FFB5FB08F8E3315AF859" box="[1192,1344,1917,1942]" captionStart="Fig" captionStartId="28.[174,205,1869,1888]" captionTargetBox="[239,1395,251,1840]" captionTargetId="figure-5@28.[227,1398,240,1871]" captionTargetPageId="28" captionText="Fig. 22. A–D, Borneosa kapit sp. nov., paratype male (23.5 × 19.1 mm) (ZRC 2021.0518), Belaga, Sarawak; E–G, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak; H, B. kapit sp. nov., paratype male (35.8 × 28.0 mm) (ZRC 2021.0521), Kapit, Sarawak; I– L, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak; M–P, S. serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. Scale bars: A, D, E, H, I, L, M, P = 1.0 mm; B, C, F, G, J, K, N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074574" httpUri="https://zenodo.org/record/8074574/files/figure.png" pageId="22" pageNumber="23">Fig. 22M–O</figureCitation>
|
||
) (versus relatively shorter in
|
||
<taxonomicName id="808789A83C24FFB5FB8AF80030FBF87A" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[1066,1249,1949,1973]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brachyphallus">
|
||
<emphasis id="75F32E393C24FFB5FB8AF80030FBF87A" box="[1066,1249,1949,1973]" italics="true" pageId="22" pageNumber="23">B. brachyphallus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C24FFB5FB4EF8033174F879" box="[1262,1390,1949,1974]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng & Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="22" pageNumber="23">Fig. 20Q–S</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<footnote id="249CEE253C24FFB5FB3CF87B31B1F838" box="[1180,1451,2020,2039]" pageId="22" pageNumber="23">
|
||
<paragraph id="4738F22B3C24FFB5FB3CF87B31B1F838" blockId="22.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="22" pageNumber="23">© 2022 Academia Sinica, Taiwan</paragraph>
|
||
</footnote>
|
||
<subSubSection id="0F9DA1A03C25FFB4FF65FF6334F8FEB9" pageId="23" pageNumber="24" type="biology_ecology">
|
||
<paragraph id="4738F22B3C25FFB4FF65FF6334F8FEB9" blockId="23.[136,737,253,374]" pageId="23" pageNumber="24">
|
||
<emphasis id="75F32E393C25FFB4FF65FF633500FEDA" box="[197,282,253,277]" italics="true" pageId="23" pageNumber="24">Biology</emphasis>
|
||
: The specimen was collected in a small tributary with rock substrates and dense forest cover in the centre of
|
||
<collectingRegion id="85433CC93C25FFB4FEE8FEA3360DFE9A" box="[328,535,317,341]" country="Indonesia" name="Kalimantan Utara" pageId="23" pageNumber="24">Kalimantan Utara</collectingRegion>
|
||
(R. Diesel, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |