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<document id="FBB14B63E30A61AFDD6F2842E87C86D4" ID="10.11646/zootaxa.4140.1.1" ID-DOI="10.11646/zootaxa.4140.1.1" ID-GBIF-Dataset="01995ca4-7d04-4402-b21a-926957b1a391" ID-ISSN="1175-5326" ID-Zenodo-Dep="256507" ID-ZooBank="AD1F6B4E-3B0D-422C-B47C-0495BD385857" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469426945103" checkinUser="plazi" docAuthor="Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. &amp; Blakemore, Robert J." docDate="2016" docId="B5727931FFA17917FF3C2706612E880A" docLanguage="en" docName="zootaxa.4140.1.1.pdf" docOrigin="Zootaxa 4140 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Amynthas bangoianus Michaelsen 1934" docType="treatment" docVersion="22" lastPageNumber="17" masterDocId="494B0149FFB17907FFAB254162078C42" masterDocTitle="A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="17" updateTime="1698365898991" updateUser="plazi">
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<mods:title id="61C138A8BFB5E7B6E894AB1EDC4A7993">A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)</mods:title>
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<mods:namePart id="6CB0630B4A60D8CA1B7EC8FC9E869164">Nguyen, Tung T.</mods:namePart>
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<mods:namePart id="7A4F26D43A3F39DEEDFC93CFD10E3134">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="8A134C3405356C3318FCD274FAF47753">Blakemore, Robert J.</mods:namePart>
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<treatment id="B5727931FFA17917FF3C2706612E880A" ID-DOI="http://doi.org/10.5281/zenodo.5467050" ID-GBIF-Taxon="123207266" ID-Zenodo-Dep="5467050" LSID="urn:lsid:plazi:treatment:B5727931FFA17917FF3C2706612E880A" httpUri="http://treatment.plazi.org/id/B5727931FFA17917FF3C2706612E880A" lastPageNumber="17" pageId="16" pageNumber="17">
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<paragraph id="3D64C827FFA17917FF3C270660878E22" blockId="16.[151,640,583,608]" box="[151,640,583,608]" pageId="16" pageNumber="17">
<heading id="662C7F4BFFA17917FF3C270660878E22" bold="true" box="[151,640,583,608]" fontSize="10" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="FADBB3A4FFA17917FF3C270660878E22" authority="Michaelsen, 1934" authorityName="Michaelsen" authorityYear="1934" box="[151,640,583,608]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="bangoianus">
<emphasis id="0FAF1435FFA17917FF3C270660878E22" bold="true" box="[151,640,583,608]" pageId="16" pageNumber="17">
<emphasis id="0FAF1435FFA17917FF3C2706639D8E22" bold="true" box="[151,410,583,608]" italics="true" pageId="16" pageNumber="17">Amynthas bangoianus</emphasis>
(Michaelsen, 1934)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFA17917FF3C27D1632F8EA1" pageId="16" pageNumber="17" type="reference_group">
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<taxonomicName id="FADBB3A4FFA17917FF3C27D160EA8EE4" authority="Michaelsen, 1934b: 505" authorityName="Michaelsen" authorityPageNumber="505" authorityYear="1934" box="[151,749,656,678]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="bangoiana" subGenus="Pheretima">
<emphasis id="0FAF1435FFA17917FF3C27D163078EE4" box="[151,256,656,678]" italics="true" pageId="16" pageNumber="17">Pheretima</emphasis>
(
<emphasis id="0FAF1435FFA17917FEA527D163718EE4" box="[270,374,656,678]" italics="true" pageId="16" pageNumber="17">Pheretima</emphasis>
)
<emphasis id="0FAF1435FFA17917FE2E27D163F68EE4" box="[389,497,656,678]" italics="true" pageId="16" pageNumber="17">bangoiana</emphasis>
<treatmentCitation id="BC7AEE36FFA17917FE5327D160EA8EE4" author="Michaelsen" box="[504,749,656,678]" page="505" pageId="16" pageNumber="17" year="1934">
<bibRefCitation id="594AB5D6FFA17917FE5327D160BB8EE4" author="Michaelsen" box="[504,700,656,678]" pageId="16" pageNumber="17" refString="Michaelsen, W. (1934 b) Oligochaten von Franzosisch-Indochina. Archive de Zoologie Experimentale et Generale, 76, 493 - 546." type="journal article" year="1934" yearSuffix="b">Michaelsen, 1934b</bibRefCitation>
: 505
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</taxonomicName>
, Figs. VIVIII;
<treatmentCitation id="BC7AEE36FFA17917FC3A27D1665A8EE4" author="Nakamura" box="[913,1117,656,678]" page="40" pageId="16" pageNumber="17" year="1999">
<bibRefCitation id="594AB5D6FFA17917FC3A27D166318EE4" author="Nakamura" box="[913,1078,656,678]" pageId="16" pageNumber="17" refString="Nakamura, Y. (1999) Checklist of earthworms of Pheretima genus group (Megascolecidae: Oligochaeta) of the world. Edaphologia, 64, 1 - 78." type="journal article" year="1999">Nakamura 1999</bibRefCitation>
: 40
</treatmentCitation>
.
<emphasis id="0FAF1435FFA17917FF3C27EF638A8E86" box="[151,397,686,708]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="FADBB3A4FFA17917FF3C27EF62FC8E86" box="[151,251,686,708]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="16" pageNumber="17" phylum="Annelida" rank="genus">Amynthas</taxonomicName>
bangoianus—
</emphasis>
<treatmentCitation id="BC7AEE36FFA17917FE2627EF60978E86" author="Sims" box="[397,656,686,708]" page="236" pageId="16" pageNumber="17" year="1972">
<bibRefCitation id="594AB5D6FFA17917FE2627EF60678E86" author="Sims" box="[397,608,686,708]" pageId="16" pageNumber="17" refString="Sims, R. W. &amp; Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4, 169 - 268. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x" type="journal article" year="1972">Sims &amp; Easton 1972</bibRefCitation>
: 236
</treatmentCitation>
(
<emphasis id="0FAF1435FFA17917FD3527EF60E98E86" box="[670,750,686,708]" italics="true" pageId="16" pageNumber="17">sieboldi</emphasis>
-group);
<treatmentCitation id="BC7AEE36FFA17917FCEC27EF66268E86" author="Blakemore" box="[839,1057,686,708]" page="11" pageId="16" pageNumber="17" year="2007">
<bibRefCitation id="594AB5D6FFA17917FCEC27EF61FA8E86" author="Blakemore" box="[839,1021,686,708]" pageId="16" pageNumber="17" refString="Blakemore, R. J. (2007 a) Updated checklist of Pheretimoids (Oligochaeta: Megascolecidae: Pheretima auct.) taxa. www. annelida. net / earthworm / Pheretimoids. pdf (accessed 6 Jun 2016)" type="journal article" year="2007" yearSuffix="a">Blakemore 2007a</bibRefCitation>
: 11
</treatmentCitation>
;
<bibRefCitation id="594AB5D6FFA17917FB8727EF66E48E86" author="Blakemore" box="[1068,1251,686,708]" pageId="16" pageNumber="17" refString="Blakemore, R. J. (2008 b) List of earthworms from Cambodia, Laos, and Vietnam (Excel spreadsheet). In: A Series of Searchable Texts on Earthworm Biodiversity, Ecology and Systematics from Various Regions of the World - 2 nd Edition (2008). VermEcology, Yokohama. CD - ROM. Avaliable from: http: // www. annelida. net / earthworm / Vietnam % 20 + % 20 Lao s % 20 + % 20 Cambodia 1. xls (Accessed 21 Jul. 2016)" type="journal article" year="2008" yearSuffix="b">Blakemore 2008b</bibRefCitation>
;
<treatmentCitation id="BC7AEE36FFA17917FB4627EF62E58EA1" author="Blakemore" page="21" pageId="16" pageNumber="17" year="2016">
<bibRefCitation id="594AB5D6FFA17917FB4627EF679E8E86" author="Blakemore" box="[1261,1433,686,708]" pageId="16" pageNumber="17" refString="Blakemore, R. J. (2016) Darwin's earthworms (Annelida, Oligochaeta, Megadrilacea, Acanthodrilidae, Octochaetidae, Megascolecidae, Glossoscolecidae, Lumbricidae) with review of cosmopolitan Metaphire peguana - species group from Philippines. Opuscula Zoologic a, 47 (1), 09 - 30. http: // dx. doi. org / 10.18348 / opzool. 2016.1.9" type="journal article" year="2016">Blakemore 2016</bibRefCitation>
: 21
</treatmentCitation>
, fig. 8.
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</paragraph>
</subSubSection>
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<materialsCitation id="8DB3C27AFFA17917FF3C264E60AD8F6A" ID-GBIF-Occurrence="1288043366" box="[151,682,783,808]" country="Vietnam" location="Ba Ngoi" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<emphasis id="0FAF1435FFA17917FF3C264E63328F6A" bold="true" box="[151,309,783,808]" pageId="16" pageNumber="17">
<typeStatus id="E2607685FFA17917FF3C264E62D58F6A" box="[151,210,783,808]" pageId="16" pageNumber="17">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="45CC88B7FFA17917FE97264E639C8F6A" box="[316,411,783,808]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
(
<collectingRegion id="FF1F06C5FFA17917FE02264E60378F65" box="[425,560,783,808]" country="Vietnam" name="Khanh Hoa" pageId="16" pageNumber="17">Khanh Hoa</collectingRegion>
:
<location id="38049EFCFFA17917FD912651609A8F6A" LSID="urn:lsid:plazi:treatment:B5727931FFA17917FF3C2706612E880A:38049EFCFFA17917FD912651609A8F6A" box="[570,669,783,808]" country="Vietnam" name="Ba Ngoi" pageId="16" pageNumber="17" stateProvince="Khanh Hoa">Ba Ngoi</location>
).
</materialsCitation>
</paragraph>
<paragraph id="3D64C827FFA17917FF6C267560708F0E" blockId="16.[151,1436,783,1096]" box="[199,631,820,845]" pageId="16" pageNumber="17">
<materialsCitation id="8DB3C27AFFA17917FF6C267560708F0E" ID-GBIF-Occurrence="1288043382" box="[199,631,820,845]" collectionCode="MNHN" country="France" location="France" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<emphasis id="0FAF1435FFA17917FF6C267563728F0F" bold="true" box="[199,373,820,845]" pageId="16" pageNumber="17">
<typeStatus id="E2607685FFA17917FF6C267563058F0F" box="[199,258,820,845]" pageId="16" pageNumber="17">Type</typeStatus>
material.
</emphasis>
<collectionCode id="5BCA50E2FFA17917FED7267463D68F0E" box="[380,465,821,844]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
(576),
<collectingCountry id="45CC88B7FFA17917FD88267460748F0E" box="[547,627,821,844]" name="France" pageId="16" pageNumber="17">France</collectingCountry>
.
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</paragraph>
<paragraph id="3D64C827FFA17917FF6C261666488F32" blockId="16.[151,1436,783,1096]" box="[199,1103,855,880]" pageId="16" pageNumber="17">
<materialsCitation id="8DB3C27AFFA17917FF6C261666488F32" ID-GBIF-Occurrence="1288043385" box="[199,1103,855,880]" country="Vietnam" location="Khanh Hoa" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Khanh Hoa">
<emphasis id="0FAF1435FFA17917FF6C261660778F32" bold="true" box="[199,624,855,880]" pageId="16" pageNumber="17">
Records from
<collectingCountry id="45CC88B7FFA17917FEDA261663DE8F32" box="[369,473,855,880]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
.
<collectingRegion id="FF1F06C5FFA17917FE4E261660778F32" box="[485,624,855,880]" country="Vietnam" name="Khanh Hoa" pageId="16" pageNumber="17">Khanh Hoa</collectingRegion>
</emphasis>
(
<collectingRegion id="FF1F06C5FFA17917FDD4261960B18F32" box="[639,694,856,880]" country="Myanmar" name="Chin" pageId="16" pageNumber="17">Cam</collectingRegion>
Ranh: Ba Ngoi) (Michaelsen 1934).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFA17917FF6C263D60478FFA" pageId="16" pageNumber="17" type="distribution">
<paragraph id="3D64C827FFA17917FF6C263D609A8FD7" blockId="16.[151,1436,783,1096]" box="[199,669,892,917]" pageId="16" pageNumber="17">
<emphasis id="0FAF1435FFA17917FF6C263D63678FD7" bold="true" box="[199,352,892,917]" pageId="16" pageNumber="17">Distribution.</emphasis>
Only known from
<collectingCountry id="45CC88B7FFA17917FD9C263D609D8FD7" box="[567,666,892,917]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="3D64C827FFA17917FF6C26DE60478FFA" blockId="16.[151,1436,783,1096]" box="[199,576,927,952]" pageId="16" pageNumber="17">
<emphasis id="0FAF1435FFA17917FF6C26DE63A78FFA" bold="true" box="[199,416,927,952]" pageId="16" pageNumber="17">Vietnamese name.</emphasis>
Giun ba ngòi.
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFA17917FF6C2685612E880A" pageId="16" pageNumber="17" type="discussion">
<paragraph id="3D64C827FFA17917FF6C2685612E880A" blockId="16.[151,1436,783,1096]" pageId="16" pageNumber="17">
<emphasis id="0FAF1435FFA17917FF6C2685633C8F9F" bold="true" box="[199,315,964,989]" pageId="16" pageNumber="17">Remarks.</emphasis>
The species has been considered as a synonym of
<taxonomicName id="FADBB3A4FFA17917FC1D268566768F9F" box="[950,1137,964,989]" class="Oligochaeta" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="bahli">
<emphasis id="0FAF1435FFA17917FC1D268566768F9F" box="[950,1137,964,989]" italics="true" pageId="16" pageNumber="17">Metaphire bahli</emphasis>
</taxonomicName>
(see
<bibRefCitation id="594AB5D6FFA17917FB12268567328F9E" author="Thai" box="[1209,1333,964,989]" pageId="16" pageNumber="17" refString="Thai, T. B. (1983) Earthworms of Vietnam (Systematic, fauna, distribution and zoogeographic). PhD dissertation in Zooglogy. Lomonosov Moscow State University, Russia [in Russian]" type="book" year="1983">Thai 1983</bibRefCitation>
,
<bibRefCitation id="594AB5D6FFA17917FAEC268567968F9E" author="Thai" box="[1351,1425,964,988]" pageId="16" pageNumber="17" refString="Thai, T. B. (2000 a) Species diversity of earthworms in Vietnam. Proceedings of the national workshop on the basic issues in life science. Hanoi Science and Technics Publishing House, 307 - 311." type="proceedings paper" year="2000" yearSuffix="a">2000a</bibRefCitation>
). However,
<bibRefCitation id="594AB5D6FFA17917FEA626A6602A8842" author="Michaelsen" box="[269,557,999,1024]" pageId="16" pageNumber="17" refString="Michaelsen, W. (1934 b) Oligochaten von Franzosisch-Indochina. Archive de Zoologie Experimentale et Generale, 76, 493 - 546." type="journal article" year="1934" yearSuffix="b">Michaelsen (1934b: 508)</bibRefCitation>
states that male pores are simple and that &quot;Kopulationstaschen&quot; are absent, hence the species belongs to
<taxonomicName id="FADBB3A4FFA17917FE58214D60668867" box="[499,609,1036,1061]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="16" pageNumber="17" phylum="Annelida" rank="genus">
<emphasis id="0FAF1435FFA17917FE58214D60668867" box="[499,609,1036,1061]" italics="true" pageId="16" pageNumber="17">Amynthas</emphasis>
</taxonomicName>
rather than to
<taxonomicName id="FADBB3A4FFA17917FCBC214D618C8867" box="[791,907,1036,1061]" class="Oligochaeta" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="16" pageNumber="17" phylum="Annelida" rank="genus">
<emphasis id="0FAF1435FFA17917FCBC214D618C8867" box="[791,907,1036,1061]" italics="true" pageId="16" pageNumber="17">Metaphire</emphasis>
</taxonomicName>
. Furthermore, the genital markings are quite different from those typical of
<taxonomicName id="FADBB3A4FFA17917FE4421706122880A" authority="Blakemore 2016" authorityName="Blakemore" authorityYear="2016" box="[495,805,1071,1096]" class="Oligochaeta" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="bahli">
<emphasis id="0FAF1435FFA17917FE442170604A880A" box="[495,589,1071,1096]" italics="true" pageId="16" pageNumber="17">M. bahli</emphasis>
(
<bibRefCitation id="594AB5D6FFA17917FDF7216E611A880A" author="Blakemore" box="[604,797,1071,1096]" pageId="16" pageNumber="17" refString="Blakemore, R. J. (2016) Darwin's earthworms (Annelida, Oligochaeta, Megadrilacea, Acanthodrilidae, Octochaetidae, Megascolecidae, Glossoscolecidae, Lumbricidae) with review of cosmopolitan Metaphire peguana - species group from Philippines. Opuscula Zoologic a, 47 (1), 09 - 30. http: // dx. doi. org / 10.18348 / opzool. 2016.1.9" type="journal article" year="2016">Blakemore 2016</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>