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<document id="CCB81C42B18375954A539E09E26D7E30" ID-CLB-Dataset="8173" ID-DOI="10.5852/ejt.2021.735.1237" ID-GBIF-Dataset="4074b185-a219-4e7e-b1bd-1bc4bd3ef9e7" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542891" ID-ZooBank="8364CCA1-11A1-437F-944D-BA70C690D7F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613471756163" checkinUser="plazi" docAuthor="Rodríguez-Melgarejo, Maryzender &amp; Chani-Posse, Mariana" docDate="2021" docId="0A25E34BFFC2FFE59439FCB0FB34C117" docLanguage="en" docName="ejt-735_15-33.pdf" docOrigin="European Journal of Taxonomy 735" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophionthus asenjoi Rodríguez-Melgarejo &amp; Chani-Posse 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="26" masterDocId="F61C9B33FFCBFFEE964BFF85FF9BC553" masterDocTitle="On the rare Neotropical genus Ophionthus Bernhauer (Coleoptera: Staphylinidae: Staphylininae): redescription of the type species and description of a new species" masterLastPageNumber="33" masterPageNumber="15" pageNumber="24" updateTime="1698929669084" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B54A63F37736A6A48F94B6CDFFAE0A75">On the rare Neotropical genus Ophionthus Bernhauer (Coleoptera: Staphylinidae: Staphylininae): redescription of the type species and description of a new species</mods:title>
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<mods:namePart id="0AE30122E33C0A0E02198303592D80D8">Rodríguez-Melgarejo, Maryzender</mods:namePart>
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<mods:affiliation id="B90BE33055E0CD67E4E650BFEDED6BAF">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14 - 0434, Lima 14, Perú. Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina.</mods:affiliation>
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<mods:namePart id="39DE6B019ECF1F73846F6181309907EC">Chani-Posse, Mariana</mods:namePart>
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<mods:affiliation id="789D6EC92E46BF0EAB574E5240E38B8C">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14 - 0434, Lima 14, Perú. Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina.</mods:affiliation>
<mods:nameIdentifier id="100B3605A5EEA7AC6A525AD8F2509351" type="email">mchani@mendoza-conicet.gob.ar</mods:nameIdentifier>
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<mods:title id="1CF51ED18D3EF2EA587652E641F8AC4A">European Journal of Taxonomy</mods:title>
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<paragraph id="8233525DFFC2FFE79439FCB0FC59C603" blockId="9.[371,1217,821,918]" box="[626,962,821,848]" pageId="9" pageNumber="24">
<heading id="D97BE531FFC2FFE79439FCB0FC59C603" box="[626,962,821,848]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="24" reason="2">
<taxonomicName id="458C29DEFFC2FFE79439FCB0FCC4C61C" ID-CoL="8Y739" authority="Rodríguez-Melgarejo &amp; Chani-Posse, 2021" authorityName="Rodríguez-Melgarejo &amp; Chani-Posse" authorityYear="2021" box="[626,863,821,848]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="asenjoi" status="sp. nov.">
<emphasis id="B0F88E4FFFC2FFE79439FCB0FCC4C61C" bold="true" box="[626,863,821,848]" italics="true" pageId="9" pageNumber="24">Ophionthus asenjoi</emphasis>
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<taxonomicNameLabel id="ABCB3334FFC2FFE7952CFCB3FC59C603" box="[871,962,822,848]" pageId="9" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="CA9601D6FFC2FFE79493FCF9FCC7C6C5" box="[728,860,892,918]" pageId="9" pageNumber="24" type="description">
<paragraph id="8233525DFFC2FFE79493FCF9FCC7C6C5" blockId="9.[371,1217,821,918]" box="[728,860,892,918]" pageId="9" pageNumber="24">
<figureCitation id="1AB74ED8FFC2FFE79493FCF9FCDAC6C5" box="[728,833,892,918]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[189,232,1916,1942]" captionStartId-1="11.[189,232,1972,1998]" captionTargetBox-0="[202,1398,532,1882]" captionTargetBox-1="[189,1398,1237,1937]" captionTargetId-0="figure-102@10.[189,1398,532,1883]" captionTargetId-1="figure-257@11.[189,1399,1237,1937]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." captionText-1="Fig. 4. Distribution map of Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.4542901" figureDoi-1="http://doi.org/10.5281/zenodo.4542903" httpUri-0="https://zenodo.org/record/4542901/files/figure.png" httpUri-1="https://zenodo.org/record/4542903/files/figure.png" pageId="9" pageNumber="24">Figs 34</figureCitation>
,
<figureCitation id="1AB74ED8FFC2FFE79506FCF9FCC7C6C5" box="[845,860,892,918]" captionStart="Fig" captionStartId="13.[189,232,1815,1841]" captionTargetBox="[230,1362,326,1764]" captionTargetId="figure-11@13.[206,1382,310,1780]" captionTargetPageId="13" captionText="Fig. 6 (continued on next page). Strict consensus of the 28 MP trees obtained showing the systematic placement of Ophionthus Bernhauer, 1908 within the Neotropical lineage of Philonthina Kirby, 1837 (tree length = 579; consistency index (CI) = 0.24; retention index (RI) = 0.71. Black circles: exclusive synapomorphies; open circles, non-exclusive synapomorphies. Numbers above circles are character numbers; numbers below circles represent character states. Support values are indicated in bold below the branches as follows: Bremer/Standard Bootstrap/Jack-knife (cut = 50 for resampling support values)." figureDoi="http://doi.org/10.5281/zenodo.4545915" httpUri="https://zenodo.org/record/4545915/files/figure.png" pageId="9" pageNumber="24">6</figureCitation>
</paragraph>
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<subSubSection id="CA9601D6FFC2FFE796F6FC38FB7AC11F" pageId="9" pageNumber="24" type="diagnosis">
<paragraph id="8233525DFFC2FFE796F6FC38FEADC684" blockId="9.[189,310,957,983]" box="[189,310,957,983]" pageId="9" pageNumber="24">
<heading id="D97BE531FFC2FFE796F6FC38FEADC684" bold="true" box="[189,310,957,983]" fontSize="11" level="3" pageId="9" pageNumber="24" reason="3">
<emphasis id="B0F88E4FFFC2FFE796F6FC38FEADC684" bold="true" box="[189,310,957,983]" pageId="9" pageNumber="24">Diagnosis</emphasis>
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<paragraph id="8233525DFFC2FFE796F6FC69FB7AC11F" blockId="9.[187,1399,1004,1101]" pageId="9" pageNumber="24">
<taxonomicName id="458C29DEFFC2FFE796F6FC69FE39C155" authority="Rodríguez-Melgarejo &amp; Chani-Posse, 2021" authorityName="Rodríguez-Melgarejo &amp; Chani-Posse" authorityYear="2021" box="[189,418,1004,1030]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="asenjoi" status="sp. nov.">
<emphasis id="B0F88E4FFFC2FFE796F6FC69FE39C155" box="[189,418,1004,1030]" italics="true" pageId="9" pageNumber="24">Ophionthus asenjoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABCB3334FFC2FFE797E2FC69FD9FC155" box="[425,516,1004,1030]" pageId="9" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="458C29DEFFC2FFE794D5FC69FCD2C155" authorityName="Bernhauer" authorityYear="1908" box="[670,841,1004,1030]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="serpentinus">
<emphasis id="B0F88E4FFFC2FFE794D5FC69FCD2C155" box="[670,841,1004,1030]" italics="true" pageId="9" pageNumber="24">O. serpentinus</emphasis>
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by having the head with one pair of interocular punctures, antennae with the last three antennomeres much lighter than the preceding segments (
<figureCitation id="1AB74ED8FFC2FFE7935FFB8AFAF5C17A" box="[1300,1390,1039,1065]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="9" pageNumber="24">Fig. 3A</figureCitation>
) and the abdominal terga IV and V each without a distinct posterior basal transverse carina.
</paragraph>
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<subSubSection id="CA9601D6FFC2FFE796F6FBF1FAECC075" pageId="9" pageNumber="24" type="etymology">
<paragraph id="8233525DFFC2FFE796F6FBF1FEDFC1DD" blockId="9.[189,324,1140,1166]" box="[189,324,1140,1166]" pageId="9" pageNumber="24">
<heading id="D97BE531FFC2FFE796F6FBF1FEDFC1DD" bold="true" box="[189,324,1140,1166]" fontSize="11" level="3" pageId="9" pageNumber="24" reason="3">
<emphasis id="B0F88E4FFFC2FFE796F6FBF1FEDFC1DD" bold="true" box="[189,324,1140,1166]" pageId="9" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFC2FFE796F6FB27FAECC075" blockId="9.[187,1400,1186,1318]" pageId="9" pageNumber="24">
This new species is dedicated to the outstanding Peruvian coleopterist Angélico Asenjo, a great friend and colleague, in recognition of his tireless contribution to the study of
<taxonomicName id="458C29DEFFC2FFE795A7FB40FB16C18C" authorityName="Latreille" authorityYear="1802" box="[1004,1165,1221,1247]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="24" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
from Latin America and other regions, and as a small gesture of thanks for his continuous support of and collaboration with the first author, MR. This specific epithet is regarded as a Latinized masculine noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="CA9601D6FFC2FFE796F6FAC8FD67C315" pageId="9" pageNumber="24" type="materials_examined">
<paragraph id="8233525DFFC2FFE796F6FAC8FE31C034" blockId="9.[189,426,1357,1383]" box="[189,426,1357,1383]" pageId="9" pageNumber="24">
<heading id="D97BE531FFC2FFE796F6FAC8FE31C034" bold="true" box="[189,426,1357,1383]" fontSize="11" level="3" pageId="9" pageNumber="24" reason="3">
<emphasis id="B0F88E4FFFC2FFE796F6FAC8FE31C034" bold="true" box="[189,426,1357,1383]" pageId="9" pageNumber="24">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFC2FFE796A1FAF9FD67C315" blockId="9.[189,1399,1404,1606]" pageId="9" pageNumber="24">
<materialsCitation id="32E45800FFC2FFE796A1FAF9FD67C315" ID-GBIF-Occurrence="3037423301" collectingDate="28. VI" collectionCode="IFML" collectorName="W. Weyrauch" country="Peru" elevation="3000" location="PERU" pageId="9" pageNumber="24" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B0F88E4FFFC2FFE796A1FAF9FE96C0EA" bold="true" pageId="9" pageNumber="24">
<typeStatus id="5D37ECFFFFC2FFE796A1FAF9FEC7C0C5" box="[234,348,1404,1430]" pageId="9" pageNumber="24" type="holotype">Holotype</typeStatus>
<collectingCountry id="FA9B12CDFFC2FFE796F6FA1AFE96C0EA" box="[189,269,1439,1465]" name="Peru" pageId="9" pageNumber="24">PERU</collectingCountry>
</emphasis>
<specimenCount id="948A99D4FFC2FFE7976EFA25FEA0C0E9" box="[293,315,1440,1466]" pageId="9" pageNumber="24" type="female"></specimenCount>
[specimen glued to a white strip]; “N-PERU: / Bambamarca, /
<quantity id="4574FFB8FFC2FFE79255FA1AFBE8C0EA" box="[1054,1139,1439,1465]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="24" unit="m" value="3000.0">
<elevation id="09A1B56EFFC2FFE79255FA1AFBE8C0EA" box="[1054,1139,1439,1465]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="24" unit="m" value="3000.0">3000m</elevation>
</quantity>
,
<date id="F632749DFFC2FFE79235FA1BFB58C0EA" box="[1150,1219,1438,1465]" pageId="9" pageNumber="24">
<collectingDate id="E6768D75FFC2FFE79235FA1BFB58C0EA" box="[1150,1219,1438,1465]" pageId="9" pageNumber="24">28.VI</collectingDate>
</date>
[June]. [19]56 / leg.
<collectorName id="2F79378BFFC2FFE796A4FA46FE0BC08F" box="[239,400,1474,1501]" pageId="9" pageNumber="24">W. Weyrauch</collectorName>
” [typewritten on white label], “ex- col / Weyrauch [printed on white label with black borders]”, “COLECCION / Fundación M. Lillo / 4000 S. M. Tucumán / TUCUMAN - ARGENTINA [printed on white greenish label]”, “HOLOTYPE /
<taxonomicName id="458C29DEFFC2FFE79573F98CFBB9C371" authorityName="Rodríguez-Melgarejo &amp; Chani-Posse" authorityYear="2021" box="[824,1058,1545,1571]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="asenjoi">
<emphasis id="B0F88E4FFFC2FFE79573F98CFBB9C371" box="[824,1058,1545,1571]" italics="true" pageId="9" pageNumber="24">Ophionthus asenjoi</emphasis>
</taxonomicName>
/ Rodríguez-Melgarejo &amp; / Chani-Posse 2020 [printed on red label]”;
<collectionCode id="E49DCA98FFC2FFE794E5F9A9FD63C315" box="[686,760,1580,1606]" collectionName="IFML" pageId="9" pageNumber="24">IFML</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA9601D6FFC2FFE596F6F9E8FB34C117" lastPageId="11" lastPageNumber="26" pageId="9" pageNumber="24" type="description">
<paragraph id="8233525DFFC2FFE796F6F9E8FED5C3D4" blockId="9.[189,334,1645,1671]" box="[189,334,1645,1671]" pageId="9" pageNumber="24">
<heading id="D97BE531FFC2FFE796F6F9E8FED5C3D4" bold="true" box="[189,334,1645,1671]" fontSize="11" level="3" pageId="9" pageNumber="24" reason="3">
<emphasis id="B0F88E4FFFC2FFE796F6F9E8FED5C3D4" bold="true" box="[189,334,1645,1671]" pageId="9" pageNumber="24">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFC2FFE796F6F919FDD8C3E5" blockId="9.[189,579,1692,1719]" box="[189,579,1692,1719]" pageId="9" pageNumber="24">
DIMENSIONS. BL = 10.5 (
<figureCitation id="1AB74ED8FFC2FFE79793F919FDADC3E5" box="[472,566,1692,1718]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="9" pageNumber="24">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8233525DFFC2FFE796F6F958FB1EC26D" blockId="9.[189,1398,1757,1854]" pageId="9" pageNumber="24">
COLOURATION. Head, thorax, and abdomen piceous, except apical half of tergum VII, apical third of sternum VII, and segments VIII and IX yellow; mandibles, maxillary and labial palps, elytra and legs dark brown; antennomeres 18 dark brown to black and 911 yellowish (
<figureCitation id="1AB74ED8FFC2FFE79252F8A1FBECC26D" box="[1049,1143,1828,1854]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="9" pageNumber="24">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8233525DFFC2FFE496F6F8E3FB1FC4E3" blockId="9.[185,1398,1893,1990]" lastBlockId="10.[187,1400,265,432]" lastPageId="10" lastPageNumber="25" pageId="9" pageNumber="24">HEAD. Wider than long (HW/HL =1.2),moderately wider than pronotum (HW/PW =1.2). Eyes moderately shorter than temples (YL/TL = 0.8) seen from above. Epicranium with one pair of interocular punctures; each side of vertex with 45 postocular punctures. Antennae with antennomere 1 more than 2× as long as antennomere 2 (A1/A2 = 2.2) and as long as antennomeres 2 and 3 combined (A1/(A2+A3) = 1), antennomere 2 distinctly shorter than antennomere 3 (A2/A3 = 0.7). Labial palpus with palpomere 2 1.4× as long as palpomere 1 and 0.6× as long as palpomere 3. Maxillary palpus with palpomere 1 distinctly short; palpomere 2 curved, 2× as long as palpomere 3; palpomere 4 slightly longer than palpomere 2 and about 2× as long as palpomere 3. Neck about 0.4× as wide as head at widest point.</paragraph>
<caption id="D6F302D5FFC1FFE496F6F8F9FA8CC2E9" ID-DOI="http://doi.org/10.5281/zenodo.4542901" ID-Zenodo-Dep="4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="10" pageNumber="25" startId="10.[189,232,1916,1942]" targetBox="[202,1398,532,1882]" targetPageId="10">
<paragraph id="8233525DFFC1FFE496F6F8F9FA8CC2E9" blockId="10.[189,1399,1916,1979]" pageId="10" pageNumber="25">
<emphasis id="B0F88E4FFFC1FFE496F6F8F9FE97C2C5" bold="true" box="[189,268,1916,1942]" pageId="10" pageNumber="25">Fig. 3.</emphasis>
<taxonomicName id="458C29DEFFC1FFE4975CF8F8FE65C2C5" authority="Rodríguez-Melgarejo &amp; Chani-Posse, 2021" authorityName="Rodríguez-Melgarejo &amp; Chani-Posse" authorityYear="2021" box="[279,510,1917,1943]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="asenjoi" status="sp. nov.">
<emphasis id="B0F88E4FFFC1FFE4975CF8F8FE65C2C5" box="[279,510,1917,1943]" italics="true" pageId="10" pageNumber="25">Ophionthus asenjoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABCB3334FFC1FFE4944CF8F8FDFFC2C4" box="[519,612,1917,1943]" pageId="10" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, ♀ (IFML).
<emphasis id="B0F88E4FFFC1FFE49539F8F9FC1CC2C5" bold="true" box="[882,903,1916,1942]" pageId="10" pageNumber="25">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B0F88E4FFFC1FFE492D6F8F8FB34C2C4" bold="true" box="[1181,1199,1917,1943]" pageId="10" pageNumber="25">B</emphasis>
. Original labels.
<emphasis id="B0F88E4FFFC1FFE496F4F825FF49C2E9" bold="true" box="[191,210,1952,1978]" pageId="10" pageNumber="25">C</emphasis>
. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8233525DFFC0FFE596F6FE8FFDFDC414" blockId="11.[189,1398,265,327]" pageId="11" pageNumber="26">PRONOTUM. Longer than wide (PL/PW = 1.3), with two dorsal rows of four punctures each on disk of pronotum, 34 sublateral punctures.</paragraph>
<paragraph id="8233525DFFC0FFE596F6FEF3FC4AC4DC" blockId="11.[189,977,373,400]" box="[189,977,373,400]" pageId="11" pageNumber="26">
LEGS. First metatarsomere longer than fifth (S1/S5 = 1.7) (
<figureCitation id="1AB74ED8FFC0FFE5952CFEF0FC5FC4DC" box="[871,964,373,399]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="11" pageNumber="26">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8233525DFFC0FFE596F6FE3AFA8EC48A" blockId="11.[189,1301,446,473]" box="[189,1301,446,473]" pageId="11" pageNumber="26">ELYTRON. Longer than wide (EL/EW = 1.6), at sides about 1.6× as long as elytron along suture.</paragraph>
<paragraph id="8233525DFFC0FFE596F6FD82FC63C771" blockId="11.[189,1016,519,546]" box="[189,1016,519,546]" pageId="11" pageNumber="26">ABDOMEN. Posterior basal transverse carina distinct only on tergum III.</paragraph>
<paragraph id="8233525DFFC0FFE596F6FDD4FDEAC739" blockId="11.[189,625,592,619]" box="[189,625,592,619]" pageId="11" pageNumber="26">MALE SEXUAL CHARACTERS. Unknown.</paragraph>
<paragraph id="8233525DFFC0FFE596F6FD1FFC05C7E0" blockId="11.[189,926,665,692]" box="[189,926,665,692]" pageId="11" pageNumber="26">
FEMALE SEXUAL CHARACTERS. Similar to those of
<emphasis id="B0F88E4FFFC0FFE594A0FD1FFC05C7E0" box="[747,926,666,692]" italics="true" pageId="11" pageNumber="26">
<taxonomicName id="458C29DEFFC0FFE594A0FD1FFC01C7E0" authorityName="Bernhauer" authorityYear="1908" box="[747,922,666,692]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="serpentinus">O. serpentinus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8233525DFFC0FFE596F6FD67FD24C7AF" blockId="11.[189,703,738,764]" box="[189,703,738,764]" pageId="11" pageNumber="26">
<heading id="D97BE531FFC0FFE596F6FD67FD24C7AF" bold="true" box="[189,703,738,764]" fontSize="11" level="3" pageId="11" pageNumber="26" reason="3">
<emphasis id="B0F88E4FFFC0FFE596F6FD67FD24C7AF" bold="true" box="[189,703,738,764]" pageId="11" pageNumber="26">Geographical distribution and bionomics</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFC0FFE596F6FC94FB34C117" blockId="11.[189,1399,785,1093]" pageId="11" pageNumber="26">
<taxonomicName id="458C29DEFFC0FFE596F6FC94FE39C679" authority="Rodríguez-Melgarejo &amp; Chani-Posse, 2021" authorityName="Rodríguez-Melgarejo &amp; Chani-Posse" authorityYear="2021" box="[189,418,785,811]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="asenjoi" status="sp. nov.">
<emphasis id="B0F88E4FFFC0FFE596F6FC94FE39C679" box="[189,418,785,811]" italics="true" pageId="11" pageNumber="26">Ophionthus asenjoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABCB3334FFC0FFE597E2FC94FD98C678" box="[425,515,785,811]" pageId="11" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
is only known at present from its type locality in Bambamarca, department of
<collectingRegion id="40489CBFFFC0FFE59694FCB1FEFBC61D" box="[223,352,820,846]" country="Peru" name="Cajamarca" pageId="11" pageNumber="26">Cajamarca</collectingRegion>
, Northern
<collectingCountry id="FA9B12CDFFC0FFE597A9FCB1FD83C61D" box="[482,536,820,846]" name="Peru" pageId="11" pageNumber="26">Peru</collectingCountry>
at
<quantity id="4574FFB8FFC0FFE5940AFCB1FD06C61D" box="[577,669,820,846]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="26" unit="m" value="3000.0">3000 m</quantity>
[m a.s.l.] (
<figureCitation id="1AB74ED8FFC0FFE5956EFCB1FCEAC61D" box="[805,881,820,846]" captionStart="Fig" captionStartId="11.[189,232,1972,1998]" captionTargetBox="[189,1398,1237,1937]" captionTargetId="figure-257@11.[189,1399,1237,1937]" captionTargetPageId="11" captionText="Fig. 4. Distribution map of Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4542903" httpUri="https://zenodo.org/record/4542903/files/figure.png" pageId="11" pageNumber="26">Fig. 4</figureCitation>
). In addition, it is known that on the same collection day as that of the
<typeStatus id="5D37ECFFFFC0FFE59453FCD2FDE4C622" box="[536,639,855,881]" pageId="11" pageNumber="26" type="holotype">holotype</typeStatus>
of
<taxonomicName id="458C29DEFFC0FFE594E0FCDDFCBDC622" authority="Rodríguez-Melgarejo &amp; Chani-Posse, 2021" authorityName="Rodríguez-Melgarejo &amp; Chani-Posse" authorityYear="2021" box="[683,806,856,882]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="asenjoi" status="sp. nov.">
<emphasis id="B0F88E4FFFC0FFE594E0FCDDFCBDC622" box="[683,806,856,882]" italics="true" pageId="11" pageNumber="26">O. asenjoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABCB3334FFC0FFE5957BFCDDFC16C621" box="[816,909,856,882]" pageId="11" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
(i.e.,
<date id="F632749DFFC0FFE5959DFCD2FBEEC621" box="[982,1141,855,882]" pageId="11" pageNumber="26" value="1956-06-28">28 Jun. 1956</date>
), Weyrauch collected two specimens of
<emphasis id="B0F88E4FFFC0FFE597D6FCFEFD02C6C7" box="[413,665,891,917]" italics="true" pageId="11" pageNumber="26">
<collectingRegion id="40489CBFFFC0FFE597D6FCFEFDBBC6C6" box="[413,544,891,917]" country="Peru" name="Cajamarca" pageId="11" pageNumber="26">Cajamarca</collectingRegion>
triseriata
</emphasis>
<bibRefCitation id="E61D2FACFFC0FFE594EFFCFEFCD7C6C6" author="Roewer C. F." box="[676,844,890,917]" pageId="11" pageNumber="26" pagination="67 - 94" refId="ref8656" refString="Roewer C. F. 1957. Arachnida Arthrogastra aus Peru, III. Senckenbergiana biologica 38 (1): 67 - 94." type="journal article" year="1957">Roewer, 1957</bibRefCitation>
(Arachnida: Opiliones) in “Nord-Peru: Cerro Macheipungo,
<quantity id="4574FFB8FFC0FFE59738FC1BFE2FC6E4" box="[371,436,925,952]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="26" unit="km" value="4.0">4 km</quantity>
NW Bambamarca” (
<bibRefCitation id="E61D2FACFFC0FFE594F6FC1BFC03C6EB" author="Roewer C. F." box="[701,920,925,952]" pageId="11" pageNumber="26" pagination="67 - 94" refId="ref8656" refString="Roewer C. F. 1957. Arachnida Arthrogastra aus Peru, III. Senckenbergiana biologica 38 (1): 67 - 94." type="journal article" year="1957">Roewer 1957: 75</bibRefCitation>
), which is georeferenced as
<geoCoordinate id="E7B8349AFFC0FFE5934FFC19FAEAC6EB" box="[1284,1393,924,952]" degrees="06" direction="south" minutes="41" orientation="latitude" pageId="11" pageNumber="26" precision="925" value="-6.6833334">06°41 S</geoCoordinate>
,
<geoCoordinate id="E7B8349AFFC0FFE596F6FC3AFEB4C688" box="[189,303,959,987]" degrees="78" direction="west" minutes="32" orientation="longitude" pageId="11" pageNumber="26" precision="925" value="-78.53333">78°32 W</geoCoordinate>
(
<bibRefCitation id="E61D2FACFFC0FFE5970AFC44FE60C688" author="Breure A." box="[321,507,960,987]" pageId="11" pageNumber="26" pagination="4 - 24" refId="ref7179" refString="Breure A. 2012. Weyrauch's type localities: a clarification; with illustrations of types of Orthalicoidea (Mollusca, Gastropoda, Stylommatophora) in the Tucuman museum. Folia Conchyliologica 17: 4 - 24." type="journal article" year="2012">Breure 2012: 5</bibRefCitation>
). Therefore, it is highly probable that this locality is the same or close to where the
<typeStatus id="5D37ECFFFFC0FFE5970BFC61FE3CC6AD" box="[320,423,996,1022]" pageId="11" pageNumber="26" type="holotype">holotype</typeStatus>
of
<taxonomicName id="458C29DEFFC0FFE5979DFC61FDC8C6AD" authority="Rodríguez-Melgarejo &amp; Chani-Posse, 2021" authorityName="Rodríguez-Melgarejo &amp; Chani-Posse" authorityYear="2021" box="[470,595,996,1022]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="asenjoi" status="sp. nov.">
<emphasis id="B0F88E4FFFC0FFE5979DFC61FDC8C6AD" box="[470,595,996,1022]" italics="true" pageId="11" pageNumber="26">O. asenjoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABCB3334FFC0FFE59414FC61FD25C6AD" box="[607,702,996,1022]" pageId="11" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
was collected. Bambamarca belongs to the ecoregion of Peruvian Yungas (
<bibRefCitation id="E61D2FACFFC0FFE597DDFB82FDF0C172" author="Olson D. M. &amp; Dinerstein E. &amp; Wikramanayake E. D. &amp; Burgess N. D. &amp; Powell G. V. &amp; Underwood E. C. &amp; D'Amico J. &amp; Itoua I. &amp; Strand H. E. &amp; Morrison J. C. &amp; Loucks C. J. &amp; Allnutt T. F. &amp; Ricketts T. H. &amp; Kura Y. &amp; Lamoreux J. F. &amp; Wettengel W. W. &amp; Hedaeo P. &amp; Kassen K. R." box="[406,619,1031,1057]" pageId="11" pageNumber="26" pagination="933 - 938" refId="ref8386" refString="Olson D. M., Dinerstein E., Wikramanayake E. D., Burgess N. D., Powell G. V., Underwood E. C., D'Amico J., Itoua I., Strand H. E., Morrison J. C., Loucks C. J., Allnutt T. F., Ricketts T. H., Kura Y., Lamoreux J. F., Wettengel W. W., Hedaeo P. &amp; Kassen K. R. 2001. Terrestrial ecoregions of the world: a new map of life on earth: a new global map of terrestrial ecoregions provides an innovative tool for conserving biodiversity. BioScience 51 (11): 933 - 938. https: // doi. org / 10.1641 / 0006 - 3568 (2001) 051 [0933: TEOTWA] 2.0. CO; 2" type="journal article" year="2001">
Olson
<emphasis id="B0F88E4FFFC0FFE597A1FB8DFDBCC172" box="[490,551,1031,1057]" italics="true" pageId="11" pageNumber="26">et al.</emphasis>
2001
</bibRefCitation>
) or Puna province (
<bibRefCitation id="E61D2FACFFC0FFE59516FB82FB97C172" author="Morrone J. J." box="[861,1036,1031,1057]" pageId="11" pageNumber="26" pagination="1 - 110" refId="ref8350" refString="Morrone J. J. 2014. Biogeographical regionalisation of the Neotropical region. Zootaxa 3782 (1): 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
) and is nowadays agricultural land, surrounded by an urban zone and Andean shrubland ecosystem (
<bibRefCitation id="E61D2FACFFC0FFE595BAFBAFFB38C117" author="MINAM" box="[1009,1187,1066,1092]" pageId="11" pageNumber="26" refId="ref8217" refString="MINAM [Ministerio del Ambiente del Peru]. 2019. Mapa nacional de ecosistemas del Peru. Available from mapa-nacional-de-ecosistemas-del-peru [accessed 3 Jun. 2020]." type="book" year="2019">MINAM 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>