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<document id="314E714203D80264948F393EC77F5BD8" ID-DOI="10.11646/zootaxa.3683.2.3" ID-GBIF-Dataset="bb1cc4ed-5a7d-4c04-a042-2deb834b1d21" ID-ISSN="1175-5326" ID-Zenodo-Dep="219725" ID-ZooBank="1CDD7727-E1BF-4C46-B8FD-A13522FFC698" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460644285648" checkinUser="plazi" docAuthor="Dominiak, Patrycja &amp; Alwin, Alicja" docDate="2013" docId="081B3D0DFFF8FFC342AB310800108DC1" docLanguage="en" docName="zt03683p144.pdf" docOrigin="Zootaxa 3683 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea nauta Dominiak, sp. nov." docType="treatment" docUuid="2733A340-8ADE-46EF-AC4B-7504DC249E61" docUuidSource="ZooBank" docVersion="10" lastPageNumber="139" masterDocId="F4224575FFFEFFC5423C302F02178965" masterDocTitle="Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)" masterLastPageNumber="144" masterPageNumber="133" pageNumber="139" updateTime="1698336543250" updateUser="plazi">
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<mods:title id="F38CD818AF070850D6E36C12952E503F">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
</mods:titleInfo>
<mods:name id="44A6F8511F1E08EB90C11713B4D90B06" type="personal">
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<mods:namePart id="684F5E64809344424B63A6E5A38B533B">Dominiak, Patrycja</mods:namePart>
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<mods:namePart id="E72E904F80B2D9CF4DCDB0017CAA7364">Alwin, Alicja</mods:namePart>
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<mods:title id="7F8D0851A19CD45E84F38D24E0BB0630">Zootaxa</mods:title>
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<mods:part id="14A9CA28915148345F8DEB5E2AE2819C">
<mods:date id="1DDB010FF959A17F20A644A9127A7027">2013</mods:date>
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<mods:number id="E5EA17C20CD157B87C45BE22095FE7AF">3683</mods:number>
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<mods:identifier id="C617C0FDF790CBC7D9D82C61B9D48E40" type="DOI">10.11646/zootaxa.3683.2.3</mods:identifier>
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<treatment id="081B3D0DFFF8FFC342AB310800108DC1" ID-DOI="http://doi.org/10.5281/zenodo.4500016" ID-GBIF-Taxon="119571715" ID-Zenodo-Dep="4500016" LSID="urn:lsid:zoobank.org:act:2733A340-8ADE-46EF-AC4B-7504DC249E61" httpUri="http://treatment.plazi.org/id/081B3D0DFFF8FFC342AB310800108DC1" lastPageNumber="139" pageId="6" pageNumber="139">
<subSubSection id="C8A8DF90FFF8FFC342AB3108030E8807" pageId="6" pageNumber="139" type="nomenclature">
<paragraph id="800D8C1BFFF8FFC342AB3108005E8824" blockId="6.[151,585,295,354]" box="[151,585,295,321]" pageId="6" pageNumber="139">
<heading id="DB453B77FFF8FFC342AB3108005E8824" bold="true" box="[151,585,295,321]" fontSize="11" level="1" pageId="6" pageNumber="139" reason="1">
<emphasis id="B2C65009FFF8FFC342AB3108005E8824" bold="true" box="[151,585,295,321]" pageId="6" pageNumber="139">
<taxonomicName id="47B2F798FFF8FFC342AB310803F38824" ID-CoL="9HN7L" authority="Dominiak" authorityName="Dominiak" box="[151,484,295,321]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="nauta" status="sp. nov.">
<emphasis id="B2C65009FFF8FFC342AB310803778824" bold="true" box="[151,352,295,321]" italics="true" pageId="6" pageNumber="139">Dasyhelea nauta</emphasis>
Dominiak
</taxonomicName>
<taxonomicNameLabel id="A9F5ED72FFF8FFC343D73108005E8824" box="[491,585,295,321]" pageId="6" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="800D8C1BFFF8FFC342AB3165030E8807" blockId="6.[151,585,295,354]" box="[151,281,330,354]" pageId="6" pageNumber="139">
(
<figureCitation id="1889909EFFF8FFC342A3316503068807" box="[159,273,330,354]" captionStart="FIGURES 9 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 15. Dasyhelea nauta Dominiak sp. nov. (9 12), holotype male: 9. flagellomeres 9 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Figs 912</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342AB31BC03B38897" pageId="6" pageNumber="139" type="materials_examined">
<paragraph id="800D8C1BFFF8FFC342AB31BC03B38897" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342AB31BC035288CE" bold="true" box="[151,325,402,427]" pageId="6" pageNumber="139">
<typeStatus id="5F0932B9FFF8FFC342AB31BC02C788CE" box="[151,208,403,427]" pageId="6" pageNumber="139">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFF8FFC3437331BD03AB88CF" box="[335,444,402,426]" pageId="6" pageNumber="139" type="holotype">Holotype</typeStatus>
: adult male.
<emphasis id="B2C65009FFF8FFC3406D31BC00B188CE" bold="true" box="[593,678,403,427]" pageId="6" pageNumber="139">Ye m e n.</emphasis>
Al Mukalla, N 14˚33' E 49˚08',
<date id="F40CAADBFFF8FFC3461D31BD06DE88CF" box="[1057,1225,402,426]" pageId="6" pageNumber="139" value="2003-07" valueMax="2003-08-31" valueMin="2003-07-00">VIIVIII.2003</date>
, light trap, leg. A. van Harten (UG).
<typeStatus id="5F0932B9FFF8FFC3435D319803C588AA" box="[353,466,439,463]" pageId="6" pageNumber="139" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B2C65009FFF8FFC343E03198002688AA" bold="true" box="[476,561,439,463]" pageId="6" pageNumber="139">
<collectingCountry id="F8A5CC8BFFF8FFC343E03198003D88AA" box="[476,554,439,463]" name="Yemen" pageId="6" pageNumber="139">Yemen</collectingCountry>
.
</emphasis>
Al Mukalla, N 14˚33' E 49˚08',
<date id="F40CAADBFFF8FFC341A73199065588AB" box="[923,1090,438,462]" pageId="6" pageNumber="139" value="2003-07" valueMax="2003-08-31" valueMin="2003-07-00">VIIVIII.2003</date>
, light trap,
<specimenCount id="96B44792FFF8FFC346833199070088AB" box="[1215,1303,438,462]" pageId="6" pageNumber="139" type="male">3 males</specimenCount>
, leg. A. van Harten (
<collectionCode id="E6A314DEFFF8FFC342CD31F503508897" box="[241,327,474,498]" pageId="6" pageNumber="139">BMNH</collectionCode>
&amp; UG).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB31D101EC8B5F" pageId="6" pageNumber="139" type="diagnosis">
<paragraph id="800D8C1BFFF8FFC342FB31D101EC8B5F" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB31D103568B72" bold="true" box="[199,321,510,535]" pageId="6" pageNumber="139">Diagnosis.</emphasis>
Adult male of
<taxonomicName id="47B2F798FFF8FFC3403C31D100D48B72" box="[512,707,510,535]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="nauta">
<emphasis id="B2C65009FFF8FFC3403C31D100D48B72" box="[512,707,510,535]" italics="true" pageId="6" pageNumber="139">Dasyhelea nauta</emphasis>
</taxonomicName>
has unique eight-shaped median projections of aedeagus and symmetrical parameres. It is also distinct in having short distal flagellomeres.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB326906078D71" pageId="6" pageNumber="139" type="description">
<paragraph id="800D8C1BFFF8FFC342FB326902CF8ACD" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB326903B18B3A" bold="true" box="[199,422,582,607]" pageId="6" pageNumber="139">Description. Male.</emphasis>
Head. Flagellum length
<quantity id="474A21FEFFF8FFC340F6326901658B3B" box="[714,882,582,606]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="6" pageNumber="139" unit="mm" value="0.48" valueMax="0.51" valueMin="0.45">0.450.51 mm</quantity>
, AR 0.580.66 (n=4). Four distal flagellomeres short, with one row of long setae (
<figureCitation id="1889909EFFF8FFC3402F324500408BE7" box="[531,599,618,642]" captionStart="FIGURES 9 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 15. Dasyhelea nauta Dominiak sp. nov. (9 12), holotype male: 9. flagellomeres 9 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 9</figureCitation>
). Last flagellomere without apical prolongation. Clypeus composed of two sclerites, with 56 strong setae. Maxillary palp (
<figureCitation id="1889909EFFF8FFC3409232A100E98BC3" box="[686,766,654,678]" captionStart="FIGURES 9 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 15. Dasyhelea nauta Dominiak sp. nov. (9 12), holotype male: 9. flagellomeres 9 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 10</figureCitation>
) with third palpal segment cylindrical, 4354 μm long, PR III 2.743.19 (n=4). Thorax. Transverse suture absent, number of supraalar setae vary from 6 to 7. Scutellum dark, with 6 strong setae. Wing with one nearly obliterated radial cell. Wing length
<quantity id="474A21FEFFF8FFC3463D32F606B18B94" box="[1025,1190,729,753]" metricMagnitude="-4" metricUnit="m" metricValue="8.3" metricValueMax="8.6" metricValueMin="8.0" pageId="6" pageNumber="139" unit="mm" value="0.83" valueMax="0.86" valueMin="0.8">0.800.86 mm</quantity>
, CR 0.480.50 (n=4). Legs uniform in coloration,
<collectingCountry id="F8A5CC8BFFF8FFC343D732D100198A70" box="[491,526,766,789]" name="Chile" pageId="6" pageNumber="139">TR</collectingCountry>
I 2.52.6 (n=4),
<collectingCountry id="F8A5CC8BFFF8FFC340E932D100EE8A70" box="[725,761,766,789]" name="Chile" pageId="6" pageNumber="139">TR</collectingCountry>
II 2.63.0 (n=4),
<collectingCountry id="F8A5CC8BFFF8FFC341F932D101FE8A70" box="[965,1001,766,789]" name="Chile" pageId="6" pageNumber="139">TR</collectingCountry>
III 2.32.4 (n=4). Genitalia (
<figureCitation id="1889909EFFF8FFC3470A32D207998A70" box="[1334,1422,765,789]" captionStart="FIGURES 9 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 15. Dasyhelea nauta Dominiak sp. nov. (9 12), holotype male: 9. flagellomeres 9 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 11</figureCitation>
). Posterior margin of tergite 9 rounded, with a pair of short apicolateral processes. Sternite 9 slightly expanded. Gonostylus long and slender, gently arched. Parameres symmetrical, stout basally and very thin in apical part, fused (
<figureCitation id="1889909EFFF8FFC342DA3343032A8AE1" box="[230,317,876,900]" captionStart="FIGURES 9 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 15. Dasyhelea nauta Dominiak sp. nov. (9 12), holotype male: 9. flagellomeres 9 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 12</figureCitation>
). Median projections of aedeagus strongly sclerotized, eight-shaped, with short and sharp apical tooth.
</paragraph>
<paragraph id="800D8C1BFFF8FFC342FB339A00498AA9" blockId="6.[151,1437,402,1188]" box="[199,606,948,973]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB339A03F68AA8" bold="true" box="[199,481,948,973]" pageId="6" pageNumber="139">Female and immatures.</emphasis>
Unknown.
</paragraph>
<paragraph id="800D8C1BFFF8FFC342FB33F806078D71" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB33F803F58A95" bold="true" box="[199,482,983,1008]" pageId="6" pageNumber="139">Derivation of the name.</emphasis>
The Latin word
<taxonomicName id="47B2F798FFF8FFC3409C33F600F78A95" box="[672,736,985,1008]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="nauta">
<emphasis id="B2C65009FFF8FFC3409C33F600F78A95" box="[672,736,985,1008]" italics="true" pageId="6" pageNumber="139">nauta</emphasis>
</taxonomicName>
,-
<emphasis id="B2C65009FFF8FFC340CC33F6011C8A95" box="[752,779,985,1008]" italics="true" pageId="6" pageNumber="139">ae</emphasis>
means sailor or shipowner. The species is named after the
<typeStatus id="5F0932B9FFF8FFC342AB33D202D08D70" box="[151,199,1021,1045]" pageId="6" pageNumber="139">type</typeStatus>
locality, Al Mukalla, an important sea port in the province of Hadhramaut.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB343003AE8D5D" box="[199,441,1055,1080]" pageId="6" pageNumber="139" type="distribution">
<paragraph id="800D8C1BFFF8FFC342FB343003AE8D5D" blockId="6.[151,1437,402,1188]" box="[199,441,1055,1080]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB343003778D5D" bold="true" box="[199,352,1055,1080]" pageId="6" pageNumber="139">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFF8FFC3435B340E03A58D5D" box="[359,434,1057,1080]" name="Yemen" pageId="6" pageNumber="139">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB346B00108DC1" pageId="6" pageNumber="139" type="discussion">
<paragraph id="800D8C1BFFF8FFC342FB346B00108DC1" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB346B03588D38" bold="true" box="[199,335,1092,1117]" pageId="6" pageNumber="139">Comments.</emphasis>
The characteristic armature of male genitalia, including the shape of both aedeagus and parameres, distinguishes this species from all other
<taxonomicName id="47B2F798FFF8FFC3413A3447016B8DE5" box="[774,892,1128,1152]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B2C65009FFF8FFC3413A3447016B8DE5" box="[774,892,1128,1152]" italics="true" pageId="6" pageNumber="139">Dasyhelea</emphasis>
</taxonomicName>
. The built of antenna, with very short distal flagellomeres, is also distinctive.
</paragraph>
</subSubSection>
</treatment>
</document>