treatments-xml/data/03/D5/74/03D574268E5EFFBEFF1AFCCADA40FDD5.xml
2024-06-21 12:22:17 +02:00

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XML

<document id="6E815C93487161D39525DDDAB0048794" ID-CLB-Dataset="55717" ID-DOI="10.5281/zenodo.5341014" ID-GBIF-Dataset="b9741f08-928d-4423-9e6f-478cebcd02cc" ID-ISSN="0253-116X" ID-Zenodo-Dep="5341014" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630189109691" checkinUser="marcus" docAuthor="Assing, V., Schülke, M., M. R &amp; A. S" docDate="2013" docId="03D574268E5EFFBEFF1AFCCADA40FDD5" docLanguage="en" docName="LinzerbiolBeitr.2013.45.1.141-154.pdf" docOrigin="Linzer biologische Beiträge 45 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Pinophilinus spinosus Assing &amp; Schülke &amp; M.R &amp; A.S 2013, nov.sp." docType="treatment" docVersion="9" lastPageNumber="149" masterDocId="FFEC0C5E8E5BFFB6FFB9FFFBD97DFFE6" masterDocTitle="On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera)" masterLastPageNumber="154" masterPageNumber="141" pageNumber="146" updateTime="1699181010772" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A8750996948C6ADCD174D267913EAADF">On the Staphylinidae of Saudi Arabia, with descriptions of two new species (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="D80C088361D27EE2B7C20C0AC005494C">Assing, V.</mods:namePart>
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<mods:namePart id="5D1315744AB427422D7FDEF490309DB5">Schülke, M.</mods:namePart>
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<mods:namePart id="D0CC9D89424AF4B84BAF79BF6490E734">M. R</mods:namePart>
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<mods:namePart id="43F0912D64E893AFD26FB5729ED32293">A. S</mods:namePart>
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<treatment id="03D574268E5EFFBEFF1AFCCADA40FDD5" ID-DOI="http://doi.org/10.5281/zenodo.6521790" ID-GBIF-Taxon="195191924" ID-Zenodo-Dep="6521790" LSID="urn:lsid:plazi:treatment:03D574268E5EFFBEFF1AFCCADA40FDD5" httpUri="http://treatment.plazi.org/id/03D574268E5EFFBEFF1AFCCADA40FDD5" lastPageId="8" lastPageNumber="149" pageId="5" pageNumber="146">
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<paragraph id="8BC3C5308E5EFFB3FF1AFCCADB4DFCA1" blockId="5.[163,745,817,839]" box="[163,560,817,839]" pageId="5" pageNumber="146">
<heading id="D08B725C8E5EFFB3FF1AFCCADB4DFCA1" box="[163,560,817,839]" fontSize="8" level="2" pageId="5" pageNumber="146" reason="2">
<emphasis id="B90819228E5EFFB3FF1AFCCADB4DFCA1" bold="true" box="[163,560,817,839]" pageId="5" pageNumber="146">
<taxonomicName id="4C7CBEB38E5EFFB3FF1AFCCAD8A2FCA1" ID-CoL="77LDH" authority="ASSING" authorityName="Assing &amp; Schülke &amp; M.R &amp; A.S" authorityYear="2013" box="[163,479,817,839]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="B90819228E5EFFB3FF1AFCCAD8F7FCA1" bold="true" box="[163,394,817,839]" italics="true" pageId="5" pageNumber="146">Pinophilinus spinosus</emphasis>
ASSING
</taxonomicName>
<taxonomicNameLabel id="A23BA4598E5EFFB3FE5DFCCADB4DFCA1" box="[484,560,817,839]" pageId="5" pageNumber="146" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="8BC3C5308E5EFFB3FD8FFCCADB94FCA1" blockId="5.[163,745,817,839]" box="[566,745,817,839]" pageId="5" pageNumber="146">
<heading id="D08B725C8E5EFFB3FD8FFCCADB94FCA1" box="[566,745,817,839]" fontSize="8" level="2" pageId="5" pageNumber="146" reason="2">
(
<figureCitation id="1347D9B58E5EFFB3FD84FCCADBEEFCA1" box="[573,659,817,839]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="5" pageNumber="146">Figs 1-5</figureCitation>
,
<figureCitation id="1347D9B58E5EFFB3FD26FCC9DBA2FCA1" box="[671,735,817,839]" captionStart="Map 1" captionStartId="7.[163,208,1544,1564]" captionTargetBox="[162,1070,861,1530]" captionTargetId="figure-281@7.[164,1068,1140,1529]" captionTargetPageId="7" captionText="Map 1: Distribution of Arabian Pinophilinus species: P. schatzmayri (KOCH) (triangle); P. spinosus nov.sp. (circles); P. socotranus ASSING (diamond)." figureDoi="http://doi.org/10.5281/zenodo.5341018" httpUri="https://zenodo.org/record/5341018/files/figure.png" pageId="5" pageNumber="146">Map 1</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="C36696BB8E5EFFB3FF1AFCACDAE8FC49" pageId="5" pageNumber="146" type="materials_examined">
<paragraph id="8BC3C5308E5EFFB3FF1AFCACDAE8FC49" blockId="5.[163,1070,854,943]" pageId="5" pageNumber="146">
Type material:
<materialsCitation id="3B14CF6D8E5EFFB3FE25FCACDB2EFC7F" ID-GBIF-Occurrence="3762959311" collectingDate="2011-05-11" collectingDateMax="2011-09-22" collectingDateMin="2011-05-11" collectionCode="KSMA, cAss" collectorName="Sharaf &amp; M. R. Sharaf" country="Saudi Arabia" determinerName="V. Assing" elevation="740" latitude="19.838806" location="Shada Mnt." longLatPrecision="1" longitude="41.31172" municipality="Al Mukhwah Dhi Ayn Arch. vill." pageId="5" pageNumber="146" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Al Bahah" typeStatus="holotype">
<typeStatus id="54C77B928E5EFFB3FE25FCACD88DFC8D" box="[412,496,855,875]" pageId="5" pageNumber="146" type="holotype">Holotype</typeStatus>
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: “
<collectingCountry id="F36B85A08E5EFFB3FD9AFCACDBB7FC8D" box="[547,714,855,875]" name="Saudi Arabia" pageId="5" pageNumber="146">SAUDI ARABIA</collectingCountry>
<quantity id="4C8468D58E5EFFB3FD6AFCACDA6FFC8D" box="[723,786,855,875]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="146" unit="m" value="740.0">
<elevation id="005122038E5EFFB3FD6AFCACDA6FFC8D" box="[723,786,855,875]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="146" unit="m" value="740.0">740 m</elevation>
</quantity>
,
<collectingRegion id="49B80BD28E5EFFB3FCA7FCACDA07FC8D" box="[798,890,855,875]" country="Saudi Arabia" name="Al Bahah" pageId="5" pageNumber="146">Al Bahah</collectingRegion>
,
<collectingMunicipality id="6BA75F4A8E5EFFB3FC3EFCACD847FC64" pageId="5" pageNumber="146">Al Mukhwah Dhi Ayn Arch. vill.</collectingMunicipality>
,
<geoCoordinate id="EE48A3F78E5EFFB3FEF1FC95D8C9FC64" box="[328,436,878,898]" degrees="19" direction="north" minutes="55" orientation="latitude" pageId="5" pageNumber="146" precision="15" seconds="46" value="19.929443">19°55'46''N</geoCoordinate>
,
<geoCoordinate id="EE48A3F78E5EFFB3FE7AFC95DB51FC64" box="[451,556,878,898]" degrees="41" direction="east" minutes="26" orientation="longitude" pageId="5" pageNumber="146" precision="15" seconds="30" value="41.44167">41°26'30''E</geoCoordinate>
,
<date id="FFC2E3F08E5EFFB3FD85FC95DBE0FC64" box="[572,669,878,898]" pageId="5" pageNumber="146" value="2011-05-11">
<collectingDate id="EF861A188E5EFFB3FD85FC95DBE0FC64" box="[572,669,878,898]" pageId="5" pageNumber="146" value="2011-05-11">11.V.2011</collectingDate>
</date>
, leg.
<collectorName id="2689A0E68E5EFFB3FD60FC95DA69FC64" box="[729,788,878,898]" pageId="5" pageNumber="146">Sharaf</collectorName>
/ Holotypus ♂
<originalDetermination id="A8627DCC8E5EFFB3FC03FC96D857FC7F" pageId="5" pageNumber="146">
<taxonomicName id="4C7CBEB38E5EFFB3FC03FC96D98CFC7F" authority="Assing &amp; Schülke &amp; M.R &amp; A.S, 2013" authorityName="Assing &amp; Schülke &amp; M.R &amp; A.S" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="B90819228E5EFFB3FC03FC96D98CFC7F" italics="true" pageId="5" pageNumber="146">Pinophilinus spinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23BA4598E5EFFB3FF4EFC7ED857FC7F" box="[247,298,901,921]" pageId="5" pageNumber="146" rank="species">sp. n.</taxonomicNameLabel>
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, det.
<determinerName id="7E1F29F48E5EFFB3FEE5FC7ED8CAFC7F" box="[348,439,901,921]" pageId="5" pageNumber="146">V. Assing</determinerName>
2012” (
<collectionCode id="ED6D5DF58E5EFFB3FDBBFC7EDB34FC7F" box="[514,585,901,921]" pageId="5" pageNumber="146">KSMA</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B14CF6D8E5EFFB3FDE1FC7EDAE8FC49" ID-GBIF-Occurrence="3762959342" collectingDate="2011-05-11" collectingDateMax="2011-09-22" collectingDateMin="2011-05-11" collectionCode="KSMA, cAss" collectorName="Sharaf &amp; M. R. Sharaf" country="Saudi Arabia" determinerName="V. Assing" elevation="740" latitude="19.838806" location="Shada Mnt." longLatPrecision="1" longitude="41.31172" municipality="Al Mukhwah Dhi Ayn Arch. vill." pageId="5" pageNumber="146" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Al Bahah" typeStatus="paratype">
<typeStatus id="54C77B928E5EFFB3FDE1FC7EDBD5FC7F" box="[600,680,901,921]" pageId="5" pageNumber="146" type="paratype">Paratype</typeStatus>
<specimenCount id="9D7A0EB98E5EFFB3FD16FC72DBBFFC78" box="[687,706,905,926]" pageId="5" pageNumber="146" type="female"></specimenCount>
: “
<collectingCountry id="F36B85A08E5EFFB3FD6FFC7EDA2FFC7F" box="[726,850,901,921]" pageId="5" pageNumber="146">Saudi Arabia</collectingCountry>
,
<collectingRegion id="49B80BD28E5EFFB3FCE2FC7EDAC9FC7F" box="[859,948,901,921]" country="Saudi Arabia" name="Al Bahah" pageId="5" pageNumber="146">Al Bahah</collectingRegion>
,
<location id="8EA393EB8E5EFFB3FC06FC7EDD56FC7F" LSID="urn:lsid:plazi:treatment:03D574268E5EFFBEFF1AFCCADA40FDD5:8EA393EB8E5EFFB3FC06FC7EDD56FC7F" box="[959,1067,901,921]" country="Saudi Arabia" latitude="19.838806" longLatPrecision="1" longitude="41.31172" municipality="Al Mukhwah Dhi Ayn Arch. vill." name="Shada Mnt." pageId="5" pageNumber="146" stateProvince="Al Bahah">Shada Mnt.</location>
,
<date id="FFC2E3F08E5EFFB3FF1AFC60D870FC49" box="[163,269,923,943]" pageId="5" pageNumber="146" value="2011-09-22">
<collectingDate id="EF861A188E5EFFB3FF1AFC60D870FC49" box="[163,269,923,943]" pageId="5" pageNumber="146" value="2011-09-22">22.IX.2011</collectingDate>
</date>
,
<geoCoordinate id="EE48A3F78E5EFFB3FEAFFC60D8EEFC49" box="[278,403,923,943]" degrees="19" direction="north" minutes="50" orientation="latitude" pageId="5" pageNumber="146" precision="1" seconds="19.7" value="19.838806">19°50'19.7''N</geoCoordinate>
,
<geoCoordinate id="EE48A3F78E5EFFB3FE24FC60DB6AFC49" box="[413,535,923,943]" degrees="41" direction="east" minutes="18" orientation="longitude" pageId="5" pageNumber="146" precision="1" seconds="42.2" value="41.31172">41°18'42.2''E</geoCoordinate>
,
<quantity id="4C8468D58E5EFFB3FD99FC60DB15FC49" box="[544,616,923,943]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="5" pageNumber="146" unit="m" value="1620.0">
<elevation id="005122038E5EFFB3FD99FC60DB15FC49" box="[544,616,923,943]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="5" pageNumber="146" unit="m" value="1620.0">1620 m</elevation>
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, Leg.
<collectorName id="2689A0E68E5EFFB3FD19FC60DA72FC49" box="[672,783,923,943]" pageId="5" pageNumber="146">M.R. Sharaf</collectorName>
0036” (
<collectionCode id="ED6D5DF58E5EFFB3FCE0FC60DAF7FC49" box="[857,906,923,943]" pageId="5" pageNumber="146">cAss</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36696BB8E5EFFB3FF1AFC45DB40FC17" pageId="5" pageNumber="146" type="etymology">
<paragraph id="8BC3C5308E5EFFB3FF1AFC45DB40FC17" blockId="5.[163,1070,958,1009]" pageId="5" pageNumber="146">Etymology: The specific epithet (Latin, adjective) refers to the presence of a large spine in the internal sac of the aedeagus.</paragraph>
</subSubSection>
<subSubSection id="C36696BB8E5EFFB1FF1AFBFBD80DFE9A" lastPageId="7" lastPageNumber="148" pageId="5" pageNumber="146" type="description">
<paragraph id="8BC3C5308E5EFFB3FF1AFBFBDAD5FBD2" blockId="5.[163,1070,1024,1076]" pageId="5" pageNumber="146">
Description: Body length
<quantity id="4C8468D58E5EFFB3FDB1FBFBDBFFFBF0" box="[520,642,1024,1046]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.5" metricValueMin="5.8" pageId="5" pageNumber="146" unit="mm" value="6.15" valueMax="6.5" valueMin="5.8">5.8-6.5 mm</quantity>
; length of forebody
<quantity id="4C8468D58E5EFFB3FCE0FBFBDAAEFBF0" box="[857,979,1024,1046]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="5" pageNumber="146" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2-2.4 mm</quantity>
. Habitus as in
<figureCitation id="1347D9B58E5EFFB3FF61FBE5D868FBD2" box="[216,277,1054,1076]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="5" pageNumber="146">Fig. 1</figureCitation>
. Coloration: whole body pale-reddish; legs and antennae yellow.
</paragraph>
<paragraph id="8BC3C5308E5EFFB3FF1AFBB8DBDCFAA5" blockId="5.[163,1071,1091,1347]" pageId="5" pageNumber="146">
Head (
<figureCitation id="1347D9B58E5EFFB3FF53FBB8D856FBBF" box="[234,299,1091,1113]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
) strongly transverse, approximately 1.4 times as wide as long; posterior margin concave in the middle; frons practically impunctate; punctation of vertex dense, somewhat sparser in posterior median portion, rather coarse, and defined; interstices glossy, without microsculpture, on average as broad as, or slightly narrower than diameter of punctures, somewhat broader in posterior median portion. Eyes weakly convex and rather large, distinctly longer than postocular region in dorsal view. Antenna approximately
<quantity id="4C8468D58E5EFFB3FF48FB08D83CFAEE" box="[241,321,1267,1289]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="146" unit="mm" value="1.4">1.4 mm</quantity>
long; antennomere III approximately twice as long as broad, IV-V approximately 1.5 times as long as broad, VI and VII weakly oblong, VIII approximately as long as broad, and IX and X weakly transverse.
</paragraph>
<paragraph id="8BC3C5308E5EFFB3FF1AFAA8DA4DFA26" blockId="5.[163,1070,1362,1472]" pageId="5" pageNumber="146">
Pronotum (
<figureCitation id="1347D9B58E5EFFB3FEAFFAA9D829FA8E" box="[278,340,1362,1384]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
) indistinctly oblong, at anterior angles approximately as wide as head, and distinctly tapering posteriad in posterior half; posterior angles abruptly rounded, moderately marked; punctation similar to that of head; interstices without microsculpture, on average somewhat narrower than diameter of punctures.
</paragraph>
<paragraph id="8BC3C5308E5EFFB3FF1AFA2BDD60F9C6" blockId="5.[163,1070,1488,1568]" pageId="5" pageNumber="146">
Elytra (
<figureCitation id="1347D9B58E5EFFB3FF56FA2BD856FA00" box="[239,299,1488,1510]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
) short and narrow, approximately 0.75 times as long and 0.81-0.82 times as broad as pronotum, slightly widened posteriad; humeral angles almost obsolete; punctures similar to those of pronotum, or slightly larger; interstices without microsculpture.
</paragraph>
<caption id="DF0395B88E5DFFB0FF1AFA80D8E8FA5A" ID-DOI="http://doi.org/10.5281/zenodo.5341016" ID-Zenodo-Dep="5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="6" pageNumber="147" startId="6.[163,203,1403,1423]" targetBox="[176,1060,209,1383]" targetPageId="6">
<paragraph id="8BC3C5308E5DFFB0FF1AFA80D8E8FA5A" blockId="6.[163,1070,1402,1468]" pageId="6" pageNumber="147">
<emphasis id="B90819228E5DFFB0FF1AFA80D98CFA69" bold="true" box="[163,241,1403,1423]" pageId="6" pageNumber="147">Figs 1-8</emphasis>
:
<taxonomicName id="4C7CBEB38E5DFFB0FF47FA81D8BAFA69" authority="Assing &amp; Schülke &amp; M.R &amp; A.S, 2013" authorityName="Assing &amp; Schülke &amp; M.R &amp; A.S" authorityYear="2013" box="[254,455,1402,1423]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="B90819228E5DFFB0FF47FA81D8BAFA69" box="[254,455,1402,1423]" italics="true" pageId="6" pageNumber="147">Pinophilinus spinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23BA4598E5DFFB0FE69FA80DB6EFA69" box="[464,531,1403,1423]" pageId="6" pageNumber="147" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="B90819228E5DFFB0FDA6FA80DB40FA69" bold="true" box="[543,573,1403,1423]" pageId="6" pageNumber="147">1-5</emphasis>
) and
<taxonomicName id="4C7CBEB38E5DFFB0FDCCFA81DA36FA69" authority="(KOCH)" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[629,843,1402,1423]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5DFFB0FDCCFA81DB83FA69" box="[629,766,1402,1423]" italics="true" pageId="6" pageNumber="147">P. schatzmayri</emphasis>
(KOCH)
</taxonomicName>
(
<emphasis id="B90819228E5DFFB0FCE0FA80DA05FA69" bold="true" box="[857,888,1403,1423]" pageId="6" pageNumber="147">6-8</emphasis>
): (
<emphasis id="B90819228E5DFFB0FC2DFA80DACAFA69" bold="true" box="[916,951,1403,1423]" pageId="6" pageNumber="147">1, 6</emphasis>
) habitus; (
<emphasis id="B90819228E5DFFB0FBA4FA80D9D3FA40" bold="true" pageId="6" pageNumber="147">2, 7</emphasis>
) forebody; (
<emphasis id="B90819228E5DFFB0FE9DFA69D83AFA40" bold="true" box="[292,327,1426,1446]" pageId="6" pageNumber="147">3, 8</emphasis>
) abdomen; (
<emphasis id="B90819228E5DFFB0FE07FA69D8B4FA40" bold="true" box="[446,457,1426,1446]" pageId="6" pageNumber="147">4</emphasis>
) male sternite VIII; (
<emphasis id="B90819228E5DFFB0FD28FA69DBE1FA40" bold="true" box="[657,668,1426,1446]" pageId="6" pageNumber="147">5</emphasis>
) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC3C5308E5DFFB0FF1AFA30DD64FA07" blockId="6.[163,1049,1483,1505]" box="[163,1049,1483,1505]" pageId="6" pageNumber="147">Hind wings completely reduced. Metatarsomere I as long as the combined length of II-IV.</paragraph>
<paragraph id="8BC3C5308E5DFFB1FF1AFA0BDBF4FF19" blockId="6.[163,1070,1520,1571]" lastBlockId="7.[163,1070,203,255]" lastPageId="7" lastPageNumber="148" pageId="6" pageNumber="147">
Abdomen (
<figureCitation id="1347D9B58E5DFFB0FEAFFA0BD82EF9E0" box="[278,339,1520,1542]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="6" pageNumber="147">Fig. 3</figureCitation>
) long and slender, broader than elytra; maximal width at segment VII; punctation of tergites III-V coarse and dense, that of tergites VII and VIII rather fine and much sparser, that of tergite VI intermediate; interstices without microsculpture; posterior margin of tergite VII without palisade fringe.
</paragraph>
<paragraph id="8BC3C5308E5CFFB1FF1AFEF5D80DFE9A" blockId="7.[163,1070,270,380]" pageId="7" pageNumber="148">
♂: sternite VIII oblong, distinctly tapering posteriad, and with V-shaped posterior excision (
<figureCitation id="1347D9B58E5CFFB1FF65FED0D861FEA7" box="[220,284,299,321]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="7" pageNumber="148">Fig. 4</figureCitation>
); aedeagus approximately
<quantity id="4C8468D58E5CFFB1FD96FED0DB03FEA7" box="[559,638,299,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="148" unit="mm" value="1.2">1.2 mm</quantity>
long, with slightly asymmetric and in the middle pointed ventral process (ventral view) and with large sclerotized spine in the internal sac (
<figureCitation id="1347D9B58E5CFFB1FE9CFE9DD81FFE9A" box="[293,354,358,380]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="7" pageNumber="148">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36696BB8E5CFFBEFF1AFE70D859FE9A" lastPageId="8" lastPageNumber="149" pageId="7" pageNumber="148" type="discussion">
<paragraph id="8BC3C5308E5CFFB1FF1AFE70DB1DFE1F" blockId="7.[163,1070,395,505]" pageId="7" pageNumber="148">
Comparative notes:Two
<taxonomicName id="4C7CBEB38E5CFFB1FD94FE70DBD3FE46" authority="EICHELBAUM 1908" authorityName="EICHELBAUM" authorityYear="1908" box="[557,686,395,416]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B90819228E5CFFB1FD94FE70DBD3FE46" box="[557,686,395,416]" italics="true" pageId="7" pageNumber="148">Pinophilinus</emphasis>
</taxonomicName>
species were previously known from the Arabian region,
<taxonomicName id="4C7CBEB38E5CFFB1FECCFE52DBE2FE58" authority="(KOCH 1934)" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[373,671,424,446]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5CFFB1FECCFE52DB70FE58" box="[373,525,425,446]" italics="true" pageId="7" pageNumber="148">P. schatzmayri</emphasis>
(
<bibRefCitation id="EFEDB8C18E5CFFB1FDA4FE53DBE5FE58" author="KOCH C" box="[541,664,424,446]" pageId="7" pageNumber="148" pagination="33 - 91" refId="ref7632" refString="KOCH C. (1934): Wissenschaftliche Ergebnisse der entomologischen Expeditionen seiner Durchlaucht des Fuersten Alessandro C. della Torre e Tasso nach Aegypten und auf die Halbinsel Sinai. IV. Staphylinidae (Coleoptera). - Bulletin de la Societe Royale Entomologique d'Egypte 1934: 33 - 91." type="journal article" year="1934">KOCH 1934</bibRefCitation>
)
</taxonomicName>
from the south of the Sinai Peninsula and
<taxonomicName id="4C7CBEB38E5CFFB1FF68FE3CD88CFE3A" authority="ASSING 2012" authorityName="ASSING" authorityYear="2012" box="[209,497,454,476]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="B90819228E5CFFB1FF68FE3CD822FE3A" box="[209,351,455,476]" italics="true" pageId="7" pageNumber="148">P. socotranus</emphasis>
ASSING 2012
</taxonomicName>
from
<collectingCountry id="F36B85A08E5CFFB1FD8DFE3DDBB6FE3A" box="[564,715,454,476]" name="Socotra Island" pageId="7" pageNumber="148">Socotra Island</collectingCountry>
,
<collectingCountry id="F36B85A08E5CFFB1FD60FE3DDA58FE3D" box="[729,805,454,475]" name="Yemen" pageId="7" pageNumber="148">Yemen</collectingCountry>
.
<taxonomicName id="4C7CBEB38E5CFFB1FC8BFE3DDD6EFE3D" authorityName="Assing &amp; Schülke &amp; M.R &amp; A.S" authorityYear="2013" box="[818,1043,454,475]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B90819228E5CFFB1FC8BFE3DDD6EFE3D" box="[818,1043,454,475]" italics="true" pageId="7" pageNumber="148">Pinophilinus spinosus</emphasis>
</taxonomicName>
is distinguished from these species as follows:
</paragraph>
<paragraph id="8BC3C5308E5CFFBEFF1AFDF3DBE3FEDF" blockId="7.[163,1071,520,836]" lastBlockId="8.[163,1070,203,313]" lastPageId="8" lastPageNumber="149" pageId="7" pageNumber="148">
from
<taxonomicName id="4C7CBEB38E5CFFB1FF64FDF2D80AFDFB" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[221,375,520,542]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5CFFB1FF64FDF2D80AFDFB" box="[221,375,520,542]" italics="true" pageId="7" pageNumber="148">P. schatzmayri</emphasis>
</taxonomicName>
(male unknown) by much smaller size (
<taxonomicName id="4C7CBEB38E5CFFB1FC9CFDF2DAC2FDFB" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[805,959,520,542]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5CFFB1FC9CFDF2DAC2FDFB" box="[805,959,520,542]" italics="true" pageId="7" pageNumber="148">P. schatzmayri</emphasis>
</taxonomicName>
: length of forebody
<quantity id="4C8468D58E5CFFB1FEBEFDDED82AFDDD" box="[263,343,549,571]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="148" unit="mm" value="3.4">3.4 mm</quantity>
) (compare
<figureCitation id="1347D9B58E5CFFB1FE76FDDEDB29FDDD" box="[463,596,549,571]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="7" pageNumber="148">Figs 1 and 6</figureCitation>
), much shorter antennae (
<taxonomicName id="4C7CBEB38E5CFFB1FCDCFDDDDA83FDDD" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[869,1022,550,571]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5CFFB1FCDCFDDDDA83FDDD" box="[869,1022,550,571]" italics="true" pageId="7" pageNumber="148">P. schatzmayri</emphasis>
</taxonomicName>
: antennae
<quantity id="4C8468D58E5CFFB1FF57FDB8D842FDBE" box="[238,319,579,601]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="148" unit="mm" value="2.1">2.1 mm</quantity>
long; antennomere III approximately three times as long as broad, IV-VIII distinctly oblong), larger and less convex eyes, finer setation of the head and pronotum (
<taxonomicName id="4C7CBEB38E5CFFB1FEA0FD85D8C9FD75" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[281,436,638,659]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5CFFB1FEA0FD85D8C9FD75" box="[281,436,638,659]" italics="true" pageId="7" pageNumber="148">P. schatzmayri</emphasis>
</taxonomicName>
: head and pronotum with long and dark lateral setae), the posteriorly more distinctly narrowed pronotum (
<taxonomicName id="4C7CBEB38E5CFFB1FD21FD60DA4DFD56" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[664,816,667,688]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5CFFB1FD21FD60DA4DFD56" box="[664,816,667,688]" italics="true" pageId="7" pageNumber="148">P. schatzmayri</emphasis>
</taxonomicName>
: lateral margins in anterior two thirds of pronotum almost parallel), and the less coarse punctation of the abdomen; from
<taxonomicName id="4C7CBEB38E5CFFB1FF65FD2DD814FD0D" authorityName="ASSING" authorityYear="2012" box="[220,361,726,747]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="B90819228E5CFFB1FF65FD2DD814FD0D" box="[220,361,726,747]" italics="true" pageId="7" pageNumber="148">P. socotranus</emphasis>
</taxonomicName>
by much paler coloration (
<taxonomicName id="4C7CBEB38E5CFFB1FD3EFD2DDA69FD0D" authorityName="ASSING" authorityYear="2012" box="[647,788,726,747]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="B90819228E5CFFB1FD3EFD2DDA69FD0D" box="[647,788,726,747]" italics="true" pageId="7" pageNumber="148">P. socotranus</emphasis>
</taxonomicName>
: body dark-brown), denser punctation of the whole body, distinctly longer antennae with more slender antennomeres (
<taxonomicName id="4C7CBEB38E5CFFB1FF12FCEAD844FCC0" authorityName="ASSING" authorityYear="2012" box="[171,313,785,806]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="B90819228E5CFFB1FF12FCEAD844FCC0" box="[171,313,785,806]" italics="true" pageId="7" pageNumber="148">P. socotranus</emphasis>
</taxonomicName>
: antenna approximately
<quantity id="4C8468D58E5CFFB1FD83FCEBDBF7FCC0" box="[570,650,784,806]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="148" unit="mm" value="1.2">1.2 mm</quantity>
long; antennomere III approximately 1.5 times as long as broad, IV-VII weakly oblong), much larger eyes (
<taxonomicName id="4C7CBEB38E5CFFB1FC2FFCD4DD55FCA2" authorityName="ASSING" authorityYear="2012" box="[918,1064,815,836]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="B90819228E5CFFB1FC2FFCD4DD55FCA2" box="[918,1064,815,836]" italics="true" pageId="7" pageNumber="148">P. socotranus</emphasis>
</taxonomicName>
: shorter than postocular region in dorsal view), the more convex (cross-section) and posteriorly less strongly narrowed pronotum, the longer, more slender, and less convex elytra, the larger and deeper posterior excision of the male sternite VIII, and by the completely different morphology of the aedeagus.
</paragraph>
<caption id="DF0395B88E5CFFB1FF1AF9F3DBB4F9D5" ID-DOI="http://doi.org/10.5281/zenodo.5341018" ID-Zenodo-Dep="5341018" httpUri="https://zenodo.org/record/5341018/files/figure.png" pageId="7" pageNumber="148" startId="7.[163,208,1544,1564]" targetBox="[162,1070,861,1530]" targetPageId="7">
<paragraph id="8BC3C5308E5CFFB1FF1AF9F3DBB4F9D5" blockId="7.[163,1071,1543,1587]" pageId="7" pageNumber="148">
<emphasis id="B90819228E5CFFB1FF1AF9F3D995F9FA" bold="true" box="[163,232,1544,1564]" pageId="7" pageNumber="148">Map 1</emphasis>
: Distribution of Arabian
<taxonomicName id="4C7CBEB38E5CFFB1FE54F9FCDBCEF9FA" box="[493,691,1543,1564]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90819228E5CFFB1FE54F9FCDB1CF9FA" box="[493,609,1543,1564]" italics="true" pageId="7" pageNumber="148">Pinophilinus</emphasis>
species
</taxonomicName>
:
<taxonomicName id="4C7CBEB38E5CFFB1FD7AF9F3DADBF9FA" authority="(KOCH)" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[707,934,1543,1565]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E5CFFB1FD7AF9F3DA2CF9FA" box="[707,849,1543,1565]" italics="true" pageId="7" pageNumber="148">P. schatzmayri</emphasis>
(KOCH)
</taxonomicName>
(triangle);
<taxonomicName id="4C7CBEB38E5CFFB1FBA3F9F3D98CF9D5" authority="Assing &amp; Schülke &amp; M.R &amp; A.S, 2013" authorityName="Assing &amp; Schülke &amp; M.R &amp; A.S" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="B90819228E5CFFB1FBA3F9F3D98CF9D5" italics="true" pageId="7" pageNumber="148">P. spinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23BA4598E5CFFB1FF4EF9E4D844F9D5" box="[247,313,1567,1587]" pageId="7" pageNumber="148" rank="species">nov.sp.</taxonomicNameLabel>
(circles);
<taxonomicName id="4C7CBEB38E5CFFB1FE2DF9E5DB23F9D4" authority="ASSING" authorityName="ASSING" authorityYear="2012" box="[404,606,1566,1587]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="B90819228E5CFFB1FE2DF9E5DB6FF9D5" box="[404,530,1566,1587]" italics="true" pageId="7" pageNumber="148">P. socotranus</emphasis>
ASSING
</taxonomicName>
(diamond).
</paragraph>
</caption>
<paragraph id="8BC3C5308E53FFBEFF1AFEB2D859FE9A" blockId="8.[163,1070,329,380]" pageId="8" pageNumber="149">
For illustrations of
<taxonomicName id="4C7CBEB38E53FFBEFED7FEB2DB75FEB8" baseAuthorityName="Koch" baseAuthorityYear="1934" box="[366,520,329,350]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="schatzmayri">
<emphasis id="B90819228E53FFBEFED7FEB2DB75FEB8" box="[366,520,329,350]" italics="true" pageId="8" pageNumber="149">P. schatzmayri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7CBEB38E53FFBEFD86FEB2DBB0FEB8" authorityName="ASSING" authorityYear="2012" box="[575,717,329,350]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="B90819228E53FFBEFD86FEB2DBB0FEB8" box="[575,717,329,350]" italics="true" pageId="8" pageNumber="149">P. socotranus</emphasis>
</taxonomicName>
see
<figureCitation id="1347D9B58E53FFBEFCB9FEB2DA2AFEB9" box="[768,855,329,351]" captionStart="Figs 1-8" captionStartId="6.[163,203,1403,1423]" captionTargetBox="[176,1060,209,1383]" captionTargetId="figure-0@6.[171,1060,209,1395]" captionTargetPageId="6" captionText="Figs 1-8: Pinophilinus spinosus nov.sp. (1-5) and P. schatzmayri (KOCH) (6-8): (1, 6) habitus; (2, 7) forebody; (3, 8) abdomen; (4) male sternite VIII; (5) aedeagus in ventral view. Scale bars: 1-3, 6-8: 1.0 mm; 4-5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5341016" httpUri="https://zenodo.org/record/5341016/files/figure.png" pageId="8" pageNumber="149">Figs 6-8</figureCitation>
and ASSING (2012), respectively.
</paragraph>
</subSubSection>
<subSubSection id="C36696BB8E53FFBEFF1AFE70DA40FDD5" pageId="8" pageNumber="149" type="distribution">
<paragraph id="8BC3C5308E53FFBEFF1AFE70DA40FDD5" blockId="8.[163,1070,395,563]" pageId="8" pageNumber="149">
Distribution and natural history The type specimens were collected in two localities in
<collectingRegion id="49B80BD28E53FFBEFE76FE53DB4EFE58" box="[463,563,424,446]" country="Saudi Arabia" name="Al Bahah" pageId="8" pageNumber="149">Al Bahah</collectingRegion>
, southwestern
<collectingCountry id="F36B85A08E53FFBEFD74FE53DA29FE58" box="[717,852,424,446]" name="Saudi Arabia" pageId="8" pageNumber="149">Saudi Arabia</collectingCountry>
(
<figureCitation id="1347D9B58E53FFBEFCDBFE53DAD8FE58" box="[866,933,424,446]" captionStart="Map 1" captionStartId="7.[163,208,1544,1564]" captionTargetBox="[162,1070,861,1530]" captionTargetId="figure-281@7.[164,1068,1140,1529]" captionTargetPageId="7" captionText="Map 1: Distribution of Arabian Pinophilinus species: P. schatzmayri (KOCH) (triangle); P. spinosus nov.sp. (circles); P. socotranus ASSING (diamond)." figureDoi="http://doi.org/10.5281/zenodo.5341018" httpUri="https://zenodo.org/record/5341018/files/figure.png" pageId="8" pageNumber="149">Map 1</figureCitation>
), at altitudes of 740 and
<quantity id="4C8468D58E53FFBEFEAEFE3DD819FE3D" box="[279,356,454,476]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="8" pageNumber="149" unit="m" value="1620.0">1620 m</quantity>
in May and September. The reduced hind wings, the short elytra, and the absence of a palisade fringe at the posterior margin of tergite VII suggest that the distribution of
<taxonomicName id="4C7CBEB38E53FFBEFE84FDFAD8CCFDF0" authorityName="Assing &amp; Schülke &amp; M.R &amp; A.S" authorityYear="2013" box="[317,433,513,534]" class="Insecta" family="Staphylinidae" genus="Pinophilinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B90819228E53FFBEFE84FDFAD8CCFDF0" box="[317,433,513,534]" italics="true" pageId="8" pageNumber="149">P. spinosus</emphasis>
</taxonomicName>
is restricted. The
<typeStatus id="54C77B928E53FFBEFDD2FDFBDBB9FDF0" box="[619,708,512,534]" pageId="8" pageNumber="149" type="holotype">holotype</typeStatus>
was collected from leaf litter unter an old
<emphasis id="B90819228E53FFBEFF51FDE5D85DFDD5" box="[232,288,542,563]" italics="true" pageId="8" pageNumber="149">Ficus</emphasis>
tree, the
<typeStatus id="54C77B928E53FFBEFEC7FDE5D8A8FDD5" box="[382,469,542,563]" pageId="8" pageNumber="149" type="paratype">paratype</typeStatus>
from soil under a large plastic bag.
</paragraph>
</subSubSection>
</treatment>
</document>