172 lines
17 KiB
XML
172 lines
17 KiB
XML
<document id="83BC0DB7E2586FB6B642B324951A8527" ID-DOI="10.5281/zenodo.273880" ID-GBIF-Dataset="7322dbf9-e317-40ae-9876-3528e7107e1f" ID-ISSN="1175-5326" ID-Zenodo-Dep="273880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459878272435" checkinUser="plazi" docAuthor="Várkonyi, Gergely & Polaszek, Andrew" docDate="2007" docId="03D0EC22E71EFF92FF74D393FBC002C8" docLanguage="en" docName="zt01546p014.pdf" docOrigin="Zootaxa 1546" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Foenobethylus emiliacasellae Varkonyi& Polaszek, sp. nov." docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="FFE9945AE716FF9BFFE3D20BFFD30162" masterDocTitle="Rediscovery and revision of Foenobethylus Kieffer, 1913 (Hymenoptera, Bethylidae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="9" updateTime="1698227249447" updateUser="plazi">
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<mods:title id="F16185612D3036072E94FEB6C530F4BA">Rediscovery and revision of Foenobethylus Kieffer, 1913 (Hymenoptera, Bethylidae)</mods:title>
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<mods:name id="32CFC61E753C8263CD0979CB6EB9D3EB" type="personal">
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<mods:namePart id="C1CC0A71E062722852C7757B80B2DF01">Várkonyi, Gergely</mods:namePart>
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<mods:namePart id="B859A63A50111FC938DABE252BE36E4E">Polaszek, Andrew</mods:namePart>
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<mods:typeOfResource id="71D2F1E9CDD10CCB8A5945F55C628038">text</mods:typeOfResource>
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<mods:title id="2CBC047A379B010567E7E7835A05637E">Zootaxa</mods:title>
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<mods:part id="2F83BA718A9DF8AE03578A84CCF7A183">
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<mods:date id="8825635193FF8CFEF030B5D501696593">2007</mods:date>
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<mods:detail id="7C3044D0DD197407E793F5232F5DEB74" type="volume">
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<mods:number id="EC68B59E84E60450028C318F034BE53F">1546</mods:number>
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<mods:classification id="ECE5FB15034C461B04E10CE3B463B9CF">journal article</mods:classification>
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<mods:identifier id="33707217975D77FF8349EA33F4E59C59" type="DOI">10.5281/zenodo.273880</mods:identifier>
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<mods:identifier id="FC361966101F82BDB4370FD6AD4AE9D1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="332F24A2C3858B740778BB284AFF89C2" type="Zenodo-Dep">273880</mods:identifier>
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<treatment id="03D0EC22E71EFF92FF74D393FBC002C8" ID-DOI="http://doi.org/10.5281/zenodo.6237226" ID-GBIF-Taxon="119370406" ID-Zenodo-Dep="6237226" LSID="urn:lsid:plazi:treatment:03D0EC22E71EFF92FF74D393FBC002C8" httpUri="http://treatment.plazi.org/id/03D0EC22E71EFF92FF74D393FBC002C8" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<subSubSection id="C3630EBFE71EFF93FF74D393FE2200B6" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BC65D34E71EFF93FF74D393FCBE00D0" blockId="8.[151,877,408,468]" box="[151,877,408,434]" pageId="8" pageNumber="9">
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<heading id="D08EEA58E71EFF93FF74D393FCBE00D0" bold="true" box="[151,877,408,434]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="B90D8126E71EFF93FF74D393FCBE00D0" bold="true" box="[151,877,408,434]" pageId="8" pageNumber="9">
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<taxonomicName id="4C7926B7E71EFF93FF74D393FCD800D0" authority="Varkonyi& Polaszek" authorityName="Varkonyi& Polaszek" box="[151,779,408,434]" class="Insecta" family="Bethylidae" genus="Foenobethylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="emiliacasellae" status="sp. nov.">
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<emphasis id="B90D8126E71EFF93FF74D393FE2F00D0" bold="true" box="[151,508,408,434]" italics="true" pageId="8" pageNumber="9">Foenobethylus emiliacasellae</emphasis>
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Várkonyi& Polaszek
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</taxonomicName>
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<taxonomicNameLabel id="A23E3C5DE71EFF93FCECD393FCBE00D0" box="[783,877,408,434]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC65D34E71EFF93FF74D3B1FE2200B6" blockId="8.[151,877,408,468]" box="[151,497,442,468]" pageId="8" pageNumber="9">
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(
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<figureCitation id="134241B1E71EFF93FF7CD3B1FEC300B6" box="[159,272,442,468]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,255,1891,1915]" captionStartId-1="5.[151,255,1502,1526]" captionTargetBox-1="[409,1177,906,1482]" captionTargetId-1="figure@5.[409,1177,906,1482]" captionTargetPageId-1="5" captionText-0="FIGURE 2. Fore and hind wings of Foenobethylus emiliacasellae n. sp." captionText-1="FIGURE 3. Head and mesosoma of Foenobethylus emiliacasellae n. sp., showing the greatly swollen fore femur." httpUri-1="https://zenodo.org/record/273882/files/figure.png" pageId="8" pageNumber="9">
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Figs.
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<date id="FFC77BF4E71EFF93FF01D3B1FEC300B6" box="[226,272,442,468]" pageId="8" pageNumber="9" value="1928-02">2–3</date>
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</figureCitation>
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,
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<date id="FFC77BF4E71EFF93FEFFD3B1FE3A00B6" box="[284,489,442,468]" pageId="8" pageNumber="9" value="1928-12-15" valueMax="1928-12-26" valueMin="1928-12-15">12–15 and 26–28</date>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3630EBFE71EFF93FF74D3F5FBA10308" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BC65D34E71EFF93FF74D3F5FBA10308" blockId="8.[151,1436,509,658]" pageId="8" pageNumber="9">
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<emphasis id="B90D8126E71EFF93FF74D3F5FEAE0375" box="[151,381,510,535]" italics="true" pageId="8" pageNumber="9">Material examined.</emphasis>
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—
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<typeStatus id="54C2E396E71EFF93FE43D3F6FDE90375" box="[416,570,509,535]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
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ɗ:
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<collectingCountry id="F36E1DA4E71EFF93FD87D3F6FD2C0375" box="[612,767,509,535]" name="Thailand" pageId="8" pageNumber="9">THAILAND</collectingCountry>
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, Chiang Mai,
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<date id="FFC77BF4E71EFF93FC47D3F6FC3B0375" box="[932,1000,509,535]" pageId="8" pageNumber="9" value="1999-12">xii.99</date>
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–
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<date id="FFC77BF4E71EFF93FC1AD3F6FBFD0375" box="[1017,1070,509,535]" pageId="8" pageNumber="9" value="2000-02">ii.00</date>
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, R.A. Beaver, Malaise; Genitalia, subgenital plate and 8th sternum mounted separately in
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<collectingCountry id="F36E1DA4E71EFF93FC45D023FC2D0320" box="[934,1022,552,578]" name="Canada" pageId="8" pageNumber="9">Canada</collectingCountry>
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balsam on a microscope slide.
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<typeStatus id="54C2E396E71EFF93FF74D05BFEEA0308" box="[151,313,592,618]" pageId="8" pageNumber="9" type="paratype">PARATYPES</typeStatus>
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(6 ɗɗ):
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<collectingCountry id="F36E1DA4E71EFF93FE43D05BFDE90308" box="[416,570,592,618]" name="Thailand" pageId="8" pageNumber="9">THAILAND</collectingCountry>
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, Chiang Mai,
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<date id="FFC77BF4E71EFF93FD03D05BFCF70308" box="[736,804,592,618]" pageId="8" pageNumber="9" value="1999-12">xii.99</date>
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–
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<date id="FFC77BF4E71EFF93FCD6D05BFCB90308" box="[821,874,592,618]" pageId="8" pageNumber="9" value="2000-02">ii.00</date>
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, R.A. Beaver, Malaise
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</paragraph>
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</subSubSection>
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<subSubSection id="C3630EBFE71EFF92FF25D072FE120320" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BC65D34E71EFF93FF25D072FCBD03F0" blockId="8.[151,1436,509,658]" box="[198,878,632,658]" pageId="8" pageNumber="9">
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<emphasis id="B90D8126E71EFF93FF25D072FDFF03F0" box="[198,556,633,658]" italics="true" pageId="8" pageNumber="9">
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Description of Male
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<typeStatus id="54C2E396E71EFF93FE59D072FDFA03F0" box="[442,553,633,658]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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—Length of body
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<quantity id="4C81F0D1E71EFF93FCEED073FCB903F0" box="[781,874,632,658]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="9" unit="mm" value="3.1">3.1 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BC65D34E71EFF93FF74D7D8FC300769" blockId="8.[151,1436,1489,1547]" pageId="8" pageNumber="9">
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<emphasis id="B90D8126E71EFF93FF74D7D8FE9C0489" bold="true" box="[151,335,1489,1515]" pageId="8" pageNumber="9">FIGURE 12–15.</emphasis>
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<taxonomicName id="4C7926B7E71EFF93FEB5D7D8FD400488" box="[342,659,1491,1514]" class="Insecta" family="Bethylidae" genus="Foenobethylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="emiliacasellae" status="sp. nov.">
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<emphasis id="B90D8126E71EFF93FEB5D7D8FD400488" box="[342,659,1491,1514]" italics="true" pageId="8" pageNumber="9">Foenobethylus emiliacasellae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23E3C5DE71EFF93FD79D7D8FD060489" box="[666,725,1491,1515]" pageId="8" pageNumber="9" rank="species">
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<emphasis id="B90D8126E71EFF93FD79D7D8FD060489" bold="true" box="[666,725,1491,1515]" pageId="8" pageNumber="9">n. sp.</emphasis>
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</taxonomicNameLabel>
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: 12) dorsal aspect of head, 13) dorsal aspect of mesosoma, 14) lateral aspect of mesosoma, 15) ventrolateral aspect of hind trochanter and femur.
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</paragraph>
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<paragraph id="8BC65D34E71EFF93FF25D44FFEB20644" blockId="8.[151,1437,1604,2033]" pageId="8" pageNumber="9">
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<emphasis id="B90D8126E71EFF93FF25D44FFEF4073F" box="[198,295,1604,1629]" italics="true" pageId="8" pageNumber="9">Head.—</emphasis>
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Up to 1.36 times as long as wide, elongated in dorsal aspect, fairly strongly narrowing behind and evenly rounded towards occipital carina. The distance from the eyes to the occipital carina ca. 1.1 times the length of the eyes in dorsal aspect. Ocellar triangle forms a right angle at the anterior ocellus. Lateral ocelli are distributed closer to anterior ocellus than the maximum diameter of the latter. Occipital carina from hind ocelli at a distance of ca. 1.5 times of the maximum diameter of the latter. Surface of head finely coriaceous with sparse punctures.
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</paragraph>
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<paragraph id="8BC65D34E71EFF93FF25D53CFBE2061B" blockId="8.[151,1437,1604,2033]" pageId="8" pageNumber="9">Length-to-width ratios of antennomeres: scape: 3.2, pedicel: 1.5, 1st flagellomere: 1.7, 5th flagellomere: 1.6. Terminal palpomere of maxillary palps less than 3 times longer than wide.</paragraph>
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<paragraph id="8BC65D34E71EFF93FF25D58CFC460693" blockId="8.[151,1437,1604,2033]" pageId="8" pageNumber="9">
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<emphasis id="B90D8126E71EFF93FF25D58CFEB606C2" box="[198,357,1927,1952]" italics="true" pageId="8" pageNumber="9">Mesosoma.—</emphasis>
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With a length-to-depth ratio of 3.6 and length-to-width ratio of 2.4. Pronotum elongate and narrow anteriorly, without a horizontal shelf protruding above base of propleuron – the shelf is normally developed laterally. Length-to-width ratio of propodeal disc 1.15.
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</paragraph>
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<paragraph id="8BC65D34E71FFF92FF25D293FAD70030" blockId="9.[151,1437,152,938]" pageId="9" pageNumber="10">
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Radius with or without a node in its basal 1/7. Distal end of pterostigma continuing as an extremely short metacarpal vein. Fore femur greatly swollen, with a length-to-width ratio of ca. 2.0, and with a broad and oblique inner concavity, which frequently causes rolling up the fore femur as a spiral.
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<collectingCountry id="F36E1DA4E71FFF92FB72D2E3FB1F0060" box="[1169,1228,232,258]" name="India" pageId="9" pageNumber="10">Hind</collectingCountry>
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trochanter with a long and needle-like ventral spine.
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<collectingCountry id="F36E1DA4E71FFF92FDD7D31BFDBC0048" box="[564,623,272,298]" name="India" pageId="9" pageNumber="10">Hind</collectingCountry>
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femur with an oblique furrow beneath, with a long basal spine on its inner edge and with a small, broad but sharp-edged median tooth on the outer lateroventral edge.
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</paragraph>
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<paragraph id="8BC65D34E71FFF92FF25D36BFB4D00C0" blockId="9.[151,1437,152,938]" pageId="9" pageNumber="10">
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<emphasis id="B90D8126E71FFF92FF25D36BFEB1001B" box="[198,354,352,377]" italics="true" pageId="9" pageNumber="10">Metasoma.—</emphasis>
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Eighth sternite with distal margin broadly and fairly shallowly emarginate, with sparse setae in its distal half. Subgenital plate with distal margin deeply emarginate, centrally setose.
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</paragraph>
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<paragraph id="8BC65D34E71FFF92FF25D3BBFD6A0090" blockId="9.[151,1437,152,938]" pageId="9" pageNumber="10">
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<emphasis id="B90D8126E71FFF92FF25D3BBFEA200AB" box="[198,369,432,457]" italics="true" pageId="9" pageNumber="10">Male genitalia</emphasis>
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.—Rather short and broad, distal part of parameres with relatively short setae. Volsella with digitus clearly visible. Aedeagus as illustrated.
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</paragraph>
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<paragraph id="8BC65D34E71FFF92FF25D00BFE120320" blockId="9.[151,1437,152,938]" pageId="9" pageNumber="10">
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<emphasis id="B90D8126E71FFF92FF25D00BFEE4037B" box="[198,311,512,537]" italics="true" pageId="9" pageNumber="10">Colour.—</emphasis>
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Brown. Basal four antennomeres, palps, mandibles, tarsi, fore tibiae, mid and hind tibiae subbasally and apically yellow.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3630EBFE71FFF92FF25D05BFF1102E0" pageId="9" pageNumber="10" type="discussion">
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<paragraph id="8BC65D34E71FFF92FF25D05BFF1102E0" blockId="9.[151,1437,152,938]" pageId="9" pageNumber="10">
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<emphasis id="B90D8126E71FFF92FF25D05BFE9D030B" box="[198,334,592,617]" italics="true" pageId="9" pageNumber="10">Remarks.—</emphasis>
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A ribosomal DNA sequence of just over 700 bases was obtained from the 28S D2 region of this species by Peter Mayhew and colleagues. This sequence is presented in Appendix 1. Using the “blast” option in the NCBI Genbank database revealed the most similar sequence in that database to be that of a
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<taxonomicName id="4C7926B7E71FFF92FA82D0ABFF310383" class="Insecta" genus="Pristocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="B90D8126E71FFF92FA82D0ABFF310383" italics="true" pageId="9" pageNumber="10">Pristocera</emphasis>
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</taxonomicName>
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species (
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<taxonomicName id="4C7926B7E71FFF92FEB3D0C3FE060380" box="[336,469,712,738]" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Bethylidae</taxonomicName>
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:
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<taxonomicName id="4C7926B7E71FFF92FE01D0C3FD530380" box="[482,640,712,738]" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Pristocerinae">Pristocerinae</taxonomicName>
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: 86% similarity), followed by that of
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<taxonomicName id="4C7926B7E71FFF92FBA1D0C3FA4F0383" box="[1090,1436,712,737]" class="Insecta" family="Bethylidae" genus="Cephalonomia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="stephanoderis">
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<emphasis id="B90D8126E71FFF92FBA1D0C3FA4F0383" box="[1090,1436,712,737]" italics="true" pageId="9" pageNumber="10">Cephalonomia stephanoderis</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="4C7926B7E71FFF92FF7CD0FBFEF70268" box="[159,292,752,778]" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Bethylidae</taxonomicName>
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:
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<taxonomicName id="4C7926B7E71FFF92FED4D0FBFE760268" box="[311,421,752,778]" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Epyrinae">Epyrinae</taxonomicName>
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: 84% similarity). The Genbank accession numbers for these two sequences are EF013049.1 and AJ302937.1 respectively. These are the only bethylid sequences in Genbank, and while being by no means conclusive, this result provides some supporting evidence for
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<taxonomicName id="4C7926B7E71FFF92FBF4D14BFB17023B" box="[1047,1220,832,857]" class="Insecta" family="Bethylidae" genus="Foenobethylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="B90D8126E71FFF92FBF4D14BFB17023B" box="[1047,1220,832,857]" italics="true" pageId="9" pageNumber="10">Foenobethylus</emphasis>
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</taxonomicName>
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being a pristocerine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3630EBFE71FFF92FF25D19BFBC002C8" box="[198,1043,912,938]" pageId="9" pageNumber="10" type="etymology">
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<paragraph id="8BC65D34E71FFF92FF25D19BFBC002C8" blockId="9.[151,1437,152,938]" box="[198,1043,912,938]" pageId="9" pageNumber="10">
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<emphasis id="B90D8126E71FFF92FF25D19BFEB802CB" box="[198,363,912,937]" italics="true" pageId="9" pageNumber="10">Etymology. —</emphasis>
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Named after Emilia Casella, of Hampstead, London,
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<collectingCountry id="F36E1DA4E71FFF92FC39D19BFBC002C8" box="[986,1043,912,938]" name="United Kingdom" pageId="9" pageNumber="10">U.K.</collectingCountry>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |