treatments-xml/data/03/F5/94/03F5945FFF9D1E76FCB9FAD1FBB2C2E5.xml
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<document id="AB112C78541CDD5FBD34E49D10406351" ID-DOI="10.1111/j.1096-3642.2008.00466.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445358" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630458879514" checkinUser="felipe" docAuthor="Biju, S. D. &amp; Bossuyt, Franky" docDate="2009" docId="03F5945FFF9D1E76FCB9FAD1FBB2C2E5" docLanguage="en" docName="j.1096-3642.2008.00466.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00466.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Philautus ochlandrae Gururaja 2007" docType="treatment" docVersion="3" lastPageNumber="429" masterDocId="FFCCEC27FFAB1E41FFE9FFFDFF8CC65A" masterDocTitle="Systematics and phylogeny of Philautus Gistel, 1848 (Anura, Rhacophoridae) in the Western Ghats of India, with descriptions of 12 new species" masterLastPageNumber="444" masterPageNumber="374" pageNumber="428" updateTime="1699278541911" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="EF31DBA28F3EC927F63A783157E63583">Systematics and phylogeny of Philautus Gistel, 1848 (Anura, Rhacophoridae) in the Western Ghats of India, with descriptions of 12 new species</mods:title>
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<mods:namePart id="96F990538D4BF407965C3414639EA965">Biju, S. D.</mods:namePart>
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<mods:namePart id="F5DA2C2BE998FB5C92CCA4BAA9F5A490">Bossuyt, Franky</mods:namePart>
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<mods:title id="1F233A19C2D0EEA319B217E3ECDBC2E6">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="8444FA40D9D17B743616929376E8C656">2009</mods:date>
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<subSubSection id="C34676C2FF9D1E77FCB9FAD1FA07C31B" box="[848,1419,1324,1345]" pageId="54" pageNumber="428" type="nomenclature">
<paragraph id="8BE32549FF9D1E77FCB9FAD1FA07C31B" blockId="54.[848,1419,1324,1345]" box="[848,1419,1324,1345]" pageId="54" pageNumber="428">
<heading id="D0AB9225FF9D1E77FCB9FAD1FA07C31B" box="[848,1419,1324,1345]" centered="true" fontSize="9" level="2" pageId="54" pageNumber="428" reason="2">
<taxonomicName id="4C5C5ECAFF9D1E77FCB9FAD1FA07C31B" ID-CoL="4G5JN" authority="GURURAJA ET AL., 2007" authorityName="Gururaja" authorityYear="2007" box="[848,1419,1324,1345]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="375" phylum="Chordata" rank="species" species="ochlandrae">
<emphasis id="B928F95BFF9D1E77FCB9FAD1FBE3C31A" box="[848,1135,1324,1345]" italics="true" pageId="54" pageNumber="428">PHILAUTUS OCHLANDRAE</emphasis>
GURURAJA
<emphasis id="B928F95BFF9D1E77FB14FACDFAB7C31A" box="[1277,1339,1328,1344]" italics="true" pageId="54" pageNumber="428">ET AL</emphasis>
., 2007
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34676C2FF9D1E76FC2EFAAFFBB2C2E5" lastPageId="55" lastPageNumber="429" pageId="54" pageNumber="428" type="description">
<paragraph id="8BE32549FF9D1E77FC2EFAAFFA98C332" blockId="54.[967,1300,1362,1384]" box="[967,1300,1362,1384]" pageId="54" pageNumber="428">
(
<figureCitation id="136739CCFF9D1E77FC26FAAFFBD3C332" box="[975,1119,1362,1384]" captionStart="Figure 52" captionStartId="55.[144,223,1068,1087]" captionTargetBox="[148,756,195,1038]" captionTargetId="figure-553@55.[148,756,195,1038]" captionTargetPageId="55" captionText="Figure 52. Philautus ochlandrae: A, specimen (SDB 6076) from Kakkayam Reserve Forest; B, specimen (BNHS 4559) from Kakkayam Reserve Forest." figureDoi="http://doi.org/10.5281/zenodo.5445466" httpUri="https://zenodo.org/record/5445466/files/figure.png" pageId="54" pageNumber="428">FIGS 52A, B</figureCitation>
,
<figureCitation id="136739CCFF9D1E77FB87FAAFFB7BC33D" box="[1134,1271,1362,1384]" captionStart="Figure 53" captionStartId="56.[164,243,1842,1861]" captionTargetBox="[184,1423,197,1813]" captionTargetId="figure-0@56.[184,1425,194,1813]" captionTargetPageId="56" captionText="Figure 53. Distribution map. A, Philautus ochlandrae; B, Philautus ponmudi; C, Philautus signatus; D, Philautus sushili sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5445468" httpUri="https://zenodo.org/record/5445468/files/figure.png" pageId="54" pageNumber="428">53A; TABLE</figureCitation>
2)
</paragraph>
<paragraph id="8BE32549FF9D1E76FCD3FA87FC53C49F" blockId="54.[826,1441,1401,1484]" lastBlockId="55.[806,1421,198,709]" lastPageId="55" lastPageNumber="429" pageId="54" pageNumber="428">
<emphasis id="B928F95BFF9D1E77FCD3FA87FC65C3D4" box="[826,1001,1401,1423]" italics="true" pageId="54" pageNumber="428">Type material:</emphasis>
<typeStatus id="54E79BEBFF9D1E77FC14FA84FBEAC3D5" box="[1021,1126,1401,1423]" pageId="54" pageNumber="428" type="holotype">Holotype</typeStatus>
, ZSI/WGFRS/V/A/632, an adult male, SVL
<quantity id="4CA488ACFF9D1E77FBECFA6AFBE6C3F7" box="[1029,1130,1431,1453]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="54" pageNumber="428" unit="mm" value="25.6">25.6 mm</quantity>
, collected from Kakkayam Reserve Forest,
<collectingCountry id="F34B65D9FF9D1E77FBFAFA4BFBEAC396" box="[1043,1126,1462,1484]" name="India" pageId="54" pageNumber="428">Calicut</collectingCountry>
District,
<collectingRegion id="4998EBABFF9D1E77FB1BFA4BFACEC396" box="[1266,1346,1462,1484]" country="India" name="Kerala" pageId="54" pageNumber="428">Kerala</collectingRegion>
,
<collectingCountry id="F34B65D9FF9D1E77FAB4FA4BFA10C396" box="[1373,1436,1462,1484]" name="India" pageId="54" pageNumber="428">India</collectingCountry>
; (
<figureCitation id="136739CCFF9C1E76FCC7FF3BFC30C681" box="[814,956,198,220]" captionStart="Figure 52" captionStartId="55.[144,223,1068,1087]" captionTargetBox="[148,756,195,1038]" captionTargetId="figure-553@55.[148,756,195,1038]" captionTargetPageId="55" captionText="Figure 52. Philautus ochlandrae: A, specimen (SDB 6076) from Kakkayam Reserve Forest; B, specimen (BNHS 4559) from Kakkayam Reserve Forest." figureDoi="http://doi.org/10.5281/zenodo.5445466" httpUri="https://zenodo.org/record/5445466/files/figure.png" pageId="55" pageNumber="429">Figs 52A, B</figureCitation>
,
<figureCitation id="136739CCFF9C1E76FC26FF3BFC73C686" box="[975,1023,198,220]" captionStart="Figure 53" captionStartId="56.[164,243,1842,1861]" captionTargetBox="[184,1423,197,1813]" captionTargetId="figure-0@56.[184,1425,194,1813]" captionTargetPageId="56" captionText="Figure 53. Distribution map. A, Philautus ochlandrae; B, Philautus ponmudi; C, Philautus signatus; D, Philautus sushili sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5445468" httpUri="https://zenodo.org/record/5445468/files/figure.png" pageId="55" pageNumber="429">53A</figureCitation>
). This combination of characters can also be observed in variations of
<taxonomicName id="4C5C5ECAFF9C1E76FB30FF19FC03C742" authority="Biju &amp; Bossuyt, 2009" authorityName="Biju &amp; Bossuyt" authorityYear="2009" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="akroparallagi" status="sp. nov.">
<emphasis id="B928F95BFF9C1E76FB30FF19FC03C742" italics="true" pageId="55" pageNumber="429">Philautus akroparallagi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B4420FF9C1E76FC7EFEFEFC61C742" box="[919,1005,259,280]" pageId="55" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="136739CCFF9C1E76FC15FEFEFBDCC743" box="[1020,1104,259,281]" captionStart="Figure 4" captionStartId="7.[144,223,1475,1494]" captionTargetBox="[303,1263,835,1444]" captionTargetId="figure-208@7.[303,1263,835,1444]" captionTargetPageId="7" captionText="Figure 4. Philautus akroparallagi sp. nov. A, specimen SDB 3021; B, paratype, (BNHS 4388); C, paratype, (BNHS 4536); D, paratype, (BNHS 4392). All photographs are from the type locality, Ponmudi, and were collected from within a range of about 100 m2." figureDoi="http://doi.org/10.5281/zenodo.5445366" httpUri="https://zenodo.org/record/5445366/files/figure.png" pageId="55" pageNumber="429">Fig. 4D</figureCitation>
) and
<taxonomicName id="4C5C5ECAFF9C1E76FB7BFEF9FA82C743" baseAuthorityName="BOULENGER" baseAuthorityYear="1882" box="[1170,1294,260,281]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="signatus">
<emphasis id="B928F95BFF9C1E76FB7BFEF9FA82C743" box="[1170,1294,260,281]" italics="true" pageId="55" pageNumber="429">P. signatus</emphasis>
</taxonomicName>
(
<figureCitation id="136739CCFF9C1E76FAF7FEFEFA0DC743" box="[1310,1409,259,281]" captionStart="Figure 55" captionStartId="58.[164,243,1582,1601]" captionTargetBox="[168,777,193,1552]" captionTargetId="figure-427@58.[168,777,193,1552]" captionTargetPageId="58" captionText="Figure 55. Philautus signatus: A, specimen (SDB 098) from Avalanche; B, specimen (BNHS 4492) from Avalanche; C, specimen (BNHS 4558) from Udhagamandalam." figureDoi="http://doi.org/10.5281/zenodo.5445472" httpUri="https://zenodo.org/record/5445472/files/figure.png" pageId="55" pageNumber="429">Fig. 55B</figureCitation>
). However,
<taxonomicName id="4C5C5ECAFF9C1E76FC73FEDFFBBBC76D" box="[922,1079,290,311]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="ochlandrae">
<emphasis id="B928F95BFF9C1E76FC73FEDFFBBBC76D" box="[922,1079,290,311]" italics="true" pageId="55" pageNumber="429">P. ochlandrae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5C5ECAFF9C1E76FB3AFEDFFA01C76D" authority="Biju &amp; Bossuyt, 2009" authorityName="Biju &amp; Bossuyt" authorityYear="2009" box="[1235,1421,290,311]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="akroparallagi" status="sp. nov.">
<emphasis id="B928F95BFF9C1E76FB3AFEDFFA01C76D" box="[1235,1421,290,311]" italics="true" pageId="55" pageNumber="429">P. akroparallagi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B4420FF9C1E76FCCFFEBCFC06C70C" box="[806,906,321,342]" pageId="55" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
by its medium male adult size, SVL 23.9 ±
<quantity id="4CA488ACFF9C1E76FC9AFEA2FC46C72E" box="[883,970,351,372]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="55" pageNumber="429" unit="mm" value="1.3">1.3 mm</quantity>
,
<emphasis id="B928F95BFF9C1E76FC31FEA2FC60C72E" box="[984,1004,351,372]" italics="true" pageId="55" pageNumber="429">N</emphasis>
= 5 (vs. small male adult size, SVL 20.7 ± 1.0 mm,
<emphasis id="B928F95BFF9C1E76FC33FE83FC62C7C9" box="[986,1006,382,403]" italics="true" pageId="55" pageNumber="429">N</emphasis>
= 16), rounded snout (vs. pointed), head wider than long, HW 8.3 ±
<quantity id="4CA488ACFF9C1E76FB1AFE61FAC6C7EB" box="[1267,1354,412,433]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="55" pageNumber="429" unit="mm" value="0.3">0.3 mm</quantity>
, HL 6.6 ±
<quantity id="4CA488ACFF9C1E76FC8CFE46FC30C78A" box="[869,956,443,465]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="429" unit="mm" value="0.6">0.6 mm</quantity>
,
<emphasis id="B928F95BFF9C1E76FC22FE46FC53C78A" box="[971,991,443,464]" italics="true" pageId="55" pageNumber="429">N</emphasis>
= 5 (vs. head length equal to width, HW 7.8 ±
<quantity id="4CA488ACFF9C1E76FC4DFE24FC77C7B5" box="[932,1019,473,495]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="55" pageNumber="429" unit="mm" value="0.4">0.4 mm</quantity>
, HL 7.8 ±
<quantity id="4CA488ACFF9C1E76FB61FE24FB53C7B5" box="[1160,1247,473,495]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="55" pageNumber="429" unit="mm" value="0.5">0.5 mm</quantity>
,
<emphasis id="B928F95BFF9C1E76FB1FFE27FA86C7B5" box="[1270,1290,474,495]" italics="true" pageId="55" pageNumber="429">N</emphasis>
= 10), and from
<taxonomicName id="4C5C5ECAFF9C1E76FC8EFE05FC6BC457" baseAuthorityName="BOULENGER" baseAuthorityYear="1882" box="[871,999,504,525]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="signatus">
<emphasis id="B928F95BFF9C1E76FC8EFE05FC6BC457" box="[871,999,504,525]" italics="true" pageId="55" pageNumber="429">P. signatus</emphasis>
</taxonomicName>
by its rounded snout (vs. pointed), canthus rostralis rounded (vs. acute), head wider than long, HW 8.3 ±
<quantity id="4CA488ACFF9C1E76FBADFDC8FB17C410" box="[1092,1179,565,586]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="55" pageNumber="429" unit="mm" value="0.3">0.3 mm</quantity>
, HL 6.6 ±
<quantity id="4CA488ACFF9C1E76FAD8FDC8FA04C410" box="[1329,1416,565,587]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="429" unit="mm" value="0.6">0.6 mm</quantity>
,
<emphasis id="B928F95BFF9C1E76FCCFFDA9FCB6C433" box="[806,826,596,617]" italics="true" pageId="55" pageNumber="429">N</emphasis>
= 5 (vs. head length subequal to width, HW 10.8 ±
<quantity id="4CA488ACFF9C1E76FC9AFD8FFC46C4D2" box="[883,970,626,648]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="55" pageNumber="429" unit="mm" value="1.1">1.1 mm</quantity>
, HL 10.6 ±
<quantity id="4CA488ACFF9C1E76FBA4FD8FFB28C4D2" box="[1101,1188,626,648]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="55" pageNumber="429" unit="mm" value="1.1">1.1 mm</quantity>
,
<emphasis id="B928F95BFF9C1E76FB46FD8EFB4FC4D2" box="[1199,1219,627,648]" italics="true" pageId="55" pageNumber="429">N</emphasis>
= 8), and absence of a lingual papilla on the tongue (vs. tongue with a lingual papilla).
</paragraph>
<caption id="DF2375C1FF9D1E77FF4DF8A7FAF2C134" ID-DOI="http://doi.org/10.5281/zenodo.5445464" ID-Zenodo-Dep="5445464" box="[164,1406,1882,1902]" httpUri="https://zenodo.org/record/5445464/files/figure.png" pageId="54" pageNumber="428" startId="54.[164,243,1882,1901]" targetBox="[323,1284,1537,1853]" targetPageId="54">
<paragraph id="8BE32549FF9D1E77FF4DF8A7FAF2C134" blockId="54.[164,1406,1882,1902]" box="[164,1406,1882,1902]" pageId="54" pageNumber="428">
<emphasis id="B928F95BFF9D1E77FF4DF8A7FE95C137" bold="true" box="[164,281,1882,1901]" pageId="54" pageNumber="428">Figure 51.</emphasis>
Holotype of
<taxonomicName id="4C5C5ECAFF9D1E77FE41F8A6FD1AC137" box="[424,662,1883,1902]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="428" phylum="Chordata" rank="species" species="nerostagona">
<emphasis id="B928F95BFF9D1E77FE41F8A6FD1AC137" box="[424,662,1883,1902]" italics="true" pageId="54" pageNumber="428">Philautus nerostagona</emphasis>
</taxonomicName>
: A, paratype (BNHS 4245); B, holotype (BNHS 4244) from Kalpetta.
</paragraph>
</caption>
<paragraph id="8BE32549FF9C1E76FCCFFD0EFC0EC51F" blockId="55.[806,1421,754,837]" pageId="55" pageNumber="429">
<emphasis id="B928F95BFF9C1E76FCCFFD0EFBE9C55D" box="[806,1125,754,776]" italics="true" pageId="55" pageNumber="429">
Description of the
<typeStatus id="54E79BEBFF9C1E76FBE8FD0FFBECC55D" box="[1025,1120,754,775]" pageId="55" pageNumber="429" type="holotype">holotype</typeStatus>
:
</emphasis>
The detailed description and illustrations were published by
<bibRefCitation id="EFCD58B8FF9C1E76FB36FCEDFCF3C51F" author="Gururaja KV &amp; Dinesh KP &amp; Palot MJ &amp; Radhakrishnan C &amp; Ramachandra TV" pageId="55" pageNumber="429" pagination="1 - 16" refId="ref43481" refString="Gururaja KV, Dinesh KP, Palot MJ, Radhakrishnan C, Ramachandra TV. 2007 b. A new species of Philautus Gistel (Amphibia: Anura: Rhacophoridae) from southern Western Ghats, India. Zootaxa 1621: 1 - 16." type="journal article" year="2007">
Gururaja
<emphasis id="B928F95BFF9C1E76FABCFCECFA01C57F" box="[1365,1421,784,806]" italics="true" pageId="55" pageNumber="429">et al.</emphasis>
(2007b)
</bibRefCitation>
.
</paragraph>
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<paragraph id="8BE32549FF9C1E76FF79FBD1FDA2C220" blockId="55.[144,759,1068,1146]" pageId="55" pageNumber="429">
<emphasis id="B928F95BFF9C1E76FF79FBD1FE89C265" bold="true" box="[144,261,1068,1087]" pageId="55" pageNumber="429">Figure 52.</emphasis>
<taxonomicName id="4C5C5ECAFF9C1E76FEF8FBD1FD8DC265" box="[273,513,1068,1087]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="ochlandrae">
<emphasis id="B928F95BFF9C1E76FEF8FBD1FD8DC265" box="[273,513,1068,1087]" italics="true" pageId="55" pageNumber="429">Philautus ochlandrae</emphasis>
</taxonomicName>
: A, specimen (SDB 6076) from Kakkayam Reserve Forest; B, specimen (BNHS 4559) from Kakkayam Reserve Forest.
</paragraph>
</caption>
<paragraph id="8BE32549FF9C1E76FF79FB4DFE4CC358" blockId="55.[144,758,1199,1282]" pageId="55" pageNumber="429">
<materialsCitation id="3B342F14FF9C1E76FF79FB4DFF78C358" collectionCode="WGFRS, V" pageId="55" pageNumber="429" specimenCode="A636" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54E79BEBFF9C1E76FF79FB4DFE88C29F" box="[144,260,1200,1221]" pageId="55" pageNumber="429" type="paratype">paratypes</typeStatus>
, ZSI/
<collectionCode id="ED4DBD8CFF9C1E76FEACFB52FE2CC29F" box="[325,416,1199,1221]" pageId="55" pageNumber="429">WGFRS</collectionCode>
/
<collectionCode id="ED4DBD8CFF9C1E76FE41FB52FE36C29F" box="[424,442,1199,1221]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="55" pageNumber="429" type="Museum">V</collectionCode>
/A/633 and ZSI/
<collectionCode id="ED4DBD8CFF9C1E76FD90FB52FD58C29E" box="[633,724,1199,1220]" pageId="55" pageNumber="429">WGFRS</collectionCode>
/
<collectionCode id="ED4DBD8CFF9C1E76FD35FB52FD62C29E" box="[732,750,1199,1220]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="55" pageNumber="429" type="Museum">V</collectionCode>
/ A/637,
<specimenCount id="9D5AEEC0FF9C1E76FF08FB33FED4C2BE" box="[225,344,1230,1252]" pageId="55" pageNumber="429" type="male">two males</specimenCount>
, and ZSI/
<collectionCode id="ED4DBD8CFF9C1E76FE2EFB33FDA8C2BE" box="[455,548,1230,1252]" pageId="55" pageNumber="429">WGFRS</collectionCode>
/
<collectionCode id="ED4DBD8CFF9C1E76FDC5FB33FDB3C2BE" box="[556,575,1230,1252]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="55" pageNumber="429" type="Museum">V</collectionCode>
/
<specimenCode id="DBFA8D32FF9C1E76FDAEFB33FD0DC2BE" box="[583,641,1230,1252]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="55" pageNumber="429" type="Museum">A636</specimenCode>
, a female, from the
</materialsCitation>
<typeStatus id="54E79BEBFF9C1E76FF17FB11FED3C358" box="[254,351,1260,1282]" pageId="55" pageNumber="429" type="holotype">holotype</typeStatus>
locality.
</paragraph>
<paragraph id="8BE32549FF9C1E76FF79FAD3FE74C3DB" blockId="55.[144,758,1326,1409]" pageId="55" pageNumber="429">
<emphasis id="B928F95BFF9C1E76FF79FAD3FE2EC319" box="[144,418,1326,1347]" italics="true" pageId="55" pageNumber="429">Other material studied:</emphasis>
<materialsCitation id="3B342F14FF9C1E76FE44FAD3FD22C319" box="[429,686,1326,1348]" collectionCode="BNHS" pageId="55" pageNumber="429" specimenCode="SDB 6076" specimenCount="1">
<specimenCode id="DBFA8D32FF9C1E76FE44FAD3FDAFC31E" box="[429,547,1326,1348]" collectionCode="SDB" pageId="55" pageNumber="429">SDB 6076</specimenCode>
and
<collectionCode id="ED4DBD8CFF9C1E76FD8DFAD3FD22C319" box="[612,686,1326,1347]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="55" pageNumber="429">BNHS</collectionCode>
</materialsCitation>
4559,
<specimenCount id="9D5AEEC0FF9C1E76FF79FAB0FE8EC338" box="[144,258,1356,1378]" pageId="55" pageNumber="429" type="adult">two adult</specimenCount>
<materialsCitation id="3B342F14FF9C1E76FEF9FAB1FE78C3DB" collectionCode="BNHS" country="India" location="Kakkayam Reserve Forest" pageId="55" pageNumber="429" specimenCount="1" stateProvince="Kerala">
males collected from
<location id="8E837392FF9C1E76FDFBFAB1FF50C3DA" LSID="urn:lsid:plazi:treatment:03F5945FFF9D1E76FCB9FAD1FBB2C2E5:8E837392FF9C1E76FDFBFAB1FF50C3DA" country="India" name="Kakkayam Reserve Forest" pageId="55" pageNumber="429" stateProvince="Kerala">Kakkayam Reserve Forest</location>
,
<collectingRegion id="4998EBABFF9C1E76FF00FA96FEB5C3DB" box="[233,313,1387,1409]" country="India" name="Kerala" pageId="55" pageNumber="429">Kerala</collectingRegion>
,
<collectingCountry id="F34B65D9FF9C1E76FEAEFA96FE08C3DB" box="[327,388,1387,1409]" name="India" pageId="55" pageNumber="429">India</collectingCountry>
(
<tableCitation id="C6DE10F2FF9C1E76FE7CFA96FE67C3DB" box="[405,491,1387,1409]" captionStart="Table 2" captionStartId="10.[113,177,400,421]" captionText="Table 2. Morphometric measurements (in mm) of the specimens used in this study" pageId="55" pageNumber="429">Table 2</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE32549FF9C1E76FF79FA50FE7FC0ED" blockId="55.[144,759,1452,1903]" pageId="55" pageNumber="429">
<emphasis id="B928F95BFF9C1E76FF79FA50FD88C39B" box="[144,516,1452,1474]" italics="true" pageId="55" pageNumber="429">
Diagnosis:
<taxonomicName id="4C5C5ECAFF9C1E76FEE7FA51FD88C39B" box="[270,516,1452,1473]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="ochlandrae">Philautus ochlandrae</taxonomicName>
</emphasis>
can be distinguished from known congeners by the following combination of characters: (1) medium male adult size (SVL 23.9 ±
<quantity id="4CA488ACFF9C1E76FF35F9F5FEBFC044" box="[220,307,1544,1566]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="55" pageNumber="429" unit="mm" value="1.3">1.3 mm</quantity>
,
<emphasis id="B928F95BFF9C1E76FEABF9F4FEDAC044" box="[322,342,1545,1566]" italics="true" pageId="55" pageNumber="429">N</emphasis>
= 5); (2) canthus rostralis rounded; (3) snout rounded; (4) pupil with striking golden yellow dentition-like marks in life; (5) light brown to creamy yellowish, with two invariable and distinct golden yellow streaks running from snout, along the side of head, to near the vent.
</paragraph>
<paragraph id="8BE32549FF9C1E76FF41F93DFB40C5FF" blockId="55.[144,759,1452,1903]" lastBlockId="55.[806,1421,882,934]" pageId="55" pageNumber="429">
<taxonomicName id="4C5C5ECAFF9C1E76FF41F93DFE13C08F" box="[168,415,1728,1749]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="ochlandrae">
<emphasis id="B928F95BFF9C1E76FF41F93DFE13C08F" box="[168,415,1728,1749]" italics="true" pageId="55" pageNumber="429">Philautus ochlandrae</emphasis>
</taxonomicName>
can be differentiated from all other known species of
<taxonomicName id="4C5C5ECAFF9C1E76FE42F923FD97C0A9" box="[427,539,1758,1779]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="genus">
<emphasis id="B928F95BFF9C1E76FE42F923FD97C0A9" box="[427,539,1758,1779]" italics="true" pageId="55" pageNumber="429">Philautus</emphasis>
</taxonomicName>
from the Western Ghats by its two distinct golden yellow streaks, extending from the snout along the side of head to near the vent, and by a pupil with striking golden yellow dentition-like marks in all colour morphs
<emphasis id="B928F95BFF9C1E76FCCFFC8FFC1BC5DD" box="[806,919,882,903]" italics="true" pageId="55" pageNumber="429">Variation:</emphasis>
Measurements of
<specimenCount id="9D5AEEC0FF9C1E76FB9FFC8FFA9BC5D2" box="[1142,1303,882,904]" pageId="55" pageNumber="429" type="generic">six specimens</specimenCount>
including the
<typeStatus id="54E79BEBFF9C1E76FCBCFC6CFC0AC5FC" box="[853,902,913,934]" pageId="55" pageNumber="429">type</typeStatus>
series are given in
<tableCitation id="C6DE10F2FF9C1E76FB9AFC6DFB44C5FF" box="[1139,1224,912,934]" captionStart="Table 2" captionStartId="10.[113,177,400,421]" captionText="Table 2. Morphometric measurements (in mm) of the specimens used in this study" pageId="55" pageNumber="429">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BE32549FF9C1E76FCCFFC2EFBB2C2E5" blockId="55.[806,1421,979,1215]" pageId="55" pageNumber="429">
<emphasis id="B928F95BFF9C1E76FCCFFC2EFCC6C25D" italics="true" pageId="55" pageNumber="429">
Distribution and natural history:
<taxonomicName id="4C5C5ECAFF9C1E76FB58FC2EFCC6C25D" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="55" pageNumber="429" phylum="Chordata" rank="species" species="ochlandrae">Philautus ochlandrae</taxonomicName>
</emphasis>
is known only from the
<typeStatus id="54E79BEBFF9C1E76FB73FC0FFB47C25D" box="[1178,1227,1010,1031]" pageId="55" pageNumber="429">type</typeStatus>
locality in the Kakkayam Reserve Forest of the
<collectingCountry id="F34B65D9FF9C1E76FB3AFBEDFAAAC27C" box="[1235,1318,1040,1062]" name="India" pageId="55" pageNumber="429">Calicut</collectingCountry>
District (
<figureCitation id="136739CCFF9C1E76FCC7FBD2FC1EC21E" box="[814,914,1070,1093]" captionStart="Figure 53" captionStartId="56.[164,243,1842,1861]" captionTargetBox="[184,1423,197,1813]" captionTargetId="figure-0@56.[184,1425,194,1813]" captionTargetPageId="56" captionText="Figure 53. Distribution map. A, Philautus ochlandrae; B, Philautus ponmudi; C, Philautus signatus; D, Philautus sushili sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5445468" httpUri="https://zenodo.org/record/5445468/files/figure.png" pageId="55" pageNumber="429">Fig. 53A</figureCitation>
). The
<typeStatus id="54E79BEBFF9C1E76FC0EFBD2FB94C21E" box="[999,1048,1071,1092]" pageId="55" pageNumber="429">type</typeStatus>
series was collected from the hollow tube of the internodal region of reed brake, about
<quantity id="4CA488ACFF9C1E76FC9BFB91FC3DC2DB" box="[882,945,1132,1154]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="55" pageNumber="429" unit="m" value="2.5">2.5 m</quantity>
above the ground. Breeding (amplexus) and oviposition also take place inside hollow reeds (
<bibRefCitation id="EFCD58B8FF9C1E76FCC7FB54FBBEC2E5" author="Gururaja KV &amp; Dinesh KP &amp; Palot MJ &amp; Radhakrishnan C &amp; Ramachandra TV" box="[814,1074,1193,1215]" pageId="55" pageNumber="429" pagination="1 - 16" refId="ref43481" refString="Gururaja KV, Dinesh KP, Palot MJ, Radhakrishnan C, Ramachandra TV. 2007 b. A new species of Philautus Gistel (Amphibia: Anura: Rhacophoridae) from southern Western Ghats, India. Zootaxa 1621: 1 - 16." type="journal article" year="2007">
Gururaja
<emphasis id="B928F95BFF9C1E76FC4BFB57FC56C2E4" box="[930,986,1193,1215]" italics="true" pageId="55" pageNumber="429">et al.</emphasis>
, 2007b
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>